taxonID	type	description	language	source
317087F1FFA1FFF6EB0FFEFBFC9C70E5.taxon	description	urn: lsid: zoobank. org: act: F 98 C 9058 - CAAE- 4 EED- 9416 - 61 B 94 E 66 C 28 C	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
317087F1FFA1FFF6EB0FFEFBFC9C70E5.taxon	type_taxon	TYPE SPECIES. — Assamhoplites martensis n. sp.	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
317087F1FFA1FFF6EB0FFEFBFC9C70E5.taxon	etymology	ETYMOLOGY. — The genus name is formed by the combination of Assam-, as a reference to the family Assamiidae where the new genus is allocated, and - hoplites from Ancient Greek ὁπλῖται, romanized hoplîtai, the famous soldiers of Ancient Greece who were armed with spears and shields. The spears are metaphorically associated with the highly elongated male pedipalp and the shield with the scutum magnum of these opilionids. Gender masculine.	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
317087F1FFA1FFF6EB0FFEFBFC9C70E5.taxon	diagnosis	COMPARATIVE DIAGNOSIS. — Even though males of both Filopalpinae genera, Assamhoplites n. gen. and Filopalpus, have characteristically long, thread-like pedipalps considerably longer than body length, the contribution of the podomeres elongation to this architecture differs in each genus. The highly elongated male pedipalps in Assamhoplites n. gen. include the strong elongations of the tibia and tarsus, where the tibia and patella have almost the same length. On the contrary, the highly elongated male pedipalp in Filopalpus does not exhibit a strong elongation in the tibia and tarsus (in fact, the tarsus is similar to that of a female), and the patella is remarkably more elongated than the tibia. Assamhoplites n. gen. has a wider granulated ocularium in males and females without other conspicuous armature, whereas Filopalpus exhibits a small and narrow ocularium strongly armed with a number of long, pointed spiniform apophyses, which are longest on the anterior and posterior ocularium rim. In fact, bodies of Filopalpus species are much more roughly granulated than Assamhoplites n. gen., generally with strong paramedian tubercles on the abdominal area. Additionally, the armature of the free tergites, in Filopalpus, is commonly differentiated from the armature of scutal area (at least in males) with a row of slender, drawn-out, pointed tubercles, longest in central third in comparison, Assamhoplites n. gen. has uniformly less body granulation, with no remarkable difference between scutal area and free tergite armature. Another remarkable difference between the two genera is in regard to the degree of sexually dimorphic male chelicerae. Males of Assamhoplites n. gen. exhibit a much bigger and more strongly armed male chelicerae compared to Filopalpus, where the elongated basichelicerite has a weakly marked bulla, is conspicuously granulated, and has a ventral surface heavily armed with many strong, pointed apophyses. The Assamhoplites n. gen. cheliceral hand is also strongly elongated and has a conspicuous modification of the fixed finger. In Filopalpus, the chelicerae are only slightly bigger in males, with few armature differences compared to the females but with neither a strong pointed apophysis in ventral basichelicerite nor a modified fixed finger in the cheliceral hand. The male genitalia of the two genera are very similar (probably signifying close phylogenetic proximity) and only show slight differences. In Assamhoplites n. gen. the apical rim of the pars distalis is slightly convex and contains two small mounts each with an apical macroseta, and the pars distalis, in an unexpanded state, is pointed downwards. In contrast, the penis in Filopalpus has a concave or straight apical rim of the pars distalis, and the pars distalis, in an unexpanded state, is pointed upwards.	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
317087F1FFA1FFF6EB0FFEFBFC9C70E5.taxon	distribution	DISTRIBUTION. — Ethiopia, Oromia province, Ambo, Wonchi crater.	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
317087F1FFA1FFFAE9F0FF5BFECF70E7.taxon	description	(Figs 1 - 5) urn: lsid: zoobank. org: act: 1 B 152754 - 004 B- 4 C 28 - A 4 E 4 - 50 C 9 AC 0 E 4 D 67	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
317087F1FFA1FFFAE9F0FF5BFECF70E7.taxon	materials_examined	TYPE MATERIAL. — Holotype. Ethiopia • 1 ♂; Oromia province, Ambo, Wonchi crater at high point, 6 N road, in moss grass; alt. 3200 m a. s. l.; 4. IV. 1988; Russel-Smith A. leg.; BE _ RMCA _ ARA. Opi. 236192. Paratypes. Ethiopia • 2 ♀; same data as for holotype; BE _ RMCA _ ARA. Opi. 247652 • 1 ♀; same data as for holotype; MACN-Ar 46909.	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
317087F1FFA1FFFAE9F0FF5BFECF70E7.taxon	etymology	ETYMOLOGY. — Patronymic in honor of the German arachnologist, Professor Jochen Martens, in recognition of his contribution to advances in arachnology. Name in the genitive case.	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
317087F1FFA1FFFAE9F0FF5BFECF70E7.taxon	diagnosis	DIAGNOSIS. — The combination of body size (3.57 mm), scutum covered by polygonal tubercles, male pedipalp three times as long as body, femur of the pedipalp notably larger than patella (60 %).	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
317087F1FFA1FFFAE9F0FF5BFECF70E7.taxon	distribution	DISTRIBUTION. — Only known from the type locality.	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
317087F1FFA1FFFAE9F0FF5BFECF70E7.taxon	description	DESCRIPTION Male (BE _ RMCA _ ARA. Opi. 236192) Measurements (mm). Total length: 3.57; carapace length: 1.30; dorsal scutum length: 3.03; carapace max. width: 1.69; dorsal scutum max. width: 2.03. Appendage measurements. Pedipalp. Trochanter: 0.52; femur: 3.69; patella: 2.32; tibia: 1.87; tarsus: 1.77; claw: 0.58. Leg I: trochanter (tr): 0.42; femur (fe): 1.82; patella (pa): 0.70; tibia (ti): 1.13; metatarsus (mt): 1.58; tarsus (tr): 1.03. Leg II: tr: 0.45; fe: 3.06; pa: 0.85; ti: 2.51; mt: 2.03; tr 1.99. Leg III: tr: 0.43; fe: 1.87; pa: 0.72; ti: 1.32; mt: 1.71; tr 1.20. Leg IV: tr: 0.56; fe: 2.52; pa: 0.82; ti: 1.89; mt: 2.67; tr: 1.42. Dorsum (Fig. 2 A-C). Outline almost rectangular with iota (ι) shape. Ocularium rounded and covered by small tubercles with setae, with eyes located on the lateral rim of the ocularium. Scutum covered by small setiferous tubercles, carapace smaller than mesotergum, anterior margin bearing a row of small tubercles, and five pointed apophyses in the following order or length and robustness: 35253. Mesotergal areas clearly defined. Posterior margin and free tergites bearing a row of setiferous tubercles and a row of small tubercles. Venter (Fig. 2 D-F). Ventral surface and anal plate covered by small setiferous tubercles. Chelicerae (Fig. 2 G-I). Basichelicerite covered dorsally by small setiferous tubercles, proventral surface with a row of conical tubercles with setae, and two small setiferous tubercles on the mesal-distal surface. Second article elongated, bearing sparse small tubercles with setae on the frontal surface and a robust spine above the fingers. Fixed finger with a central strong thickening in frontal view. Sparse long setae on the frontal surface. Pedipalps (Fig. 3 A). Long, thin, and filiform trochanter bearing four ventral setiferous tubercles, the distal one longer than the others. Femur smooth, longer than dorsal scutum, bearing sparse setae. All articles with sparse setae. Long distal claw. Legs. All articles irregularly covered with small tubercles. Tarsal count: 5: 9: 5: 6. Genitalia (Fig. 4). Pars distalis composed of an expandable capsula externa (in form of follis sensu Macías-Ordoñez et al. 2010, fig. 13.4 or “ prick funnel ” (Stacheltrichter) sensu Martens 1977) and a capsula interna reduced to a stylus. The expansion of the capsula externa exposing their internal digitiform projections. Four macrosetae on each side of lateral margin of the pars distalis, a pair of dorsal macrosetae flanking the capsula externa and two pairs of ventral small setae. Apical rim of the pars distalis slightly convex and containing two small mounts with an apical macrosetae each. Female (BE _ RMCA _ ARA. Opi. 247652) (Figs 3 B; 5) Similar to male, presenting dimorphism by the presence of a raptorial pedipalp, and chelicera small and not thickened. Measurements (mm). Total length: 3.31; carapace length: 1.10; dorsal scutum length: 3.17; carapace max. width: 1.48; dorsal scutum max. width: 1.90. Appendage measurements. Pedipalp. Trochanter: 0.33; femur: 1.05; patella: 0.51; tibia: 0.68; tarsus: 0.70; claw: 0.60. Leg I: trochanter (tr): 0.32; femur (fe): 1.32; patella (pa): 0.56; tibia (ti): 0.96; metatarsus (mt): 1.16; tarsus (tr): 0.96. Leg II: tr: 0.39; fe: 2.33; pa: 0.79; ti: 1.89; mt: 1.69; tr 1.57. Leg III: tr: 0.38; fe: 1.49; pa: 0.61; ti: 1.07; mt: 1.42; tr 1.06. Leg IV: tr: 0.43; fe: 2.03; pa: 0.65; ti: 1.51; mt: 2.10; tr: 1.28. Tarsal count: 4: 8 / 9: 5: 6 / 7. Female Chelicerae (Fig. 5 G-I). Basichelicerite shorter than that of males, dorsally covered by small setiferous tubercles, ventrally covered by a prolateral row of small conical tubercles. Second segment small, bearing sparse long setae, without spine above the fingers. Female Pedipalps (Fig. 3 E-H). Female pedipalp shorter than male, composed of a raptorial trochanter with a ventral-distal tubercle bearing setae, femur with a ventral row of small setiferous tubercles, tibia and tarsus covered with long setae, the tibia with (Fig. 3 B; 5 H) two prolateral and one retrolateral strong spines, and the tarsus two prolateral and two retrolateral spines. Ovipositor. Simple, bearing long distal bifid setae, as in Martens (2022: fig. 24).	en	Porto, Willians, Kontos, Panagiotis, Pérez-González, Abel (2024): African highland harvestman: New genus and new species of Filopalpinae Martens, 2022 (Opiliones, Assamiidae) from Wonchi crater, Oromia province, Ethiopia. Zoosystema 46 (22): 577-587, DOI: 10.5252/zoosystema2024v46a22, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a22.pdf
