identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3C4D87BDFFCCFFCBFEFEFC0D78AEFE6F.text	3C4D87BDFFCCFFCBFEFEFC0D78AEFE6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callichrotus Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Callichrotus gen. nov.</p>
            <p>(Figs 1–2)</p>
            <p> Type species.  Callichrotus gimmeli sp. nov. , by present designation. </p>
            <p> Diagnosis. The ovate, glabrous body form and antennal club of 2 antennomeres distinguish this genus from most  Picrotini apart from  Picrotus , which also has a broad lateral bead along the prothoracic carina with an abrupt inner margin with a furrow running parallel at the margin of the pronotal disc. It can be distinguished from  Picrotus by its colouration, the widely separated antennal insertions with the frons between antennal insertions not strongly constricted and wider than length of antennal club, presence of a ridge on the vertex of the head, posterolateral angles of pronotum obtuse, prothoracic hypomeron separated from the pronotum by a suture, and abdominal ventrites 1 and 2 connate. The characters marked by an asterisk (*) in the description distinguish  Callichrotus from  Picrotus . </p>
            <p>Description. Body form extremely short and stout, ovate, surfaces shining with extremely short, extremely sparse, inconspicuous, decumbent setae dorsally, erect or suberect setae totally absent; bicoloured. Head without tempora; vertex with ridge, temporal depression immediately anterior to ridge lacking; band of reticulate sculpture lacking. Frontoclypeus not projecting laterally anterior to antennal insertions; raised portion of frons between antennal insertions not strongly constricted, wider than length of antennal club. Transverse ridge above antennal insertions absent. Eye small, conical, not contacting antennal cavity; consisting of about 18 facets; interfacetal setae absent. Antenna inserted into large cavity on head; antennal club consisting of 2 antennomeres; antennomere 9 equal in width to antennomere 8 or almost imperceptibly wider. Mandible with apex bifid, subapical serrations present. Maxillary palpomere 4 subequal in length to 3; palpomere 4 conical but not subulate. Gena without antennal groove; genal spines broadly to narrowly rounded, right to obtuse. Gular sutures present, incomplete, not reaching occipital foramen. Pronotum not explanate, not constricted at base, equal in width to base of elytra, widest in basal half, distinctly transverse; anterior angles weakly to strongly projecting anteriorly (extending anterior to cervical foramen of prothorax), without a distinct platform; lateral carina present and complete, not bearing setigerous tubercles, lateral bead along the prothoracic carina broad, about as wide as antennal club, with an abrupt inner margin and furrow running parallel at the margin of the pronotal disc; disc without transverse basal impression extending across width of pronotum; paramedial carinae and paralateral plicae absent; posterolateral angles obtuse. Prothoracic hypomeron separated from pronotum by suture. Prosternum with anterior margin on same plane as disc; prosternal process with lateral marginal beads weakly present, parallel-sided, connected across apex, process without narrow longitudinal depression medially, process expanded apically, apex truncate and not crenulate, lacking setae; procoxal cavity with anterolateral notch. Scutellar shield clearly visible and transverse. Elytron lacking humeral tooth, lacking subbasal and subapical impressions; subapical gape present; punctation coarse, densely concentrated in scutellar region; vestiture originating in punctures, uniformly short and recumbent. Hind wing vestigial. Mesoventrite with mesoventrital cavity bowl-like, glabrous, flanked by sharp carinae. Mesanepisternal pit absent. Metaventrite with short, crenulate postcoxal lines; discrimen absent, posterior notch of metaventrite absent. Metendosternite with anterior tendons widely separated or absent. Tarsi 5-5- 5 in female, 5-5- 4 in male; tarsi compact, tarsomere 5 wider than preceding tarsomeres in lateral view; pro- and mesotarsomere 4 asetose; mesotarsomere 3 not lobed, with few setae; mesotarsomeres 1–3 of subequal to equal lengths with 1 longer than 2, 3 and 4 equal, 4 slightly shorter than 3, mesotarsomere 5 as wide as tarsomeres 1–4 combined. Abdominal ventrites 1–3 solidly fused with sutures nearly obliterated, with medial calli absent, lateral calli present or absent, intersegmental crenulations absent; ventrite 1 with intercoxal process broadly rounded, with postcoxal lines absent; medio-basal thickenings of ventrites 3–5 absent; apex of ventrite 5 with crenulations. Abdominal spiracles on segment VII with openings present and not larger in diameter than spiracle VI, texture smooth and atrium rounded and saclike. Aedeagus with tegminal strut absent, tegminal arms contiguous; parameres separate and articulated to phallobase with inner surface concave, fused parameres combined about 3 times longer than wide; apices multisetose; attachment point to phallobase not constricted, interparameral process present; basipenis 4 times longer than distipenis, without median carina; distipenis rectangular and elongate and about 2 times longer than wide, outer rims not crenulate, lateral lobes narrowly separated, symmetrical; internal sac with a single arrow-shaped plate. Etymology. The generic name is derived from a combination of the Latin prefix ‘ calli- ’, meaning beautiful, in reference to its striking colouration and ‘- chrotus ’, a genus ending used in one other cryptophagid. The gender is masculine.</p>
            <p> Remarks. The species upon which this genus is based was originally considered an undescribed member of  Picrotus by GIMMEL &amp; LESCHEN (2022: fig. 69) mainly by the presence of a broad lateral pronotal bead shared by both genera, and not present elsewhere in the tribe. </p>
            <p>Distribution. New Zealand.</p>
            <p> Included species.  Callichrotus gimmeli sp. nov.</p>
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	https://treatment.plazi.org/id/3C4D87BDFFCCFFCBFEFEFC0D78AEFE6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFCBFFCBFE96FDF67E06FA8F.text	3C4D87BDFFCBFFCBFE96FDF67E06FA8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callichrotus gimmeli Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Callichrotus gimmeli sp. nov.</p>
            <p>(Figs 1–2)</p>
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                 Type material.   HOLOTYPE: ♁ (NZAC), labelled: ‘ NEW ZEALAND, WD, Mt Aspiring NP Haast Hwy, Bridle Tk, Davis Flat end footbridge from hwy. 15 Jan 2019, K. Marske / Mixed beech forest Sifted leaf and woody litter in moss free areas. -44.12639, 169.34029, 403 m, L siftate. KM360 / NZ  
                <a title="Search Plazi for locations around (long 169.34029/lat -44.12639)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=169.34029&amp;materialsCitation.latitude=-44.12639">Arthropod Collection Private Bag</a>
                 92170 Auckland  
                <a title="Search Plazi for locations around (long 169.34029/lat -44.12639)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=169.34029&amp;materialsCitation.latitude=-44.12639">New Zealand</a>
                 NZAC04254545  ’. PARATYPES (11; FMNH, NZAC). SOUTH IS- LAND: NN: 1 (FMNH), Arthur Range, W side Flora Saddle, 1 Jan 1985, leaf log litter forest floor,  Nothofagus sp. mossy forest, A. Newton, M. Thayer, [41°11.399′S, 172°44.474′E], NZAC 04234533; 1, Nelson, Canaan, 13 Sep 1964, litter, L.P. Merchant, [40°56.580′S, 172°53.574′E], NZAC 04234454; 1, Nelson, Mount Arthur, Flora Track, 19 May 1966, moss, J.I. Townsend, [41°11.567′S, 172°44.067′E], NZAC 04272529. BR: 1, Boatmans Creek, 4 Oct 1971, Beech forest, litter, G. Kuschel, [42°1.333′S, 171°52.117′E], NZAC 04272530; 1, Boatmans Creek, 4.5km SE Cronadun, Nov 1971, litter,J. McBurney, [42°2.682′S, 171°53.729′E], NZAC 04234473; 2, Buller, Capleston, 6 Nov 1971, moss, J.C. Watt, [42°4.054′S, 171°55.326′E], NZAC 04234439, NZAC 04272549; 1, Buller, Capleston, Redman Creek, 8 Nov 1972, moss, J.C. Watt, [42°2.692′S, 171°53.015′E], NZAC 04234465; 1, Lewis Pass, 12 Nov 1964, moss, G. Kuschel, [42°22.712′S, 172°23.967′E], NZAC 04235177. NC: 1, Arthurs Pass, 2 Mar 1966, moss tussock, J.I. Townsend, [42°54.439′S, 171°33.566′E], NZAC 04234498. WD: 2, WD, Mount Aspiring National Park, Haast Highway, Bridle Track, Davis Flat end footbridge from highway, 15 Jan 2019, mixed beech forest sifted leaf and woody litter in moss free areas, K. Marske, [44°7.583′S, 169°20.417′E], NZAC 04254530, NZAC 04254538; 1, Fantail Falls, 3 km NE Haast Pass, sifted wood mould and litter, 84/20, J.C.Watt, [44°04.69′S 169°23.25′E], NZAC 04272550. OL: 1, Makarora, McKerrow Range, 23 Jan 1978, sifted litter, G. Kuschel, [44°14.096′S, 169°14.061′E], NZAC 04234623. 
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            <p>Description. Length 1.30–1.50 mm. Colour of body dark to light reddish-brown and tan, legs, and sometimes the antennae and elytra lighter; and each elytron with broad maculation that reaches the suture in some specimens. Dorsal setae uniformly silver, pronotum and base of elytra with long, decumbent setae, lacking on posterior surfaces of the pronotal disc and maculation of the elytron. Ventral surfaces with recumbent setae, slightly shorter in length compared with those on dorsum; finely to densely punctate on most surfaces, punctation of the vertex of the head dense but not areolate, punctation of metaventrite denser than that of prosternum and more uniform, punctation of abdominal ventrites 1 to 4 less dense and impressed than metaventrite, and on ventrite 5 denser and smaller towards apex, which lacks a patch of posteriorly-directed setae. Head rugose-punctate, with smooth and shiny interspaces, punctures not forming transverse rows; vertex with ridge; transverse ridge above antennal insertions absent. Eye wider than long. Antennomere 2 globular, as long as wide, antennomere 3 conical, longer, as wide as 2, antennomere 4 subequal in width to 3, shorter than 3 and similar in length to 5, antennomeres 6–8 similar, antennomere 9 distinctly larger than antennomere 8 and narrower than antennomeres 10 and 11, antennomere 10 weakly transverse, about as wide as antennomere 11, antennomere 11 about 1.2 times length of than 10, slightly shorter than 9 and 10 combined. Pronotum transverse, pronotal length/width ratio ~0.73, widest at middle or slightly widest at base (some specimens from BR); lateral edges parallel and convergent anteriorly; pronotal punctation similar to that of frons but more irregular over much of the disc, punctures separated by about one puncture diameter or less, with smooth and shiny interspaces; anterior part of the lateral furrow pinched; posterior margin weakly sinuate without weak scutellar lobe. Prosternal process about 0.5 times wide as prosternal length. Elytra about 1.14 times as long as combined widths, about 1.16 times width of pronotum at greatest width, about 1.81 times length of pronotum length; punctation extremely sparse, fine to coarse, densely concentrated in scutellar region where punctures are separated by up to 2 puncture diameters, denser and becoming fused towards midline, sparser on disc and absent from macula; parasutural stria absent, apical gape present. Abdominal ventrite 1 with broad, truncate intercoxal process; intercoxal process wider than mesoventrital process. Aedeagus with apices with 4 primary setae distinctly longer than parameres.</p>
            <p>Etymology. The epithet honors a colleague and friend of the first author, Matthew Gimmel, for his collaborative work on cryptophagids and contribution towards beetle systematics.</p>
            <p>Distribution. South Island: NN, BR, NC (Fig. 21).</p>
            <p> Remarks. This species is known from 12 specimens collected in the South Island. Half were taken in moss samples, a significantly higher proportion of all other species treated in this paper, including species of  Picrotus that were collected three times from moss. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFCBFFCBFE96FDF67E06FA8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFCBFFCBFC47FAD67E31F945.text	3C4D87BDFFCBFFCBFC47FAD67E31F945.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Picrotus Sharp 1886	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Picrotus Sharp, 1886</p>
            <p>(Figs 3–5)</p>
            <p> Type species.  Picrotus thoracicus Sharp, 1886 , by monotypy. </p>
            <p> Diagnosis.  Picrotus is easily distinguished from most  Picrotini by having a broad body form with a glabrous body surface, an antennal club of 2 antennomeres, and a wide lateral bead along the prothoracic carina, which it shares with  Callichrotus . It can be distinguished from  Callichrotus by the narrower frons between the antennal insertion and free abdominal ventrites. A full description of this genus is available in GIMMEL &amp; LESCHEN (2022). </p>
            <p> Included species.  Picrotus thoracicus Sharp, 1886 ;  P. wairarapa sp. nov.</p>
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	https://treatment.plazi.org/id/3C4D87BDFFCBFFCBFC47FAD67E31F945	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFCBFFC6FC09F91C7E50FC8F.text	3C4D87BDFFCBFFC6FC09F91C7E50FC8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Picrotus thoracicus Sharp 1886	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Picrotus thoracicus Sharp, 1886</p>
            <p>(Figs 3–4)</p>
            <p> Picrotus thoracicus Sharp, 1886: 395 . =  Picrotus sanguineus Broun, 1893: 1098 . Synonymy by LESCHEN </p>
            <p> (1996: 606). =  Picrotus pensus Broun, 1910: 40 . Synonymy by LESCHEN (1996: 606). </p>
            <p>Material examined (127; AMNZ, LUNZ, NZAC). NORTH ISLAND: WO: 1, Maungatautari, 20 Nov - 17 Dec 2010, C. Watts, 37°58.133′S, 175°33.683′E, AMNZ 86125; 1 (AMNZ), Mount Maungatautari track, Tari Road end, 1 Sept 2003,rotten wood,S.E.Thorpe,[38°0′S, 175°34.8′E];</p>
            <p> 1, Mount Pirongia, Grey Road track, 30 Jan 2003, on underside of log, S.E. Thorpe, [37°58.145′S, 175°8.826′E], NZAC 04234610; 1, Mount Pirongia, Mahaukura Track, 18 Nov 2005, J. Nunn Collection, [37°59.552′S, 175°7.282′E], NZAC 04234525 2 (LUNZ), Mount Pirongia, track at end of Waite Road, 6 Jan 2006, under bark dead branch on ground, S.E. Thorpe, [37°57.628′S, 175°8.162′E]; 7, Mount Pirongia, track from end of Waite Road, 8 Jan 2002, under bark of dead branch on ground, S.E. Thorpe, 37°58.2′S, 175°7.8′E, AMNZ 47938, AMNZ 47939, AMNZ 47940, AMNZ 47941, AMNZ 47942, AMNZ 47943, AMNZ 47944; 1, Mount Pirongia, Waite Road track, 13 Dec 2004, under logs, S.E. Thorpe, 37°57.867′S, 175°8.067′E, AMNZ 67397; 1, W of Otorohanga, 6 Sep 1942, C.E. Clarke, 38°11.1′S, 175°9.017′E, AMNZ 30942. TK: 1, Tahora Scenic Reserve, west end of road tunnel, sifted wood and leaf litter, K. Marske, R. Leschen, T. Buckley, 39°1.218′S, 174°47.968′E, KM196, NZAC 03019739. TO: 1, Mt Ruapehu,Rangataua Forest, 15 Dec 1981, beech leaf litter, K.A.J. Wise, 39°25.6598S, 175°27.207′E, AMNZ 38132; 1, Waituhi Saddle, 2 Sep 1993, decayed wood, J. Nunn Collection, [38°51.859′S, 175°32.733′E], NZAC 04114151. GB: 1 (FRNZ), Putere, Site 8495 – W, 16 Jan 13 - 13 Feb 2013, M. Scott, 38°56.39′S, 176°55.004′E; 1 (FRNZ), Putere, Site 8661- 16 Jan 13 - 13 Feb 2013, E M. Scott 39°1.835′S, 177°9.439′E. RI: 1, Kahuterawa, 27 Apr 1961, R.A. Cumber, [40°30.023′S, 175°35.268′E], NZAC 04238382; 1, Mataroa near Taihape, Paengaroa Scientific Reserve, 30 Mar 2006, T. Buckley, D. Seldon, R. Hoare, [39°38′S, 175°43′E], RL1106, NZAC 04114145;1, Mataroa near Taihape,Paengaroa Scientific Reserve, 30 Mar 2006, T. Buckley, D. Seldon, R. Hoare, 39°38′S, 175°43′E, RL1105, NZAC 04234436;1, Ruahine State Forest Park, Oroua River, 19 Jan 2008, leaf litter, K. Marske, R. Leschen, T. Buckley, [39°57.66′S, 176°0.841′E], KM211, NZAC 04136820; 6, RI, Ruahine State Forest Park, Oroua River, 19 Jan 2008, sifted leaf litter, K. Marske, R. Leschen, T. Buckley, 39°57.66′S, 176°0.841′E, KM211, NZAC 03018640, NZAC 04114148, NZAC 04114150, NZAC 04234441, NZAC 04234520, NZAC 04234534; 1, Wharite Summit, 23 Oct 1994, litter, J. Nunn Collection, [40°15.274′S, 175°51.454′E], NZAC 04234523. HB: 1, Wairarapa, Waewaepa Range, 4 Sep 1965, litter, J.I. Townsend, [40°25.859′S, 176°3.157′E], NZAC 04238601. WI: 4, Manawatu, Pohangina Field Centre, 13 Jan - 10 Feb 2004, I. Johnston, 40°2.917′S, 175°57.317′E, AMNZ 72127, AMNZ 72129, AMNZ 72130, AMNZ 72131, AMNZ 72132; 3, Manawatu, Pohangina Field Centre, 16 Dec 2003 - 13 Jan 2004, I. Johnston, 40°2.917′S, 175°57.317′E, AMNZ 72126, AMNZ 72128. WN: 2, Akatarawa Saddle, 17 Mar 1978, litter, S.B. Peck, [40°56.907′S, 175°6.502′E], NZAC 04238561, NZAC 04238623; 3, Akatarawa Saddle, 7 Mar 1978, litter, S.B. Peck, J. Peck, [40°56.851′S, 175°6.485′E], NZAC 04234519, NZAC 04234524, NZAC 04234539; 1, Akatarawa, Kakanui Park, 9 Mar 1978, bracket fungi, S.B. Peck, J. Peck, [40°57.891′S, 175°8.123′E], NZAC 04114157; 6, Kaitoke Regional Park, Pakuratahi Forks, 15 Apr 2005, leaf litter, R. Leschen, C. McGuiness, [41°3.374′S, 175°11.698′E], RL975, NZAC 04134363, NZAC 04224147, NZAC 04224217, NZAC 04233895, NZAC 04234035, NZAC 04272539; 2, Kapiti Island, Wilkinson Track, 21 Jan 2008, under/in dead wood, K. Marske, R. Leschen, T. Buckley, 40°51.145′S, 174°55.722′E, KM226, NZAC 03018789, NZAC 03019129; 1, Levin, Waiopehu Scientific Reserve, 16 Nov 2015, in tawa litter, J. Nunn, [40°38.338′S, 175°19.771′E], NZAC 04235146; 1, Lower Hutt, Woburn Reserve, 10 Sep 1994, J. Nunn Collection, [41°13.176′S, 174°51.817′E], NZAC 04234576; 3, Manawatu District Balance Bridge, 19 Dec 1948, leaf mould, A.E. Brookes Collection, [40°20.244′S, 175°49.1′E], NZAC 04238331, NZAC 04238384, NZAC 04238504; 1, Rimutaka Forest Park, Graces Stream Track, 15 Aug 1983, litter, J. Nunn Collection, [41°20.482′S, 174°55.779′E], NZAC 04234545; 6, Rimutuka Hill, Rimutuka Summit Track, 23 Jan 2008, sifted wood and leaf litter, K. Marske, R. Leschen, T. Buckley, [41°6.897′S, 175°13.7′E], KM240, NZAC 03018933, NZAC 04233873, NZAC 04233952, NZAC 04233923, NZAC 04233960, NZAC 04233970; 4, Rimutuka Summit Track, 15 Apr 2005, leaf litter, R. Leschen, C. McGuiness, [41°6.897′S, 175°13.7′E], RL974, NZAC 04233533, NZAC 04233539, NZAC 04233548, NZAC 04234023; 2, Summit Akatarawa Road, 27 Mar 1983, litter, J. Nunn Collection, [40°56.851′S, 175°6.485′E], NZAC 04114149, NZAC 04234452; 2, Summit Akatarawa Road, 19 Aug 1991, J. Nunn Collection, [40°56.851′S, 175°6.485′E], NZAC 04234444, NZAC 04234455; 1, Tararua Forest Park 4 km along Waiotauru Road, 16 Oct 1991, J. Nunn Collection, [40°56.765′S, 175°6.907′E], NZAC 04234448; 2, Tararua Forest Park Track, Akatarawa Saddle, 17 Jan 1984, litter, H.P. McColl, [40°56.902′S, 175°6.487′E], NZAC 04238347, NZAC 04238503; 1, Tararua Forest Park Track, Akatarawa Saddle, 8km from SH, 17 Jan 1984, litter, H.P. McColl, [40°56.851′S, 175°6.485′E], NZAC 04235251; 1, Tararua Forest Park, Akatarawa Saddle, 22 Jan 2008, sifted leaf litter, K. Marske, R. Leschen, T. Buckley, 40°56.9′S, 175°6.577′E, KM231, NZAC 03020463; 1, Tararua Forest Park, Kakanui, 21 Aug 1976, J. Nunn Collection, [40°57.868′S, 175°8.151′E], NZAC 04234447; 4, Tararua Forest Park, Mount Holdsworth, 29 Jan 2008, sifted wood and leaf litter, K. Marske, R.Leschen,T.Buckley, [40°54.408′S, 175°28.432′E], KM273, NZAC 03020609, NZAC 04135036, NZAC 04136059, NZAC 04138921; 1, Tararua Forest Park,Waiohine Gorge, Totara Flats Track, 25 Jan 2008, sifted wood and leaf litter, K. Marske, R. Leschen, T. Buckley, 40°59.628′S, 175°23.28′E, KM255, NZAC 03019253; 1, Tararua Forest Park,Waiotauru Road, 5 Jul 1999, E.Spurr, [40°58.179′S, 175°15.647′E], NZAC 04238619; 1, Tararua Forest Park, Waiotauru Road, 11 Jul 2000, E.Spurr,[40°58.179′S, 175°15.647′E], NZAC 04230219; 1, Tararua Forest Park, Waiotauru Road, 22 Oct 1991, in litter under silver beech, J. Nunn Collection, [40°58.179′S, 175°15.647′E], NZAC 04234679; 1, Tararua Range, Mangaore, Mangahao, 20 Jan 2008, dead wood, K. Marske, R. Leschen, T. Buckley, [40°34.456′S, 175°26.961′E], KM223, NZAC 04139573; 1, Tararua Range, Mount Holdsworth, 16 Apr 2005, leaf litter, R. Leschen, [40°52.46′S, 175°25.043′E], RL976, NZAC 04224175. WA: 2, Fensham Scenic Reserve, Carterton, 27 Jan 2008, ex freshly dead pohutukawa, at night, K. Marske, R. Leschen, T. Buckley, [40°59.594′S, 175°30.141′E], KM265, NZAC 04137404, NZAC 04138376; 6, Puketoi Range, near Mount Butters, 28 Jan 2008, forest patch in gully below road, sifted leaf litter and rotten wood, K. Marske, R. Leschen, T. Buckley, [40°33.723′S, 176°3.364′E], KM271, NZAC 03018630, NZAC 04135012, NZAC 04137696, NZAC 04137789, NZAC 04139191, NZAC 04139996; 1, S Rimutaka, White Stone Trig, 12 July 1992, black beech litter, K. Stokes, [41°19.383′S, 175°6.069′E], NZAC 04114155. SOUTH ISLAND: NN: N Nelson nr Riwaka, Takaka Hill, 5 Mar 1957,  Nothofagus bark, R. Crowson, [41°0′18”S,172°53′26”], NZAC 04272527; 1, Nelson, 14 Apr 1943, E.S. Gourlay, [41°16.355′S, </p>
            <p> 173°17.884′E], NZAC 04238498; 1, Nelson, Canaan, 18 Apr 1966, under bark  Nothofagus menziesii, J.C. Watt , [40°56.582′S, 172°53.349′E], NZAC 04238616;11, Takaka, near Riwaka, 5 Mar 1957,  Nothofagus bark, R. Crowson, [41°2.875′S, 172°55.359′E], NZAC 04182083, NZAC 04238615, NZAC 04238342, NZAC 04238344, NZAC 04238373, NZAC 04238383, NZAC 04238603, NZAC 04238607, NZAC 04238609, NZAC 04238614; 2, Upper Maitai, 14 Feb 1943, E.S. Gourlay, [41°17.583′S, 173°22.157′E], NZAC 04182057, NZAC 04238341; 1, Maitai, Wooded Park, Roadge S Branch, 14 Feb 1943, dead beech, [41°20.135′S, 173°20.895′E], NZAC 04238372. SD: 3, SD, Maud Island, 4-6 Mar 1953,E.S.Gourlay,[41°1.498′S, 173°53.694′E], NZAC 04238545, </p>
            <p>NZAC 04238604, NZAC 04238617;3, Mount Stokes Track,Mount Stokes Scenic Reserve., 19 Feb 2009, sifted leaf litter and rotten wood, K. Marske, R. Leschen, 41°5.253′S, 174°7.994′E, KM320, NZAC 03019183, NZAC 04234445, NZAC 04234446; 1, Port Underwood Saddle, 3 Dec 1969, litter, J.I. Townsend, [41°17.359′S, 174°6.637′E], NZAC 04238339; 1, Port Underwood Saddle, 3 km SSE Curious Cove, sifted litter, R. Leschen, M. Anderson, T. Buckley, Y. Chen, 41°16.423′S, 174°7.195′E, RL2241, NZAC 03038715; 1, Port Underwood Saddle, 3 km SSE Curious Cove, 15 Nov 1999, sifted leaf litter,R.Leschen, 41°17′S, 174°5′E, RL480, NZAC 04114152; 1, Stephens Island, 14-28 Jan 1933, E.S. Gourlay, [40°40.056′S, 174°0.019′E], NZAC 04238602.</p>
            <p> Diagnosis.  Picrotus thoracicus is easily distinguished from  P. wairarapa sp. nov. by the presence of a row of foveolate punctures with a broad medial puncture at the base of the pronotal disc and the prosternal process which narrows and is apically truncate. </p>
            <p>Redescription. Length 1.25–1.35 mm. Colour of body dark to light reddish-brown, mouthparts, legs, and antennae lighter. Surfaces of body glabrous. Dorsal setae golden, sparse, decumbent. Ventral surfaces with sparse, recumbent and semi-recumbent setae, longer in length compared with those on dorsum; finely to shallowly punctate on most surfaces, punctation of head dense, areolate, central disc of metaventrite glabrous with large punctures similar to those of prosternum, punctation of abdominal ventrites 1 to 4 more dense and impressed than metaventrite, on ventrite 5 denser and smaller towards apex, which lacks a patch of posteriorly-directed setae. Head smooth with large, ovate punctures with shiny interspaces, punctures above eye forming a transverse row; vertex without ridge; transverse ridge above antennal insertions present, clypeal constriction about as wide as length of antennomere 1. Eye ovate, contacting antennal cavity, consisting of about 12 facets. Antennomere 2 globular, as long as wide, antennomere 3 subconical, longer and as wide as 2, widths of antennomeres 3–8 equal, antennomeres 4–8 similar, antennomere 4 shorter than 3, antennomere 9 scarcely wider than 8 and narrower than 10 and 11, antennomere 10 transverse and wider than 11, antennomere 11 about as long as 10, shorter than 9 and 10 combined. Pronotum transverse, pronotal length/width ratio ~0.72, widest in basal third; lateral edges parallel in basal half and convergent anteriorly; anterior angles weakly projecting anteriorly (not extending anterior to cervical foramen of prothorax); anterior part of the lateral furrow not pinched; pronotal disc impunctate, basally impressed with a row of foveolate punctures and a broad medial puncture; posterolateral angles narrowly rounded, acute, and projecting posteriorly; posterior margin strongly sinuate with weak scutellar lobe. Prothoracic hypomeron fused to prosternum, not separated by suture. Prosternal process with well-developed lateral marginal beads, with narrow longitudinal depression medially, process not expanded apically behind procoxae, apex narrowed and then truncate, width 0.63 longer than length of prosternum; procoxal cavity without anterolateral notch. Scutellar shield mostly hidden except for extreme apex. Elytra about 0.98 times as long as combined width, about 1.05 times width of pronotum at greatest width, about 1.37 times length of pronotum; surface impunctate; parasutural stria absent. Tarsi moderately slender, tarsomere 5 as wide as preceding tarsomeres in lateral view; mesotarsomeres 1–3 of subequal to equal lengths, with 1 slightly shorter than 2, 3 and 4 equal, 4 shorter than 3, mesotarsomere 5 equal to 1–4 combined. Abdominal ventrites free; first abdominal ventrite with intercoxal process that is weakly curved at the apex, narrower than mesoventrital process. Aedeagus with parameres fused to phallobase and broadest at their bases and separated at the apex, combined length of fused parameres about 1.2 times as wide as long; apices with 4 primary setae distinctly shorter than parameres, two at apex longer than two subapical setae; interparameral process absent; basipenis 6.0 times length of distipenis; distipenis rectangular, about 1.1 times longer than wide, lateral lobes widely separated and symmetrical; internal sac with a pair of thin endophallites that extend about 1/2 the length of the sac.</p>
            <p>Distribution. North Island: WO, TO, GB, HB, RI, WI. WN, WA. South Island: NN, SD (Fig. 21).</p>
            <p> Remarks.  Picrotus thoracicus ranges from about the middle of North Island south to the north of the South Island. It may be abundant in leaf litter, dead wood, under bark, from fungus, wood and leaf mould and rarely from moss. All specimens we have examined that we consider as part of this species have the foveolate punctures at the base of the pronotal disc, though the strength and size of the punctures, including the median puncture, vary from moderately deeply impressed and circular to transverse and less impressed. Information on type material was given in LESCHEN &amp; GIMMEL (2012). </p>
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	https://treatment.plazi.org/id/3C4D87BDFFCBFFC6FC09F91C7E50FC8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFC6FFC5FC64FCDF78BEFC8F.text	3C4D87BDFFC6FFC5FC64FCDF78BEFC8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Picrotus wairarapa Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Picrotus wairarapa sp. nov.</p>
            <p>(Figs 5, 19F)</p>
            <p> Type material.   HOLOTYPE: ♁ (NZAC), labelled: ‘ New Zealand WA Hikorangi 13 Sep 1982 J. C. Watt // Sifted woodmould 82/75 // Entomology Div. D.S.I.R. New Zealand / NZ Arthropod Collection Private Bag 92170 Auckland  New Zealand NZAC04238506  ’. PARATYPES (31; NZAC). NORTH ISLAND: WA: 2, Haurangi, Aorangi Mountains, 2 Sep 1965, moss in forest, J.I. Townsend, [41°21.978′S, 175°23.713′E], NZAC 04238599, NZAC 04238610; 5, Hikorangi, 13 Sep 1982, sifted wood mould, J.C. Watt, [41°8.757′S, 175°47.162′E], NZAC 04114153, NZAC 04234428, NZAC 04234431, NZAC 04234537, NZAC 04234568; 5 (same as holotype), Hikorangi, 13 Sep 1982, litter, J.C.Watt,[41°8.801′S, 175°47.114′E], NZAC 04182063, NZAC 04238388, NZAC 04238556, NZAC 04238598, NZAC 04238605; 2, Ngaumu State Forest., 27 Jan 2008, under pine logs in pine plantation, K. Marske, R. Leschen, T. Buckley, 41°1.693′S, 175°57.411′E, KM261, NZAC 04137263, NZAC 04138374; 3, Pongaroa, 31 Dec 1980, wood mould, J.C. Watt, [40°32.570′S, 176°11.598′E], NZAC 04136583, NZAC 04138616, NZAC 04139481; 1 Pongaroa, 31 Dec 1980, litter, J.C. Watt, [40°32.570′S, 176°11.598′E], NZAC 04135077; 9, Putangirua Pinnacles Scenic Reserve, Palliser Bay, 24 Jan 2008, dead wood, K. Marske, R. Leschen, T. Buckley, 41°27.011′S, 175°14.435′E, KM250, NZAC 03019073, NZAC 04135546, NZAC 04135944, NZAC 04136715, NZAC 04136806, NZAC 04136811, NZAC 04138859, NZAC 04234039, NZAC 04235138; 5, Sutherland Vehicle Track, Aorangi Range, 24 Jan 2008, sifted wood and leaf litter in secondary forest on former grazing land, K. Marske, R. Leschen, T. Buckley, 41°25.239′S, 175°21.551′E, KM245, NZAC 03018832, NZAC 04272551, NZAC 04272552, NZAC 04272553, NZAC 04272554; 2, West of Lake Wairarapa Reserve, 20 Apr 1997, in humus under bark of dead  Pseudopanax, J. Nunn , [41°10.155′S, 175°18.676′E], NZAC 04235296, NZAC 04272546. </p>
            <p> Diagnosis.  Picrotus wairarapa sp. nov. is easily distinguished from  P. thoracicus by the lack of a basal row of foveolate punctures and broad medial puncture on the pronotum. The slightly longer prosternal process that is apically rounded or ligulate and the longer parameral setae will also distinguish this species from  P. thoracicus . </p>
            <p> Description. Length 1.10–1.75 mm. With characters of  P. thoracicus , except as follows. Head with clypeal constriction slightly wider than length of antennomere 1. Pronotum with pronotal length/width ratio ~0.64; pronotal base weakly impressed, without a row of foveolate punctures and without a broad medial puncture. Prosternal process with apex rounded or ligulate, width 1.16 times shorter than total length of prosternum. Elytra about as long as combined width, about 1.10 times width of pronotum at greatest width, about 1.70 times length of pronotum. First abdominal ventrite with intercoxal process subrounded at apex. Aedeagus with parameres fused to phallobase and broadest at their bases and not separated at the apex, combined length of fused parameres about 0.8 times as long as wide; apices with 3–4 primary setae distinctly as long as parameres, and of equal lengths; basipenis about 6.0 times length of distipenis; distipenis rectangular and elongate, about 1.5 times as long as wide; internal sac with a pair of thin endophallites that extend about 2/3 the length of the sac. </p>
            <p>Etymology. The specific epithet is based on its distribution in the Wairarapa; noun in apposition.</p>
            <p>Distribution. North Island: WA (Fig. 21).</p>
            <p> Remarks.  Picrotus wairarapa sp. nov. is restricted to the Wairarapa in the southeast of the North Island. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFC6FFC5FC64FCDF78BEFC8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFC5FFC5FEE3FCD67B1BF9A5.text	3C4D87BDFFC5FFC5FEE3FCD67B1BF9A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus Broun 1893	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus Broun, 1893</p>
            <p>(Figs 6–19)</p>
            <p> Type species.  Thortus ovalis Broun, 1893 , by monotypy. </p>
            <p> Diagnosis. This genus is easily distinguished from most other New Zealand  Picrotini by having glabrous to semiglabrous body surfaces, the antenna inserted into a deep cavity, often accompanied by a constricted frons that is narrower than width of the antennal club, a reduced number of eye facets in many species, and the abdominal postcoxal lines usually present and the absence of hind wings. It can be distinguished from  Callichrotus and  Picrotus by having an elongate body, an antennal club of 3 antennomeres and a narrow lateral bead along the prothoracic carina. The genitalia also differ by having asetose or microsetose apices of the parameres. </p>
            <p> Remarks. Based on the cladistic study below,  Thortus is monophyletic, and we adhere to the definition of the genus as outlined in GIMMEL &amp; LESCHEN (2022). </p>
            <p> Included species.  Thortus amoenus (Broun, 1912) ;  T. bullerensis sp. nov. ;  T. crowsoni sp. nov. ;  T. helmorei sp. nov. ;  T. latus sp. nov. ;  T. lobatus sp. nov. ;  T. luscus sp. nov. ;  T. michauxi sp. nov. ;  T. ovalis Sharp ;  T. parallelus sp. nov. ;  T. simplex sp. nov. ;  T. sulcatus sp. nov. ;  T. tioripatea sp. nov.</p>
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	https://treatment.plazi.org/id/3C4D87BDFFC5FFC5FEE3FCD67B1BF9A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFC5FFC2FEA5F98B7FAAFBAF.text	3C4D87BDFFC5FFC2FEA5F98B7FAAFBAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus amoenus (Broun 1912)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus amoenus (Broun, 1912)</p>
            <p>(Fig. 6)</p>
            <p> Cryptophagus amoenus Broun, 1912: 423 . </p>
            <p> Thortus amoenus : GIMMEL &amp; LESCHEN (2022: 106). </p>
            <p>Material examined (58; AMNZ, BMNH, FRNZ, NZAC). NORTH ISLAND: TO: 1 (FRNZ), Pukemako, 12 Feb - 19 Mar 2020, 37°51.648′S, 175°32.027′E; 1 (FRNZ), Pukemako pitfall 2 14 Dec - 22 Feb 2023, C.Wardhaugh, 37°51.648′S, 175°32.027′E; 1 (BMNH),Waimarino, Jan 1911, [39°26.368′S, 175°7.732′E]. GB: 1 (FRNZ), Putere, Site 8495, 16 Jan 13 - 13 Feb 2013, E M. Scott, 38°56.39′S, 176°55.004′E. RI: 1, Vinegar Hill Reserve, Nov 1999, tawa, titoki, mahoe forest remnant, G.M. Coombe, 39°56.220′S, 175°38.35′E, AMNZ 34211; 1, Wharite Summit, 23 Oct 1993, J. Nunn Collection, [40°15.274′S, 175°51.454′E], NZAC 04238460. WI: 1, Bruce Park, SH1, 3-6 Feb 2000, rimu/tawa forest,J.W.Early, 39°57.555′S, 175°32.083′E, AMNZ 27647;1, Manawatu Gorge, near Woodville,Manawatu Dist., North Is, 12.12.1948, leaf mould,</p>
            <p> A. E.Brookes, 40°18.96′S, 175°48.96′E, NZAC 04069899. WA: 2, Puketoi Range, near Mount Butters, 28 Jan 2008, forest patch in gully below road, sifted leaf litter and rotten wood, K. Marske, R. Leschen, T. Buckley, 40°33.723′S, 176°3.364′E, KM271, NZAC 04234449, NZAC 04234522;2, Sutherland Vehicle Track, Aorangi Range, 24 Jan 2008, sifted wood and leaf litter in secondary forest on former grazing land, K. Marske, R. Leschen, T.Buckley, 41°25.239′S, 175°21.551′E, KM245, NZAC 03019393, NZAC 03026611; 1, Waepae Forest, Puketoi Range, 28 Jan 2008, dead wood along disused forest track, K. Marske, R. Leschen, T. Buckley, 40°33.210′S, 176°1.893′E, KM268, NZAC 03019561. WN: 1, Balance Bridge Reserve, 3 Jan 1975, litter, J.C. Watt, [40°20.244′S, 175°49.1′E], NZAC 04234429; 1, Catchpool Loop Walk, Catchpool Stream, Rimutaka Forest Park, 22 Jan 2008, sifted wood and leaf litter, K. Marske, R. Leschen, T. Buckley, 41°21.067′S, 174°55.964′E, KM233, NZAC 03020596; 1, Kaitoke, Pakuratahi Forks, 18 Jul 1993, in  Cortaderia litter, J. Nunn Collection, [41°3.374′S, 175°11.698′E], NZAC 04234613; 1, Kapiti Island, [40°51.617′S, 174°54.317′E], NZAC 03019658; 1, Karori Reservoir, 16 Oct 1994, forest leaf litter, J. Nunn Collection, [41°18.408′S, 174°44.354′E], NZAC 04234457; 1, Keith Gorge Reserve, Silverstream, 17Apr 1995, leaf litter, J. Nunn, [41°8.254′S, 174°58.561′E], NZAC 04235060; 1, Khandallah Domain, 5 Apr 1994, decayed wood, J. Nunn [41°14.721′S, 174°47.815′E], NZAC 04234450; 2, Levin,Kimberley Reserve, 6 Nov 1980, litter, C.F. Butcher, [40°58.179′S, 175°15.647′E], NZAC 04234435, NZAC 04234437; 1, Manawatu District Balance Bridge, 19 Dec 1948, leaf mould, A.E. Brookes Collection, [40°20.244′S, 175°49.1′E], NZAC 04234434; 1, Paraparaumu, Nikau Reserve, 3 Oct 1983, J. Nunn Collection, [40°54.581′S, 175°2.563′E], NZAC 04234432; 1, Ruahine Range, Wharite Peak, 19 Nov 2015, forest litter, J. Nunn, [40°15.274′S, 175°51.454′E], NZAC 04235090; 1, Tararua Forest Park 4 km along Waiotauru Road, 16 Nov 1991, J. Nunn Collection, [40°58.179′S, 175°15.647′E], NZAC 04234440; 1, Tararua Forest Park, Fields Hut, 8 Dec 1953, lichen sample, B.A. Holloway, [40°58.179′S, 175°15.647′E], NZAC 04234611; 1, Tararua Forest Park, Waiotauru Road, 25 Aug 1998, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234430; 1, Tararua Forest Park, Waiotauru Road, 4 May 1999, E.Spurr, [40°58.179′S, 175°15.647′E], NZAC 04114146; 1, Tararua Forest Park,Waiotauru Road, 11May 1999,E.Spurr,[40°58.179′S, 175°15.647′E], NZAC 04114156; 2, WN, Tararua Forest Park, Waiotauru Road, 25 May 1999, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234442, NZAC 04272533; 1, Tararua Forest Park, Waiotauru Road, 1 Jun 1999, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234615; 1, Tararua Forest Park, Waiotauru Road, 7 Jun 1999, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234438; 1, Tararua Forest Park, Waiotauru Road, 22 Jun 1999, E.Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234527;2, Tararua Forest Park,Waiotauru Road, 5 Jul 1999, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234451, NZAC 04234542; 2, Tararua Forest Park, Waiotauru Road, 20 Jul 1999, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234475, NZAC 04272534; 1, Tararua Forest Park, Waiotauru Road, 25 May 2000, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234577; 3, Tararua Forest Park, Waiotauru Road, 15 Jun 2000, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234553, NZAC 04234556, NZAC 04234632; 1, Tararua Forest Park, Waiotauru Road, 20 Jun 2000, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234483; 1, Tararua Forest Park, Waiotauru Road, 27 Jun 2000, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234609; 2, Tararua Forest Park, Waiotauru Road, 11 Jul 2000, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04114147, NZAC 04234627; 1, Forest Park, Waiotauru Road, 18 Jul 2000, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234555; 2, Tararua Forest Park, Waiotauru Road, 15 Aug 2000, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04114154, NZAC 04234528; 1, Tararua Forest Park, Waiotauru Road, 21 Aug 2000, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234541; 1, Tararua Forest Park, Waiotauru Road, 29 Aug 2000, E. Spurr, [40°58.179′S, 175°15.647′E], NZAC 04234642; 2, Tinokori Hill, 13 Jun 1991, decayed wood, J. Nunn Collection, [41°16.047′S, 174°46.630′E], NZAC 04234453, NZAC 04234517; 1, Wiltons Bush, 4 Jul 1992, J. Nunn Collection, [41°15.991′S, 174°45.164′E], NZAC 04234456. SOUTH ISLAND: SD: 1, Port Underwood Saddle, NE Picton, 10 Dec 2007, sifted leaf litter and woody debris, K. Marske, J. Allwood, [41°16.961′S, 174°6.593′E], KM094, NZAC 03018835. MB: 1 (FRNZ), Blenheim, Site BZ102 - East D. Henley 11 Jan - 7 Feb 2011, 41°22.204′S, 173°58.687′E. </p>
            <p>area on disc of pronotum; E – aedeagus, dorsal view; F – parameres, dorsal view; G – distipenis, dorsal view.</p>
            <p> Diagnosis.  Thortus amoenus is recognized by having the parasutural stria complete and the eye having 10–12 facets. It can be distinguished from  T. sulcatus by the lack of a median pronotal impression and from  T. lobatus by the wider clypeal constriction. </p>
            <p>Redescription. Length 1.17–1.50 mm. Colour of body dark to light reddish-brown, mouthparts, legs, and antennal club lighter (sometimes entire antenna). Microsculpture absent, surfaces of body semiglabrous. Dorsal setae silvery-gold, sparse, dual, consisting of very long decumbent and scattered erect and suberect setae. Ventral surfaces with shorter, suberect setae; punctation ovate, mesoventrite foveolate, central disc of metaventrite with punctures larger and more impressed than those of prosternum, punctation of abdominal ventrites 1 to 4 smaller than those on metaventrite, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head lengthened, with relatively long gena about 1.5 times length of antennomere 1; frons constricted, narrowed to a width about half the length of antennomere 1; supra-antennal ridge with well-developed rim, bead present; vertex delimited anteriorly by a transverse crenulate line, anterior surface nearly impunctate and glabrous, posterior surface with broad, shallow, foveolate punctures; gular punctation deep, ovate, diameters similar to those on side of head. Eye consisting of 10–12 facets. Antennomere 2 cylindrical, slightly wider than long, antennomere 3 subconical, longer and as wide as 2 and longer and slightly wider than 4, proportions of antennomeres 4–8 equal, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse and as wide as 11, antennomere 11 longer than 10, shorter than 9 and 10 combined. Pronotum subquadrate, pronotal length/width ratio ~1.04, widest just behind middle; anterior margin straight; anterior angles right, not protruding or acute; lateral edges more or less parallel and not gradually converging anteriorly, curved posteriorly; pronotal disc with punctures regular, uniform, separated by a distance of up to 1 times their diameter, rarely with a median glabrous strip; median impression absent; not or weakly transversely depressed at base; basomedial macropuncture absent; posterolateral angles obtuse, rounded, not projecting posteriorly; posterior margin sinuate with a weak scutellar lobe. Prosternal process with well-developed lateral marginal beads, without longitudinal depression medially, process weakly widened posteriorly behind procoxae, apex rounded, width subequal to width of procoxa, 0.46 times shorter than length of prosternum; procoxal cavity without anterolateral notch. Scutellar shield rectangular, 3.3 times wider than long. Elytra about 1.85 times as long as combined width, about 1.24 times width of pronotum at greatest width, about 2.18 times length of pronotum; setation dual, consisting of very long, decumbent setae and fewer suberect setae; humeral plica present; parasutural stria present, complete, interspace setose; punctation similar to pronotum, but separated by a distance of up to 1–3 times their diameter. Metaventrital process with anterior margin rounded, marginal bead connected at middle. Tarsi moderately slender, 5-5- 5 in female, 5-5- 4 in male, male pro- and mesotarsomeres 1–3 lacking well-defined tenent setae; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 equal to tarsomeres 1–4 combined. First abdominal ventrite with postcoxal lines acuminate and parallel, extending to mid-length of sclerite; intercoxal process rounded at apex. Aedeagus with relatively long parameres articulated to phallobase, paramere about 3 times as long as wide; apices asetose, with short, membranous extensions; basipenis about 4.73 times as long as distipenis; distipenis about 1.33 times as long as wide, lateral lobes not separated, basal plate absent; internal sac with a pair of short slender endophallites that are anteriorly widened and spatulate.</p>
            <p>Distribution. North Island: TO, GB, RI, WI, WA, WN. South Island: SD, MB (Fig. 21).</p>
            <p> Remarks.  Thortus amoenus is distributed in the southern portions of North Island. It is found in leaf litter, leaf mould, and decayed wood. Several specimens were collected in traps baited with 1080 (sodium fluoroacetate), a mammal- -specific poison (e.g., SPURR &amp; BERBEN 2004). </p>
            <p> Some specimens have a medial glabrous strip on the pronotal disc, which is also a variation present in specimens of  T. lobatus . </p>
            <p> The holotype (BMNH), collected from Makatote (Tongariro), was confirmed in LESCHEN &amp; GIMMEL (2012). A second BMNH specimen labelled by Broun with his number (3229) also had attached to it handwritten labels with the combination ‘ Pithortus  amoenus ’ and collected from Waimarino which refers to place names in the central North Island ranging from Hawkes Bay westward to Tongariro. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFC5FFC2FEA5F98B7FAAFBAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFC2FFC1FC63FBBD78CFF78F.text	3C4D87BDFFC2FFC1FC63FBBD78CFF78F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus bullerensis Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus bullerensis sp. nov.</p>
            <p>(Fig. 7)</p>
            <p> Type material.   HOLOTYPE: ♁ (NZAC), labelled: ‘Dublin Terrace Buller 25.11.61 leaf litter J. I. Townsend. [handwritten] //  Thortus sp. nov. [handwritten] det. R. A. Crowson // Entomology Div. D.S.I.R. New Zealand / NZ Arthropod Collection Private Bag 92170 Auckland New Zealand NZAC04235080  .’ PARATYPE (NZAC). SOUTH ISLAND: BR: Norris Creek, Buller Road, 14 Oct 1970, litter, J.I. Townsend, [42°6.245′S, 172°12.171′E], NZAC 04272538. </p>
            <p> Diagnosis.  Thortus bullerensis sp. nov. is a unicolourous species with a well-developed eye, a narrowed frons, the gena short, the pronotum widest in the anterior 1/3, and the parasutural stria incomplete. It is most similar to  T. parallelus , which has the pronotum widest at the middle and the marginal bead of the metaventrital process present. Description. Length 1.35 mm. Colour of body unicolourous red-brown, with lighter antennal club, mouthparts and legs. Body surface semiglabrous, microsculpture absent from most surfaces, very weak and present on ventral surfaces of the head, hypomeron and lateral portions of the abdominal ventrites. Dorsal setae silver, consisting of short scattered suberect and subdecumbent setae. Ventral surfaces with suberect setae; mesoventrite foveolate, not strongly impressed, disc of metaventrite with punctures variable, some more impressed than those on prosternum, punctation of abdominal ventrites 1 to 4 generally smaller or equal to those on metaventrite, setae longer at sides, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head lengthened with short gena, subequal to length of antennomere 1; frons constricted, narrowed to width just less than half the length of antennomere 1; supra-antennal ridge with well-developed rim, bead present; vertex delimited anteriorly by a weakly transverse crenulate line, anterior surface glabrous, posterior surface with broad foveolate punctures; gular punctation deep and ovate, diameters larger than those on sides of head. Eye consisting of approximately 13 facets. Antennomere 2 barrel-shaped, wider than long, antennomere 3 subconical, longer and narrower than 2 and longer and slightly wider than 4, proportions of antennomeres 4–8 equal, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse same width as 11, antennomere 11 longer than 10, shorter than 9 and 10 combined. Pronotum transverse, pronotal length/width ratio ~0.81, widest at apical third; anterior margin convex; anterior angles obtuse, not projecting and rounded; lateral edges weakly convex, weakly curved anteriorly, gradually converging posteriorly; pronotal disc with punctation subuniform and foveolate, shallow, absent in central disc with a broad median glabrous area, separated by a distance of up to 1–2 times their diameter and larger at the base; median impression absent; transversely depressed at basal 1/5; basomedial macropuncture present, triangulate; posterolateral angles almost right, sharp, slightly projecting posteriorly; posterior margin sinuate with a weak scutellar lobe. Prosternal process with well-developed lateral marginal beads, without longitudinal depression medially, process subparallel-sided and slightly converging posteriorly, slightly expanded posteriorly behind procoxae, apex curved, width slightly narrower than procoxa and about half total prosternal length; procoxal cavity without anterolateral notch. Scutellary shield trapezoidal and transverse, 2.1 times wider than long. Elytra about 1.37 times as long as combined width, about 1.11 times width of pronotum at greatest width, about 1.90 times length of pronotum; setation dual consisting of relatively short suberect and shorter subdecumbent setae; humeral plica present; parasutural stria present and incomplete, present to about basal 1/3; punctation less dense than pronotum, except at base where it is more impressed. Metaventrital process with anterior margin straight, marginal bead present at sides, absent at middle. Tarsi moderately slender, 5-5- 5 in female, 5-5- 4 in male, tenent setae not observed; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 subequal to tarsomeres 1–4 combined. First abdominal ventrite with broad rounded intercoxal process, postcoxal lines long, divergent, extending middle of sclerite. Aedeagus with relatively short parameres that are fused at their bases and fixed to phallobase, paramere about as wide as long; apices asetose (?), without membranous extensions; basipenis about 2.5 times length of distipenis; distipenis about 1.60 times longer than width, lateral lobes separated, basal plate absent; internal sac with a pair of short slender endophallites. </p>
            <p>Etymology. The specific epithet is taken from the Buller region; adjective.</p>
            <p>Distribution. South Island: BR (Fig. 21).</p>
            <p> Remarks.  Thortus bullerensis sp. nov. is endemic to the Buller region of the South Island. It was collected from leaf litter. The two specimens were completely disarticulated. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFC2FFC1FC63FBBD78CFF78F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFC1FFDFFC6DFF5E7FAAF784.text	3C4D87BDFFC1FFDFFC6DFF5E7FAAF784.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus crowsoni Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus crowsoni sp. nov.</p>
            <p>(Fig. 8)</p>
            <p> Type material.   HOLOTYPE: ♀ (NZAC, card-mounted after dissection), labelled: ‘NZ: S. Canterbury,  Geraldine , Peel For. Nat. Res. under bark of Kanikoreas matais, 27. xii. 1956, SC2 R. Crowson // Entomology Div. D.S.I.R. New Zealand // NZ Arthropod Collection Private Bag 92170 Auckland New Zealand NZAC04235098.’ </p>
            <p> Diagnosis.  Thortus crowsoni sp. nov. is a bicolourous species with a unifacetted eye and a broad body. It is most similar to  T. ovalis but has well-developed punctures on the pronotum and the head and has a broader clypeal constriction, and lack of a fine, transverse line on the vertex. Description. Length 1.70 mm. Colour of body dark reddish-brown, with lighter elytra, antennal club, mouthparts and legs. Body surfaces semiglabrous, microsculpture absent. Dorsal setae silver, consisting of short scattered recurved setae. Ventral surfaces with suberect setae; punctation ovate on head and prosternum, mesoventrite foveolate, not strongly impressed, disc of metaventrite with ovate punctures that are slightly less impressed than those on prosternum, punctation of abdominal ventrites 1 to 4 generally smaller or equal to those on metaventrite, weak, setae not longer at sides, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head not lengthened with relatively short gena, with length narrowed subequal to length of antennomere 1; frons wide and not constricted, narrowed to width to about equal in length to antennomere 1; supra-antennal ridge with well-developed rim, bead absent; vertex not delimited anteriorly by a weak transverse crenulate line, surface rugulose with deep punctures of mixed sizes, two rows of weak transverse rows, the largest fused with their anterior rims connected and forming a crenulate line; gular punctation deep and ovate, diameters less than those on sides of head. Eye unifacetted. Antennomere 2 barrel-shaped, slightly wider than long, antennomere 3 subconical, longer and narrower than 2 and longer and slightly wider than 4, proportions of antennomeres 4–8 nearly the same, antennomere 9 slightly wider than 8 and narrower than 10 and 11, antennomere 10 not transverse and about the same width as 11, antennomere 11 longer than 10, shorter than 9 and 10 combined. Pronotum transverse, pronotal length/width ratio ~0.86, widest near middle; anterior margin weakly convex; anterior angles obtuse, not projecting and rounded; lateral edges weakly convex, curved anteriorly at anterior 1/3, gradually converging posteriorly; pronotal disc with punctation not uniform and foveolate, enlarged towards base, shallow, present in central disc (without a median glabrous strip), separated by a distance of up to 1 times their diameter; median impression absent; transversely depressed at basal 1/5; basomedial macropuncture absent; posterolateral angles almost right, sharp, slightly projecting posteriorly; posterior margin sinuate with a scutellar lobe. Prosternal process with well-developed lateral marginal beads, with longitudinal depression medially, process subparallel- -sided, slightly expanded posteriorly behind procoxae, apex curved, width slightly narrower than procoxa and 0.5 times length of prosternum; procoxal cavity without anterolateral notch. Scutellary shield subtrapezoidal and transverse, 1.7 times as wide as long. Elytra about 1.20 times as long as combined width, about 1.14 times greatest width of pronotum, about 1.58 times length of pronotum; setation uniform consisting of relatively short, curved setae; humeral plica absent; parasutural stria absent; punctation present, sparse and not as impressed as that on pronotum. Metaventrital process with anterior margin straight, marginal bead present at sides, absent at middle. Tarsi moderately slender, 5-5- 5 in female, male unknown; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 slightly longer than tarsomeres 1–4 combined. First abdominal ventrite with broad rounded intercoxal process, postcoxal lines long, weakly convex, extending middle of sclerite. </p>
            <p> Etymology. The specific epithet is a patronym for Roy Crowson, collector of the holotype, and modernist reviser of  Coleoptera classification. The specimen, along with others in yellow-aged spirits, was gifted to the RABL during his first visit to Glasgow in 1992. </p>
            <p>Distribution. South Island: SC (Fig. 21).</p>
            <p> Remarks.  Thortus crowsoni sp. nov. is known from a single specimen collected with specimens of  Thortus ovalis . The misspelling on the label may refer either to kahikatea (  Dacrycarpus dacrydioides (A.Rich.) de Laub. ) or matai (  Prumnopitys taxifolia (Banks &amp; Sol. ex D. Don) de Laub. ). </p>
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	https://treatment.plazi.org/id/3C4D87BDFFC1FFDFFC6DFF5E7FAAF784	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFDEFFDEFE8AFF5E7FAAFB6F.text	3C4D87BDFFDEFFDEFE8AFF5E7FAAFB6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus helmorei Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus helmorei sp. nov.</p>
            <p>(Figs 9, 19D)</p>
            <p> Type material.   HOLOTYPE: ♁ (AMNZ), labelled: ‘NEW ZEALAND, ND  Waipoua Forest , 10.i.1967, R. Rowe. By giant kauri. Litter. L 5787 // AMNZ36232  ’. PARATYPES (4; AMNZ, ANIC, NZAC). NORTH ISLAND: ND: 1 (ANIC), Waipoua SF, Yakas Tree Tk., 350 m, 11-14.iv.1980, broadlf.-podocarp,A. Newton, M.Thayer [35°36.99′S 173°31.91′E], litter at bases of  Metrosideros robusta trunks; 1, Waipoua Forest, 10 Jan 1967, litter by giant kauri, R.J. Rowe, 35°37′S, 173°33.067′E, AMNZ 36233; 2, Waipoua State Forest, SH12, 20 Sep 1977, rotten stumps trees, D.W. Helmore, [35°38.772′S, 173°33.321′E], NZAC 04235069, NZAC 04272548. </p>
            <p> Diagnosis.  Thortus helmorei sp. nov. is a unicolourous species with the eye composed of a single facet. The only known Northland species of the genus, it can be distinguished from other unifacetted species by the absence of abdominal postcoxal lines. </p>
            <p>Description. Length 1.70–1.75 mm. Colour of body unicolourous red-brown, with lighter antennal club, mouthparts and legs. Body surface glabrous, microsculpture absent from most surfaces, very weak and present on ventral surfaces of the head, hypomeron and lateral portions of the abdominal ventrites. Dorsal setae golden, consisting of short slightly curved and suberect setae. Ventral surfaces with short and suberect setae; punctation ovate on head (more impressed than on frons) and prosternum, mesoventrite foveolate, strongly impressed, disc of metaventrite with punctures variable, more closely spaced than those on the central disc of the prosternum, punctation of abdominal ventrites 1 to 4 generally smaller or equal to those on metaventrite, setae longer at sides, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head not lengthened with a relatively short gena that is subequal in length to antennomere 1; frons not greatly constricted, narrowed to a width equal to length of antennomere 1; supra-antennal ridge with well-developed rim, bead absent; vertex delimited anteriorly by a transverse crenulate line, anterior surface punctured, posterior surface glabrous with small punctures posteriorly; gular punctation deep and ovate, diameters about equal to those on sides of head. Eye consisting of a single facet. Antennomere 2 barrel-shaped, wider than long, antennomere 3 subconical, equal in length and narrower than 2 and longer and as wide than 4, proportions of antennomeres 4–8 about the same, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse and about the same width as 11, antennomere 11 longer than 10, subequal to 9 and 10 combined. Pronotum transverse, pronotal length/width ratio ~0.92, widest at apical third; anterior margin convex; anterior angles obtuse, not projecting and rounded; lateral edges weakly convex, weakly curved anteriorly in anterior 1/4, gradually and weakly converging posteriorly; pronotal disc with punctation more or less uniform and more larger and foveolate towards base, shallow to sparse to weak in central disc (without a median glabrous strip), separated by a distance of up to 3 times their diameter at anterolateral areas and larger and closer at base; median impression absent; transversely depressed at basal 1/5; basomedial macropuncture vaguely indicated; posterolateral angles obtuse and rounded, not projecting posteriorly; posterior margin sinuate with scutellar lobe. Prosternal process with weak lateral marginal beads, without strong longitudinal depression medially, process subparallel-sided and slightly expanded posteriorly behind procoxae, apex rounded, width slightly narrower than procoxa, 0.5 times as wide as prosternum; procoxal cavity with weak anterolateral notch. Scutellary shield suboval and transverse, 1.9 times as wide as long. Elytra about 1.27 times as long as combined widths, about 1.11 times as greatest width of pronotum, about 1.54 times longer than length of pronotum; setation uniform consisting of relatively short slightly curved setae; humeral plica absent; parasutural stria absent; punctation less dense than pronotum, impressed near base. Metaventrital process with anterior margin curved, marginal bead weakly indicated at middle. Tarsi moderately slender, 5-5- 5 in female, 5-5- 4 in male, male protarsomeres 1–3 and mesotarsomere 1 with tenent setae; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 subequal to tarsomeres 1–4 combined. First abdominal ventrite with broad rounded intercoxal process, postcoxal lines absent. Aedeagus with relatively moderately long parameres weakly articulated to phallobase, paramere about 2 times as long as wide; apices more or less acute with two microsetae, membranous extensions absent; basipenis about 2.29 times as long as distipenis; distipenis about 1.90 times as long as wide, lateral lobes not well separated, basal plate absent; internal sac (?) with a single endophallite.</p>
            <p>Etymology. Rarely into the field but quietly working at his bench, Des Helmore, artist, former illustrator at the NZAC and friend of the first author, is honored with the specific epithet.</p>
            <p>Distribution. South Island: ND (Fig. 21).</p>
            <p> Remarks.  Thortus helmorei sp. nov. is known from five specimens, two collected from a rotten tree stump and litter at the base of a  Metrosideros robusta A.Cunn. trunk. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFDEFFDEFE8AFF5E7FAAFB6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFDEFFDDFC44FAFF7F31FE8F.text	3C4D87BDFFDEFFDDFC44FAFF7F31FE8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus latus Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus latus sp. nov.</p>
            <p>(Fig. 10)</p>
            <p>
                 Type material.   HOLOTYPE: ♀ (NZAC), labelled: ‘NEW ZEALAND,NN,  
                <a title="Search Plazi for locations around (long 172.73753/lat -41.1908)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=172.73753&amp;materialsCitation.latitude=-41.1908">Mount Arthur</a>
                 , 13 Jan 2020, sifted dead wood, Y. Chen, R. Leschen, H. Lindsay, 41°11.448′S, 172°44.252′E, RL2270 // NZ Arthropod Collection Private Bag 92170 Auckland New Zealand NZAC04272556  .’ PARATYPE (NZAC). SOUTH ISLAND: NN: 1, Kaituna,Aorere, 6 May 1965, litter, G. Kuschel, [40°42.900′S, 172°35.457′E], NZAC 04235091. 
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            <p> Diagnosis.  Thortus latus sp. nov. is a unicolourous species with a short gena, a weakly indicated or absent parasutural stria and a notch at the lateral edges of the supra-antennal ridge. It can be recognized from similar-looking species by its broad body and eye composed of 6 or 7 facets. </p>
            <p>Description. Length 1.25 mm. Colour of body unicolourous red-brown, with lighter antennal club, mouthparts and legs. Body surface glabrous, microsculpture absent. Dorsal setae silver, consisting of sparse, short suberect curved setae. Ventral surfaces with sparse suberect curved setae; punctation ovate on prosternum, mesoventrite weakly foveolate, not strongly impressed, disc of metaventrite with punctures variable, about as impressed as those on prosternum, punctation of abdominal ventrites 1 to 4 generally weaker than on metaventrite, setae slightly longer at sides, punctation on ventrite 5 denser with a patch of posteriorly- -directed setae. Head not lengthened with a relatively short gena that is subequal to the length of antennomere 1; frons moderately constricted, narrowed to a width equal in length to antennomere 1; supra-antennal ridge with well-developed rim, terminating at a notch laterad, bead absent; vertex delimited anteriorly by a transverse crenulate line, anterior surface punctured, posterior surface glabrous virtually impunctate; gular punctation irregular and ovate, diameters larger than those on sides of head. Eye consisting of 6 or 7 facets. Antennomere 2 barrel-shaped, wider than long, antennomere 3 subconical, as long as and slightly narrower than 2 and longer and slightly wider than 4, proportions of antennomeres 4–8 same, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse and about the same width as 11, antennomere 11 longer than 10, slightly shorter than 9 and 10 combined. Pronotum transverse, pronotal length/width ratio ~0.84, widest at middle; anterior margin convex; anterior angles obtuse, not projecting and rounded; lateral edges weakly convex, weakly curved anteriorly from middle, parallel-sided at basal half; pronotal disc with punctation subuniform and foveolate, shallow, sparse (without a median glabrous strip), separated by a distance of up to 1–2 times their diameter and larger at the base; median impression absent; transversely depressed at basal 1/5; basomedial macropuncture weakly indicated; posterolateral angles almost right, sharp, slightly projecting posteriorly; posterior margin sinuate with scutellar lobe. Prosternal process with well-developed lateral marginal beads, without longitudinal depression medially, process subparallel-sided and slightly converging posteriorly, not expanded posteriorly behind procoxae, apex curved, width wider than procoxa and 1.75 times longer than length of prosternum; procoxal cavity without anterolateral notch. Scutellary shield trapezoidal and transverse, 1.8 times as wide as long. Elytra about 1.28 times as long as combined widths, about 1.11 times as wide as width of pronotum, about 1.09 times longer than length of pronotum; setation uniform consisting of relatively short suberect and curved setae; humeral plica present; parasutural stria incomplete, barely visible, punctation less dense than pronotum, but at base slightly more impressed. Metaventrital process with anterior margin straight. Tarsi moderately slender, female 5-5-5, 5-5- 4 in male, tenent setae not present, but long curled setae present on pro- and mesotarsomeres 1–4 of both sexes; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 subequal to combined lengths of tarsomeres 1–4. First abdominal ventrite with broad rounded intercoxal process, postcoxal lines long, weakly convergent, extending middle of sclerite. Aedeagus with relatively long parameres articulated to phallobase, paramere about 3 times longer than wide; apices not expanded and subacute; basipenis about 2.87 times as long as distipenis; distipenis about 1.33 times as long as wide, lateral lobes not separated, basal plate absent; internal sac (?) with a single, elongate endophallite.</p>
            <p> Etymology. The specific epithet is the Latin adjective  ‘ latus ’, meaning ‘broad’, in reference to the distinctive body form. </p>
            <p> Remarks.  Thortus latus sp. nov. is known from a single specimen collected from forest litter. The aedeagus was damaged during dissection. </p>
            <p>Distribution. South Island: NN (Fig. 21).</p>
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	https://treatment.plazi.org/id/3C4D87BDFFDEFFDDFC44FAFF7F31FE8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFDDFFDAFC57FED678C2FB2F.text	3C4D87BDFFDDFFDAFC57FED678C2FB2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus lobatus Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus lobatus sp. nov.</p>
            <p>(Fig. 11)</p>
            <p>
                 Type material.   HOLOTYPE: ♁ (NZAC), labelled: ‘ NEW ZEALAND, WD Ship Cr 6 Nov 2007 R. Leschen C. Carlton RL1299 (RL007) // leaf litter and rotten log berlesate 43 45.556S 169 8.986E // NZ Arthropod Collection Private Bag 92170 Auckland  
                <a title="Search Plazi for locations around (long 169.14977/lat -43.759266)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=169.14977&amp;materialsCitation.latitude=-43.759266">New Zealand</a>
                 NZAC04235171  ’. PARATYPES (21; NZAC): NORTH ISLAND: TK: 1, Egmont National Park, Waiwhakaiho Track, 16 Jan 2008, sifted leaf litter, K. Marske, R. Leschen, T. Buckley, 39°13.371′S, 174°7.444′E, KM180, NZAC 04272537; 1, Pouakai Range, 9-13 Jan 1978, J.C. Watt, [39°14.451′S, 174°1.232′E], NZAC 04235150. SOUTH ISLAND: NN: 1, Collingwood, Kaituna,Aorere Valley, 3 Apr 1964, litter, J.I. Townsend, [40°43.442′S, 172°35.915′E], NZAC 04235034; 1, Karamea Saddle, 13 Oct 1970, litter, J.I. Townsend, [41°30.981′S, 172°1.229′E], NZAC 04235301. BR: 1, Punakaiki, Pahautane Bay, 9 June 1983, H.P. McColl, [42°6.317′S 171°20.183′E], NZAC 04272536; 1, Punakaiki, Pororari River Walk, 27 Aug 2010, washed soil sample supplejack/nikau forest, J. Nunn Collection, [42°6.431′S, 171°20.413′E], NZAC 04235172. WD: 2, Open Bay Islands, Taumaka Island, 30 Apr 1976, litter mainly  Schefflera digitalis ,  Melicytus ramiflorus, R.H. Matlin , [43°51.644′S, 168°52.95′E], NZAC 04235169, NZAC 04272547. OL: 1, Haast Pass, Found Creek Culvert, 6 Mar 2003, sifted leaf litter and rotting logs, R. Leschen, C. McGuiness, 44°3.539′S, 169°22.778′E, RL785, NZAC 04235300. FD: 1, Bauza Island, 3 March 2010, sifted litter/rotten logs, S.A. Forgie, 45°17.768′S, 166°55.82′E, SAF030, NZAC 03027041; 1, Dusky Sound, Gilbert Islands, 25 Jan 2011, sifted litter/bark/lichen/moss/log scrapings, S.A. Forgie, 45°36.047′S, 166°39.595′E, SAF042, NZAC 03027121; 1, Eglington Valley, Cascade Creek, 21 Jan 1962, litter, J.I. Townsend, [44°53.694′S, 168°4.938′E], NZAC 04235081; 1, Routeburn Track, 11 May 200, ex Calostoma, R. Leschen, 44°49′S, 168°7′E, RL574, NZAC 04235142; 3, Secretary Island, Grono Bay, 24 Mar 1984, sifted litter, C.F. Butcher, [45°17.044′S, 166°56.321′E], NZAC 04235073, NZAC 04235078, NZAC 04235135; 2, Secretary Island, ridge towards Mount Grono, 30 Nov 1981, mats and mosses, C.F. Butcher, [45°16.553′S, 166°56.488′E], NZAC 04235111, NZAC 04235141; 2, Secretary Island, ridge towards Mount Grono, 30 Nov 1981, alpine mats, moss and tussock, C.F. Butcher, [45°16.553′S, 166°56.488′E], NZAC 04235059, NZAC 04235137; 1, Wilmot Pass, Jan 1970, mats, J.S. Dugdale, [45°30.482′S, 167°11.553′E], NZAC 04235082. 
            </p>
            <p>Additional material. SOUTH ISLAND: SD: 1, Port Liger, Marlborough, 26 Oct 1969, litter, F.Alack, 40°54.586′S, 173°58.688′E, NZAC 04235115.</p>
            <p> Diagnosis.  Thortus lobatus sp. nov. can be distinguished from most species by having complete parasutural striae and a relatively long gena. It is most similar to  T. amoenus and  T. sulcatus sp. nov. ; it can be distinguished from  T. amoenus by the narrow clypeus and from  T. sulcatus sp. nov. by lacking a median pronotal impression. </p>
            <p>Description. Length 1.20–1.55 mm. Colour of body dark reddish-brown, mouthparts, legs, and antennal club lighter. Body surface semiglabrous, microsculpture absent, apart from posterior portion of gula. Dorsal setae silvery-gold, consisting of very long decumbent and scattered suberect to erect and subdecumbent curved and straight setae. Ventral surfaces with shorter subdecumbent straight and slightly curved setae; punctation ovate, well-impressed, and not uniform in size, mesoventrite foveolate, central disc of metaventrite with punctures of different sizes, with larger ovate punctures present anteriorly and those on central disc smaller than those on prosternum, punctation of abdominal ventrites 1 to 4 about as large as those on the central disc of metaventrite, with those on ventrites 1–5 generally more closely packed and more shallower on ventrites 4 and 5, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head lengthened with relatively long gena, about 1.5 times as long as length of antennomere 1; frons constricted, narrowed to a width about 2/3 the length of antennomere 1; supra-antennal ridge with well-developed rim, bead present, well developed and relatively broad; vertex not delimited anteriorly by a transverse crenulate line, surfaces punctate to foveolate (punctures larger in a specimen from Taranaki); gular punctation irregular, deep and ovate, diameters similar to those on side of head. Eye consisting of about 11–18 facets. Antennomere 2 cylindrical, slightly longer than wide, antennomere 3 subconical, longer and about as wide as 2 and longer and about as wide as 4, proportions of antennomeres 4–8 same, antennomere 9 wider than 8 and slightly narrower than 10 and 11, antennomere 10 transverse and slightly wider than 11, antennomere 11 longer than 10, shorter than 9 and 10 combined. Pronotum subquadrate, pronotal length/width ratio ~1.03, widest at middle; anterior margin slightly curved; anterior angles obtuse and rounded; lateral edges somewhat evenly rounded and gradually converging anteriorly and posteriorly; pronotal disc with punctures more or less regular and uniform and at base weaker or absent at middle and more strongly impressed along posterior margin, separated by a distance of 0.5–1.0 times their diameter; median impression absent; not or weakly transversely depressed at base; basomedial macropuncture absent; posterolateral angles obtuse and rounded, not projecting posteriorly; posterior margin sinuate with weak scutellar lobe. Prosternal process with well-developed lateral marginal beads, without longitudinal depression medially, process weakly widened posteriorly behind procoxae, apex subacute, width 0.48 times shorter than length of prosternum; procoxal cavity without anterolateral notch. Scutellary shield ovoid, 1.77 times as wide as long. Elytra about 1.50 times as long as combined widths, about 1.17 times wider than width of pronotum, about 1.70 times longer than length of pronotum; setation dual, consisting of very long straight and curved setae; humeral plica present; parasutural stria present and complete, interspace setose; punctation dissimilar to pronotum, more impressed, larger and dense at base, separated by a distance of up to 1–4 times their diameter. Metaventrital process with anterior margin straight, marginal bead absent. Tarsi moderately slender, 5-5- 5 in female, 5-5- 4 in male, male pro- and mesotarsomeres 1 and 2 with tenent setae; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 4 slightly longer than tarsomeres 1–3 combined. First abdominal ventrite with postcoxal lines weakly convex, extending slightly beyond mid-length of sclerite; intercoxal process somewhat narrowed, rounded at apex. Aedeagus with long and slender parameres articulated to phallobase; paramere about 3 times as long as wide, apices asetose with long membranous extensions; basipenis about 3.92 times as long as distipenis; distipenis about 1.61 times as long as wide, lateral lobes not separated, basal plate absent; internal sac (?) with at least one slender endophallite.</p>
            <p> Etymology. The specific epithet is the Latin adjective  ‘ lobatus ’, meaning ‘lobate’, in reference to the modified apex of the parameres. </p>
            <p>Distribution. North Island: TK. South Island: NN,?SD, WD, FD (Fig. 21).</p>
            <p> Remarks.  Thortus lobatus sp. nov. is known from several specimens, including one from the North Island, Taranaki, with the rest from the western South Island. A single damaged specimen excluded from the type series with missing elytra from Port Ligar (Fig. 22C) may also be attributed  T. lobatus sp. nov. based on the form of the frons and the shape of the pronotum. This specimen locality is shown in Fig. 24 as ‘  Thortus nr.  lobatus ’. Specimens have been taken mainly by sifting moss, leaf litter and rotten wood. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFDDFFDAFC57FED678C2FB2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFDAFFD7FEF8FB3D781BFD6F.text	3C4D87BDFFDAFFD7FEF8FB3D781BFD6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus luscus Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus luscus sp. nov.</p>
            <p>(Fig. 12)</p>
            <p> Type material.  HOLOTYPE: ♁ (LUNZ), labelled: ‘NZ: 38°S 175.58°E MauRes, F:-; P:- (3363ha) #W651-013, 33cm diam litter: 72hr Berlese, Edge dist 34.7m (9.2m offset), RK Didham 19-Feb-08 .’ PARATYPES (8; AMNZ, FRNZ, LUNZ, NZAC). NORTH ISLAND: WO: 1 (LUNZ), Mangatautari, 20 Nov - 17 Dec 2008, south cell outside cell, C.H. Watts, [38°2.679′S, 175°33.832′E]; 1, Mangatautari, 21 Jan - 24 Feb 2008, south cell outside cell, C.H. Watts, [38°2.679′S, 175°33.832′E], NZAC 04272535; 1, Maungatautari, 18 Dec 2010 - 20 Jan 2011, C.H. Watts, 37°58.133′, 175°33.683′E, AMNZ 86330;1 (LUNZ), Maungatautari Mountain, 18 Dec 2010 - 20 Jan 2011, C.H. Watts, 38°0′S, 175°34.8′E; 1 (LUNZ, same as holotype), Mau Reserve Forest Park (Mangatautari), 19 Feb 2008, litter, R.K. Didham, 38°0′S, 175°34.8′E; 1, Sanctuary Mountain, Maungatautari, southern enclosure, 20 Nov 2020, litter, R. Leschen, Y. Chen, 38°2.679′S, 175°33.832′E, RL2265, NZAC 03038714. BP: 1 (FRNZ), Dansey [Road Scenic] Reserve, pitfall trap OP5, C.Wardhaugh, 9 Jan - 3 Feb 2023, 38°5.417′S, 176°7.45′E; 1 (FRNZ), Endean′ s Bush native forest pitfall 1, C. Wardhaugh, 13 Dec 2022 - 9 Feb 2023, 38°9.871′S, 176°1.622′E. </p>
            <p> Diagnosis.  Thortus luscus sp. nov. is unicolourous and has a unifacetted eye. The abdominal postcoxal lines are divergent, but do not reach the middle of the disc of abdominal ventrite 1. It most closely resembles  T. simplex , differing from it by having the pronotum widest at the middle and by having parasutural striae. </p>
            <p>Description. Length 1.30–1.40 mm. Colour of body unicolourous red-brown, with lighter antennal club, mouthparts and legs. Body surface glabrous, microsculpture present on ventral surfaces and distinct on the abdominal ventrites, absent or very weak and present on lateral surfaces of the head and at the sides of the pronotum. Dorsal setae silver, consisting of short scattered decumbent setae. Ventral surfaces with suberect setae; punctation ovate on prosternum, mesoventrite foveolate and strongly impressed, disc of metaventrite with punctures variable and as impressed as those on prosternum, but generally smaller, punctation of abdominal ventrites 1 to 4 smaller or equal to those on metaventrite and lass impressed, setae not longer at sides, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head not lengthened with relatively short gena that is subequal to the length of antennomere 1; frons constricted, narrowed to a width subequal in length to antennomere 1; supra-antennal ridge with well-developed rim, bead absent; vertex delimited anteriorly by a transverse crenulate line, anterior surface punctured, posterior surface subglabrous and irregularly foveolate; gular punctation semi-regular and ovate, diameters more or less equal to those on sides of head. Eye consisting of a single facet. Antennomere 2 barrel-shaped, slighter longer than wide, antennomere 3 subconical, about as long as slightly narrower than 2 and longer and wider than 4, proportions of antennomeres 4–8 nearly the same, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse and about the same width as 11, antennomere 11 longer than 10, shorter than 9 and 10 combined. Pronotum transverse, pronotal length/width ratio ~1.16, widest at middle; anterior margin convex; anterior angles obtuse, rounded and not projecting; lateral edges weakly convex, converging anteriorly, gradually converging posteriorly; pronotal disc with punctation coarse and subuniform and foveolate, shallow, sparse to absent in central disc, separated by a distance of up to 1 times their diameter and larger at the base; median impression absent; transversely depressed at basal 1/5; basomedial macropuncture present but weakly indicated and elongate and may be weakly triangulate; posterolateral angles right, sharp, not or slightly projecting posteriorly; posterior margin sinuate with a weak scutellar lobe. Prosternal process with well-developed lateral marginal beads, without longitudinal depression medially, process more or less subparallel-sided, slightly expanded posteriorly behind procoxae, apex subacute, width slightly narrower than procoxa 0.5 times as long as length of prosternum; procoxal cavity without anterolateral notch. Scutellary shield subtrapezoidal and transverse, 2.3 times as wide as long. Elytra about 1.31 times as long as combined widths, about 1.11 times wider than width of pronotum, about 1.70 times longer than length of pronotum; setation uniform consisting of relatively short decumbent setae; humeral plica present; parasutural stria present, incomplete and extending to about basal 1/3; punctation less dense than pronotum, but at base more impressed. Metaventrital process with anterior margin weakly curved, marginal bead weakly indicated at middle. Tarsi moderately slender, 5-5- 5 in female, 5-5- 4 in male, male pro- and mesotarsomeres 1–3 with tenent setae; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 subequal to tarsomeres 1–4 combined. First abdominal ventrite with intercoxal process relatively narrowed and subacute, postcoxal lines relatively short, divergent, extending to basal 1/3 rd of sclerite. Aedeagus with relatively long parameres articulated to phallobase, paramere about 2 times as long as wide; apices somewhat truncate or subrounded, without membranous projections, unisetose; basipenis about 3.43 times as long as distipenis; distipenis about 1.76 times as long as wide, lateral lobes separated, basal plate absent; internal sac with a pair of long slender endophallites, one, at least is expanded apically.</p>
            <p> Etymology. The specific epithet is the Latin adjective  ‘ luscus ’, meaning ‘one-eyed’. </p>
            <p>Distribution. North Island: WO, BP (Fig. 21).</p>
            <p> Remarks.  Thortus luscus sp. nov. is known from nine specimens, collected at Maungatautari, Endean’s Bush, and Dansey Road Scenic Reserve. It has been taken in pitfall traps and sifting litter and dead wood. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFDAFFD7FEF8FB3D781BFD6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFD7FFD6FE87FCFF78B9FB4F.text	3C4D87BDFFD7FFD6FE87FCFF78B9FB4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus michauxi Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus michauxi sp. nov.</p>
            <p>(Figs 13, 19B)</p>
            <p> Type material.   HOLOTYPE: ♁ (NZAC), labelled: ‘ Ewing I. Auckland Is. 9 Feb 73 D. S. Horning // ex mat of thick moss on prostrate trunk of  Olearia lyalli // NZ Arthropod Collection Private Bag 92170 Auckland New Zealand NZAC04235028  .’ PARATYPES (80; AMNZ, BPMN, NZAC, OMNZ). AUCKLAND ISLANDS (AU): 2,Adams Island, 20 Jan - 4 Feb 2011, tussock and megaherbs, in flat albatross nestling area, G. Elliott, K.J. Walker, 50°54.158′S, 166°0.262′E, AMNZ 85286, AMNZ 85287;1,Adams Island, 19 Jan - 3 Feb 2011, southern rata forest coastal, sooty shearwater colony, G. Elliott, K.J. Walker, 50°51.905′S,166°22.117′, AMNZ 85292; 1 (OMNZ), Adams Island, 2-7 Dec 1996, G.H. Sherley, B. Patrick, [50°50.248′S, 165°55.342′E]; 1, Adams Island, Fairchilds Garden, 20 Jan 1966, litter, G. Kuschel, 50°50.248′S, 165°55.342′E, NZAC 04235277; 2, Adams Island, Fairchilds Garden, 22 Mar 2006, megaherb leaf litter, R. Leschen, E. Edwards, 50°50.248′S, 165°55.342′E, NZAC 03019027, NZAC 03021584; 1, Adams Island,Magnetic Point, 28 Nov - 14 Dec 1996, herbfield, G.H. Sherley, [50°50.248′S, 165°55.342′E], NZAC 04235197; 21, Adams Island, Magnetic Cove Station, 28 Jan 1966, litter, G. Kuschel, [50°51.645′S, 166°5.302′E], NZAC 04231310, NZAC 04235064, NZAC 04235068, NZAC 04235112, NZAC 04235158, NZAC 04235160, NZAC 04235161, NZAC 04235163, NZAC 04235210, NZAC 04235238, NZAC 04235239, NZAC 04235245, NZAC 04235265, NZAC 04235299, NZAC 04235305, NZAC 04235313, NZAC 04235314, NZAC 04272528, NZAC 04272540, NZAC 04272541, NZAC 04272542; 1, Carnley Harbour, E Tagua Bay, 4 Feb 1973, litter, J.S. Dugdale, [50°48.633′S, 166°3.933′E], NZAC 04235268; 7, Auckland Islands, Disappointment Island, 15 Feb 1975, plants,D.S.Horning, [50°36.45′S, 165°58.283′E], NZAC 04235186, NZAC 04235212, NZAC 04235221 NZAC 04235224, NZAC 04235228, NZAC 04235310, NZAC 04235318; 18, Ewing Island, 9 Feb 1973, in bark of live  Olearia lyalli, D.S. Horning , [50°31.711′S, 166°18.215′E], NZAC 04235032, NZAC 04235039, NZAC 04235040, NZAC 04235055, NZAC 04235061, NZAC 04235067, NZAC 04235070, NZAC 04235071, NZAC 04235149, NZAC 04235162, NZAC 04235164, NZAC 04235167, NZAC 04235170, NZAC 04235247, NZAC 04235289, NZAC 04235290, NZAC 04235295, NZAC 04235319; 13 (same as holotype), Ewing Island, 9 Feb 1973, ex mat of thick moss on prostrate trunk of  Olearia layallii, D.S. Horning , [50°31.713′S, 166°18.229′E], NZAC 04235028, NZAC 04235031, NZAC 04235083, NZAC 04235084, NZAC 04235101, NZAC 04235162, NZAC 04235180, NZAC 04235183, NZAC 04235195, NZAC 04235200, NZAC 04235219, NZAC 04235256, NZAC 04235272, NZAC 04235275; 4, Ewing Island, 7 Feb 1973, in rotting bark of live  Olearia lyallii, D.S.Horning ,[50°31.713′S, 166°18.229′E], NZAC 04235037, NZAC 04235077, NZAC 04235298, NZAC 04235317; 3, Ewing Island, 9 Feb 1973, leaf litter in  Olearia layallii forest in drainage, D.S. Horning, [50°31.713′S, 166°18.229′E], NZAC 04235065, NZAC 04235132, NZAC 04235276; 1, Ewing Island, 9 Feb 1973, Stillbocarpus polaris, </p>
            <p> Olearia layalli litter, D.S. Horning, [50°31.713′S, 166°18.229′E], NZAC 04235114; 1 (BPMN), Ewing Island, Port Ross, 1 Jan 1963, moss on tree trunk, K.A.J. Wise, [50°31.733′S, 166°18.033′E]; 1, Fleming Plateau, 9 Feb 1973, swards, J.S. Dugdale, [50°47.383′S, 165°56.833′E], NZAC 04235230; 1, Mount D′ urville, 4 Jan 1973, mats, J. Farrell, [50°49.9′S, 166°10.333′E], NZAC 04235216; 1, Waterfall Inlet, 26 Mar 2006, litter along stream margins, R. Leschen, E. Edwards, 50°48.977′S, 166°12.042′E, NZAC 04235209. </p>
            <p> Diagnosis. The Auckland Islands endemic  Thortus michauxi sp. nov. is distinguished from all other  Thortus species by having a large body with a weak supra-antennal ridge which lacks a rim and abdominal ventrite 1 lacking postcoxal lines. It is also the only species of the genus with males that have a dense brush of tenent setae on tarsomeres 1–3 on the pro- and mesolegs and the aedeagus with a well-formed basal plate in the distipenis. </p>
            <p>Description. Length 2.10–2.50 mm. Colour of body dark to light reddish-brown, mouthparts, legs, and distal parts of the antennae often lighter. Body surface semiglabrous, microsculpture strigulate over all surfaces, but weaker or absent from elytra. Dorsal setae golden, sparse, dual, consisting of very short decumbent and erect setae, and longer suberect to erect setae. Ventral surfaces with more dense and shorter erect setae of variable lengths; finely to shallowly punctate, central disc of metaventrite with punctures larger and more impressed than those of prosternum, punctation of abdominal ventrites 1 to 4 less impressed than metaventrite, and on ventrite 5 denser, and with a well-defined patch of posteriorly-directed setae. Head lengthened with relatively long gena, about 2 times as long as antennomere 1; frons relatively wide and parallel-sided between antenna, width about equal to length of antennomere 1; supra-antennal ridges without a well- -developed rim and bead; vertex with regular punctation and not delimited anteriorly by a transverse crenulate line or distinctly glabrous areas; punctation of gula and sides of the head similar, distinct and ovate. Eye consisting of 12–16 facets. Antennomere 2 cylindrical, wider than long, antennomere 3 subconical, longer and as wide as 2 and longer and slightly wider than 4, proportions of antennomeres 4–8 same, antennomere 4 shorter than 3, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse and scarcely wider than 11, antennomere 11 longer than 10, shorter than 9 and 10 combined. Pronotum quadrate, pronotal length/width ratio ~1.00, shape widest at middle; anterior margin weakly sinuate; anterior angles obtuse lateral edges subparallel in basal half and gradually converging anteriorly; pronotal disc with punctures regular and uniform, widely separated by a distance between punctures up to 3–6 times their diameter; median impression absent; transversely depressed at basal 1/5; basomedial macropuncture present, ovoid and well-delineated by extensions of the posterior bead; posterolateral angles subacute and not projecting posteriorly; posterior margin evenly and weakly convex, scutellar lobe absent. Prosternal process with well-developed lateral marginal beads, without longitudinal depression medially, process not expanded apically behind procoxae, apex rounded, width subequal that of procoxa and 0.49 times as long as prosternum; procoxal cavity with weak anterolateral notch. Scutellary shield ovoid, 1.4 times as wide as long. Elytra about 1.68 times as long as combined widths, about 1.25 times as wide as width of pronotum, about 2.08 times as long as length of pronotum; setation dual, consisting of very short decumbent or erect setae, and longer suberect to erect setae; humeral plica absent; parasutural stria present in apical 1/3, with interspace asetose; punctation similar to pronotum, but less uniform, and separated by a distance of 1–4 times their diameter. Metaventrital process with anterior margin weakly curved, marginal bead connected at middle. Tarsi moderately slender, 5-5- 5 in female, 5-5- 4 in male, male pro- and mesotarsomeres 1–3 with dense tenent setae; mesotarsomeres 1–3 of subequal lengths with 1 slightly shorter than 2, 3 and 4 which are equal, mesotarsomere 5 equal to tarsomeres 1–4 combined. First abdominal ventrite without postcoxal lines; intercoxal process subacute. Aedeagus with very short parameres broadly fused to phallobase at their bases, separated at the apex, length about 2.25 times as long as wide; apices asetose, lacking membranous extensions; basipenis about 5.17 times as long as distipenis; distipenis about 1.38 times as long as wide, lateral lobes moderately separated, basal plate present; internal sac with a pair of short slender endophallites.</p>
            <p>Etymology. A high school teacher dedicated to understanding Earth history and conservation of birds, Bernard Michaux, friend and colleague of the first author, is honoured with the specific epithet.</p>
            <p>Distribution. Offshore Island: Auckland I. (AU) (Fig. 21).</p>
            <p> Remarks. Like many species of edaphic subantarctic beetles,  T. michauxi sp. nov. does not appear to be a microhabitat specialist (CARLTON &amp; LESCHEN 2001). It has been collected in leaf litter, under bark, and in moss. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFD7FFD6FE87FCFF78B9FB4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFD6FFD1FE80FB1D78CFF78F.text	3C4D87BDFFD6FFD1FE80FB1D78CFF78F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus ovalis Broun 1893	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus ovalis Broun, 1893</p>
            <p>(Figs 14, 15, 19E)</p>
            <p> Thortus ovalis Broun 1893: 1345 . </p>
            <p> Material examined (303; AMNZ, LUNZ, NZAC). SOUTH ISLAND: NN: 1, 6km up Little Wanganui River, 24 Jun 1967, litter, F. Alack, [41°22.661′S, 172°5.392′E], NZAC 04234647; 1, Karamea Area, 20 Jun 1967, litter, F. Alack, [41°14.611′S, 172°8.515′E], NZAC 04234604; 1, Karamea Bluff, 26 Jan 2005, litter, R. Leschen, T. Buckley, R. Hoare, 41°31′S, 172°1′E, RL958, NZAC 04234640; 1, Karamea Bluff, 29 Sep 1966, CLW-RRF, [41°30.926′S, 172°1.286′E], NZAC 04234582; 6 Karamea Bluff, 9 Feb 1999, litter, R. Leschen, R. Hoare, 41°31′S, 172°1′E, RL275, NZAC 04234547, NZAC 04234560, NZAC 04234636, NZAC 04234658, NZAC 04234672, NZAC 04234719; 1, Karamea Bluff, 20 Feb 1999, ex  Ramaria sp , R. Leschen, R. Hoare, 41°31′S, 172°1′E, RL346, NZAC 04235174; 14, NN, Karamea Bluff, View Hill Saddle, 2 Mar 2007,sifted wood and leaf litter,K.Marske, 41°30.981′S, 172°1.229′E, KM040, NZAC 03018796, NZAC 04234476, NZAC 04234480, NZAC 04234486, NZAC 04234531, NZAC 04234544, NZAC 04234598, NZAC 04234599, NZAC 04234614, NZAC 04234624, NZAC 04234625, NZAC 04234630, NZAC 04234655, NZAC 04234656;19, Karamea Gorge Route, along Virgin Creek, 28 Feb 2007, sifted wood and leaf litter, K. Marske, 41°14.793′S, 172°12.826′E, KM034, NZAC 03019647, NZAC 04234466, NZAC 04234496, NZAC 04234503, NZAC 04234535, NZAC 04234536, NZAC 04234548, NZAC 04234549, NZAC 04234550, NZAC 04234565, NZAC 04234567, NZAC 04234571, NZAC 04234588, NZAC 04234602, NZAC 04234605, NZAC 04234606, NZAC 04234607, NZAC 04234616, NZAC 04234648; 1, Karamea Saddle, 13 Oct 1970, J.I. Townsend, [41°30.981′S, 172°1.229′E], NZAC 04234622; 1, Mt Owen, 13 Mar 1938, C.E. Clarke, 41°33.183′S, 172°32.483′E, AMNZ 34193; 1, Mt Owen at river, 12 Mar 1938, C.E. Clarke, 41°35.4′S, 172°32.467′E, AMNZ 34194; 7, Oparara Basin, Box Canyon/Crazy Paving Caves track and carpark, 27 Feb 2007, mixed  Nothofagus and podocarp forest, sifted wood and leaf litter, K. Marske, 41°8.076′S, 172°11.525′E, KM028, NZAC 03018653, NZAC 04234443, NZAC 04234461, NZAC 04234481, NZAC 04234540, NZAC 04234595, NZAC 04234597; 6, Oparara Basin, Moria Gate Track, 1 Mar 2007, sifted wood and leaf litter, K. Marske, 41°9.084′S, 172°11.862′E, KM039, NZAC 04234521, NZAC 04234526, NZAC 04234570, NZAC 04234600, NZAC 04234703, NZAC 04234704; 1, Oparara Gorge, 9 Feb 1999, litter, R. Leschen, R. Hoare, 41°13′S, 172°09′E, RL281, NZAC 04234715; 2, Oparara River, 27 Apr 1963, litter, J.I. Townsend, [41°11.711′S, 172°11.476′E], NZAC 04234641, NZAC 04234700; 2, NN, Oparara River mouth, 1 Jan 2011, under driftwood after flood, J. Nunn, [41°12.746′S, 172°6.750′E], NZAC 04234620, NZAC 04235155; 1, Oparara Road, 27 Apr 1963, forest litter, J.I. Townsend, [41°11.711′S, 172°11.476′E], NZAC 04234618. BR: 1, 1.6km NW Capleston, Italians Creek, 21 April 1972, litter, J.S. Dugdale, [42°3.984′S, 171°55.345′E], NZAC 04234575;5, BR, 12km NE Ikamatua, 9 Feb 1965, litter, N.A. Walker, [42°11.245′S, 171°43.397′E], NZAC 04234579, NZAC 04234591, NZAC 04234608, NZAC 04234667, NZAC 04234673; 1, 3.5km N Rapahoe, 12 Jan 1998,  Nothofagus truncata leaf litter, R. Leschen, C. Carlton, 42°42′S, 171°15′E, RL051, NZAC 04234463; 1, Bell Hill Scientific Reserve, 14 Jul 1996, K.W.Drew, E.B. Spurr, [42°33.694′S, 171°34.641′E], NZAC 04234460; 1, Blackball Road, 20 Jan 2005, R. Leschen, T. Buckley, P. Lambert, 42°20.851′S, 171°23.552′E, RL926, NZAC 04234680; 3, Boatmans Creek, 4 Oct 1972, litter, G. Kuschel, [42°3.994′S, 171°55.244′E], NZAC 04134954, NZAC 04136280, NZAC 04139061; 3, Capleston, 6 Apr 1973, litter, J.C.Watt, [42°4.021′S, 171°55.243′E], NZAC 04234511, NZAC 04234516, NZAC 04234580; 1, Capleston, 4.5km SE of Cronadun, Redmans Creek, 8 Nov 1972, litter, J.C.Watt, [42°3.317′S, 171°54.217′E], NZAC 04272531; 25, Capleston, Flowers Creek, 12 Nov 1971, fungus, J.C.Watt,[42°3.209′S, 171°53.444′E], NZAC 04134341, NZAC 04234530, NZAC 04234543, NZAC 04234551, NZAC 04234552, NZAC 04234557, NZAC 04234559, NZAC 04234561, NZAC 04234562, NZAC 04234566, NZAC 04234572, NZAC 04234573, NZAC 04234574, NZAC 04234583, NZAC 04234585, NZAC 04234587, NZAC 04234589, NZAC 04234592, NZAC 04234593, NZAC 04234594, NZAC 04234596, NZAC 04234626, NZAC 04234631, NZAC 04234690; 1, Charleston, 8.5km S, 5 Nov 1965, J.I. Townsend, 41°54.346′S, 171°26.207′E, NZAC 04234663; 2, Denniston, 1 Nov 1965, forest litter, J.I. Townsend, A.C. Eyles, [41°44.167′S, 171°47.676′E], NZAC 04234472, NZAC 04234638; 6, Fletc he rs Cre ek, 7 Ma r 1972, litter, J. McBurne y, [41°59.113′S, 171°53.454′E], NZAC 04134966, NZAC 04135413, NZAC 04135699, NZAC 04136190, NZAC 04139339, NZAC 04139623; 1, Fletchers Creek, 26 Jan 1972, litter, J. S. Dugdale, [41°59.113′S, 171°53.454′E], NZAC 04136771; 1, Fletchers Creek, 1.5km W Coll Creek, 26Jan1972,litter,J.S.Dugdale,[41°59.113′S, 171°53.454′E], NZAC 04134523; 1, Fletchers Creek, SW Rotokohu, Nov 1971, J. McBurney, [41°59.113′S, 171°53.454′E], NZAC 04234698; 3, Fletchers Creek, 6km SW Rotokohu, 25 Jan 1972, litter, J. McBurney, [41°59.113′S, 171°53.454′E], NZAC 04134391, NZAC 04139128, NZAC 04139560; 2, Fletchers Creek, 6km SW Rotokohu, 26 Jan 1972, litter, J.C. Watt, [41°59.113′S, 171°53.454′E], NZAC 04134807, NZAC 04135497; 2, Grey Valley, Waipuna, 21 Apr 1971, litter, J.S. Dugdale, 42°2′S, 171°43′E, NZAC 04234578, NZAC 04234612; 2, Greymouth, Grandjeans Track, 10 Nov 2005, R. Leschen, S. Nomura, 42°28′S, 171°12′E, RL1019, NZAC 04234644, NZAC 04234722; 1, Greymouth,Rapahoe, 29 May 1963,litter,G.Kuschel,[42°42′S, 171°15′E], NZAC 04234661; 1, Hochstetter State Forest, 9 Nov 1972, litter, J.S. Dugdale, [42°22.387′S, 171°32.674′E], NZAC 04234677; 7, Hochstetter State Forest, 6km S of Ahaura, 11 Nov 1971, litter, J.C. Watt, [42°22.387′S, 171°32.674′E], NZAC 04234629, NZAC 04234633, NZAC 04234637, NZAC 04234651, NZAC 04234652, NZAC 04234657, NZAC 04234659; 3, Inangahua, 23 Jan 1957, E.S. Gourlay, [41°51.508′S, 171°57.269′E], NZAC 04234479, NZAC 04234619, NZAC 04234639; 3, Inland Pack Track, 1km N Punakaiki River, 29 Dec 2010, sifted ground litter, J. Nunn, [42°7.521′S, 171°21.423′E], NZAC 04234458, NZAC 04234617, NZAC 04234660; 1, Inland Pack Track, 1km N Punakaiki River, 29 Dec 2010, in flood debris, J. Nunn, [42°7.521′S, 171°21.423′E], NZAC 04234474; 1, Kopara Forest, 17 Jul 1996, K.W. Drew, E.B. Spurr, [42°34.042′S, 171°44.034′E], NZAC 04234459; 1, Kopara Forest, 18 Jul 1996, K.W. Drew, E.B. Spurr, [42°34.042′S, 171°44.034′E], NZAC 04234513; 1, Kopara Forest, 19 Jul 1996, K.W. Drew, E.B. Spurr, [42°34.042′S, 171°44.034′E], NZAC 04234670; 1, Lake Rotoroa, 7 Feb 1974, litter, S.B. Peck, J. Peck, [41°47.724′S, 172°35.569′E], NZAC 04234653; 2 (ANIC), Lake Rotoroa, Braeburn Track, 25-27 Mar 1980,  Nothofagus podocarp, litter, A. Newton, M. Thayer, [41°47.724′S, 172°35.569′E], NZAC 04234649, NZAC 04234701; 1, Lake Rotoroa, near road end, 5 Nov 2014,deep damp  Nothofagus litter, J.Nunn, [41°47.724′S, 172°35.569′E], NZAC 04234467; 1,Lewis Pass, 10Feb 1965,litter,N.A.Walker,[42°22.712′S, 172°23.967′E], NZAC 04234650; 3 (ANIC), Mount Robert, N slope, 21-26 Mar 1980,  Nothofagus spp. ,litter,A.Newton,M.Thayer,[41°49.330′S, 172°48.614′E], NZAC 04234499, NZAC 04234590, NZAC 04234717; 1, Norris Creek, Buller Road, 14 Oct 1970,litter,J.I.Townsend,[42°6.245′S, 172°12.171′E], NZAC 04234468; 2, Paparoa National Park, 2km NE Punakaiki, Bullock Creek Road, 12 Jan 1998,  Metrosideros robusta leaf litter, R. Leschen, C. Carlton, 42°06′S, 171°22′E, RL048, NZAC 04234507, NZAC 04234512; 1 (LUNZ), Punakaiki, 13 Jan 1983, coastal podocarp-broadleaf, litter, J.W. Early, [42°6.431′S, 171°20.413′E]; 1 (LUNZ), Punakaiki S Road, Porari Road, 6 Jun 1985, litter and moss, J. W. Early, [42°6.431′S, 171°20.413′E]; 5 (ANIC), Punakaiki, 0.8km N Bullock Creek, 23 Mar 1980, broadleaf nikau palm podocarp, litter, A. Newton, M. Thayer, [42°6.053′S, 171°21.141′E], NZAC 04234477, NZAC 04234506, NZAC 04234510, NZAC 04234705, NZAC 04234710; 1, Punakaiki, Bullock Creek, 12 Oct 1970, litter, J.I. Townsend, [42°6.053′S, 171°21.141′E], NZAC 04234711; 3 (LUNZ), Punakaiki, Bullock Creek, 1 Apr 2002, rimu, ground litter, K. Affeld, [42°6.053′S, 171°21.141′E]; 1, Punakaiki, Bullock Creek Road (within 1st km), 4 Mar 2007, dead wood with fungi, K. Marske, 42°6.026′S, 171°20.621′E, KM047, NZAC 04234708; 3, Punakaiki, Bullock Creek Road (within 1st km), 4 Mar 2007, sifted leaf litter, K. Marske, 42°6.026′S, 171°20.621′E, KM049, NZAC 04234500, NZAC 04234726, NZAC 04234727; 2, Punakaiki, Pororari River Track, near intersect with Inland Pack Track., 3 Mar 2007, sifted wood and leaf litter, K. Marske, 42°7.169′S, 171°22.156′E, KM044, NZAC 04234694, NZAC 04234706; 1, Punakaiki, Truman Track, 4 Mar 2007, dead wood with fungi, K. Marske, 42°5.629′S, 171°20.442′E, KM050, NZAC 04234720; 3, Rahu Scenic Reserve, 23 Feb 2007, beech forest, dead wood with fungi, K. Marske, 42°15.111′S, 171°57.888′E, KM004, NZAC 04234433, NZAC 04234508, NZAC 04234724; 7, Rahu Scenic Reserve, 25 Feb 2007, beech forest, sifted wood and leaf litter, K. Marske, 42°15. 111′S, 171°57.888′E, KM023, NZAC 03018756, NZAC 04234462, NZAC 04234501, NZAC 04234514, NZAC 04234674, NZAC 04234713, NZAC 04234723; 1, Rahu Scenic Reserve, Klondyke Spur Track, 23 Feb 2007, beech forest, dead wood with fungi, K. Marske, 42°18.833′S, 172°7.061′E, KM007, NZAC 04234699; 3, Reefton, 12 Apr 1977, pine forest, J.A. Wightman, [42°6.656′S, 171°52.201′E], NZAC 04234484, NZAC 04234487, NZAC 04234664; 10, Reefton, 12 Apr 1977, cutover pine, J.A. Wightman, [42°6.656′S, 171°52.201′E], NZAC 04234495, NZAC 04234628, NZAC 04234643, NZAC 04234668, NZAC 04234669, NZAC 04234671, NZAC 04234675, NZAC 04234678, NZAC 04234682, NZAC 04234684; 2, Reefton, 12 Apr 1977, burnt pine, J.A. Wightman, [42°6.656′S, 171°52.201′E], NZAC 04234646, NZAC 04234693;16, Reefton, 12 Apr 1977, beech forest, J.A.Wightman, [42°6.656′S, 171°52.201′E], NZAC 04234478, NZAC 04234485, NZAC 04234488, NZAC 04234489, NZAC 04234490, NZAC 04234491, NZAC 04234493, NZAC 04234494, NZAC 04234504, NZAC 04234505, NZAC 04234676, NZAC 04234707, NZAC 04234714, NZAC 04234716, NZAC 04234721, NZAC 04234725; 1, Reefton, 5 Nov 1958, J.M. Hoy Collection, [42°6.656′S, 171°52.201′E], NZAC 04234692; 4, Reefton, 3.5km up Rahu Road, 25 Nov 1961, litter, J.I. Townsend, [42°6.656′S, 171°52.201′E], NZAC 04234464, NZAC 04234558, NZAC 04234569, NZAC 04234635; 1, Reefton, Dublin Terrace, 25 Nov 1961, leaf litter, J.I. Townsend, [42°6.656′S, 171°52.201′E], NZAC 04234563; 1, Rotoiti, 3 Jun 1916, T. Broun Collection, [41°48.360′S, 172°50.375′E], NZAC 04234482; 1 (LUNZ), Shenandoah Saddle, 12 Nov 1981, litter, J.W. Early, [42°1.598′S, 172°14.61′E]; 1 (LUNZ), Shenandoah Saddle, 12 Nov 1981, moss, J.W. Early, [42°1.598′S, 172°14.61′E]; 1 (LUNZ), Shenandoah Saddle, 10 Jan 1982, on  Nothofagus fusca , moss, R.M. Emberson, [42°1.598′S, 172°14.61′E]; 6, St Arnaud, Lake Rotoiti, 19 Feb - 15 Mar 2004, beech forest, M. Sim, 41°49.817′S, 172°51.217′E, AMNZ 67820, AMNZ 67821, AMNZ 67822, AMNZ 67823, AMNZ 67824, AMNZ 67825; 7, Tawhai State Forest, 3km S Reefton, 9 Nov 1971, litter, J.S. Dugdale, [42°9.448′S, 171°48.324′E], NZAC 04234564, NZAC 04234654, NZAC 04234683, NZAC 04234686, NZAC 04234688, NZAC 04234689, NZAC 04234691; 1, Victoria State Forest Park, Shenandoah Saddle, 22 Feb 2007, beech forest, sifted wood and leaf litter, K. Marske, 42°1.117′S, 172°14.413′E, KM002, NZAC 03019164; 26, West Inangahua State Forest, Stoney Creek, 25 Jan 1972, litter, J.S. Dugdale, [41°59.865′S, 171°52.945′E], NZAC 04233862, NZAC 04233965, NZAC 04234051, NZAC 04224179, NZAC 04224210, NZAC 04224211, NZAC 04234546, NZAC 04134725, NZAC 04134832, NZAC 04134924, NZAC 04135096, NZAC 04135102, NZAC 04135117, NZAC 04135581, NZAC 04136335, NZAC 04136803, NZAC 04136807, NZAC 04136859, NZAC 04136868, NZAC 04137378, NZAC 04137712, NZAC 04137734, NZAC 04137809, NZAC 04138330, NZAC 04138684, NZAC 04139536; 19, Woods Creek Track, 24 Feb 2007, regenerating  Wienmannia racemosa and rahu forest, sifted leaf litter, K. Marske, 42°33.194′S, 171°20.926′E, KM013, NZAC 03019555, NZAC 04234470, NZAC 04234471, NZAC 04234497, NZAC 04234502, NZAC 04234515, NZAC 04234532, NZAC 04234538, NZAC 04234603, NZAC 04234621, NZAC 04234634, NZAC 04234645, NZAC 04234665, NZAC 04234687, NZAC 04234695, NZAC 04234696, NZAC 04234697, NZAC 04234702, NZAC 04234709. SC: 4, Geraldine, Peel Forest Scientific Reserve, 27 Dec 1956, under bark of Kanikoreas matais, R. Crowson, [43°53.338′S, 171°14.96′E], NZAC 04234662, NZAC 04234712, NZAC 04272543, NZAC 04272544. WD: 2, Okuku Scenic Reserve, 46.7km W Otira, 12 Jan 1998,  Laurelia novae-zelandiae leaf litter, R. Leschen, C. Carlton, 42°43′S, 171°14′E, RL050, NZAC 04234469, NZAC 04234492; 1, Open Bay Islands, Taumaka Island, 30 Apr 1976, litter mainly  Schefflera digitalis ,  Melicytus ramiflorus, R.H. Matlin , [43°51.644′S, 168°52.95′E], NZAC 04234518; 1, Ross, Mikonui Track, 9 Nov 2005, ants in wood, R. Leschen, S. Nomura, 43°0′S, 170°1′E, RL1015, NZAC 04234685. FD: 1, Bauza Island, 18 Mar 1984, penguin moulting, C.F. Butcher, [45°17.129′S, 166°54.326′E], NZAC 04235072; 2, Bauza Island, 27 Nov 1981, litter and moss, C.F. Butcher, [45°17.129′S, 166°54.326′E], NZAC 04234554, NZAC 04234586; 2, Bauza Island, Mar 1984, C.F. Butcher, [45°17.129′S, 166°54.326′E], NZAC 04234509, NZAC 04272545; 1, Secretary Island, Grono Bay, 28 Nov 1981, sifted litter, C.F. Butcher, [45°17.044′S, 166°56.321′E], NZAC 04234666; 3, Wilmot Pass, Jan 1970, litter, I. Townsend, [45°30.482′S, 167°11.553′E], NZAC 04234529, NZAC 04234584, NZAC 04234718; 1, Wilmot Pass, Deep Cove, Jan 1970, litter, A.C. Eyles, [45°27.887′S, 167°9.709′E], NZAC 04234681. </p>
            <p> Diagnosis.  Thortus ovalis is recognized by its multicoloured body, a narrow clypeal constriction, short gena, and an absent or weakly indicated, incomplete parasutural stria. It has a well-developed eye and can be distinguished from  T. parallelus sp. nov. and  T. bullerensis sp. nov. by the larger and more impressed median basal puncture on the pronotum and the shorter subcoxal lines on the first abdominal ventrite. </p>
            <p>Redescription. Length 1.31–1.60 mm. Colour of body variable, dark to light reddish or dark brown, sometimes the pronotum is light with darker edges, elytra usually pale to light at the base and humeral area, sometimes tricoloured, mouthparts, legs, antennal club lighter, and central parts of the ventral sclerites lighter. Body surfaces glabrous, microsculpture absent from most surfaces, very weak and present on ventral surfaces of the head, hypomeron and lateral portions of the abdominal ventrite, glabrous in southern populations (Fiordland). Dorsal setae silver, variable, sparse, usually dual, consisting of long scattered suberect and subdecumbent setae, with those at the base of the elytra longer and often more erect, some specimens with shorter setae. Ventral surfaces typically with shorter suberect setae; punctation ovate on prosternum, mesoventrite foveolate, not strongly impressed, disc of metaventrite with punctures smaller than those of prosternum, punctation of abdominal ventrites 1 to 4 smaller or equal to those on metaventrite, setae longer at sides, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head not lengthened with relatively short gena subequal to length of antennomere 1; frons constricted, narrowed to a width that is about 1/3 less than half the length of antennomere 1; supra-antennal ridge with well-developed rim, bead absent; vertex delimited anteriorly by a transverse line that may be weakly crenulate line, surfaces punctured and glabrous; gular punctation regular and ovate, diameters larger than those on sides of head. Eye consisting of approximately 15 facets. Antennomere 2 cylindrical, slightly wider than long, antennomere 3 subconical, longer and narrower than 2 and longer and slightly wider than 4, proportions of antennomeres 4–8 same, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse and slightly narrower than 11, antennomere 11 longer than 10, shorter than 9 and 10 combined. Pronotum transverse, pronotal length/width ratio ~1.13, widest at basal third or at middle and parallel-sided (mostly southern populations); anterior margin weakly convex; anterior angles obtuse, not projecting; lateral edges weakly convex and gradually converging anteriorly, weakly curved posteriorly; pronotal disc with punctation variable from shallow to well-impressed, irregular, sparse to absent in central disc (without a median glabrous strip), separated by a distance of up to 2–4 times their diameter; median impression absent; transversely, and abruptly depressed at basal 1/5; basomedial macropuncture present, forming a broad groove and well-delineated by a polished surface and lateral ridges; posterolateral angles almost right, rounded, not projecting posteriorly; posterior margin sinuate with a weak scutellar lobe. Prosternal process with weak lateral marginal beads, without longitudinal depression medially, process parallel-sided and not expanded posteriorly behind procoxae, apex weakly curved, width subequal to width of procoxa and 0.5 times as long as length of prosternum; procoxal cavity without anterolateral notch. Scutellary shield ovoid and transverse, 2.8 times as wide as long. Elytra about 1.47 times as long as combined widths, about 1.13 times as wide as pronotum, about 1.67 times as long as length of pronotum; setation variable, consisting of very long and short subdecumbent setae; humeral plica reduced; parasutural stria absent or weakly indicated in some specimens with interspaces setose; punctation similar to pronotum, may be weakly impressed in some specimens and in central surfaces. Metaventrital process with anterior margin straight or weakly curved, marginal bead weakly indicated at middle. Tarsi moderately slender, 5-5- 5 in female, 5-5- 4 in male, male pro- and mesotarsomeres 1–3 with tenent setae; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 subequal to tarsomeres 1–4 combined. First abdominal ventrite with broad and truncate intercoxal process, postcoxal lines short, divergent, extending at most to basal one-fourth of sclerite. Aedeagus with relatively short parameres articulated to phallobase, paramere wider than long; apices subacute, unisetose, without membranous projections; basipenis about 3.33 times as long as distipenis; distipenis about 1.13 times as long as wide, lateral lobes separated, basal plate absent; internal sac with a pair of short slender sclerites apical endophallites.</p>
            <p>Distribution. South Island: NN, BR, WD, SC, FD (Fig. 21).</p>
            <p> Remarks.  Thortus ovalis is found in the South Island. The variable colour, punctation and subtle differences in setation among the specimens does not appear to be geographically related; though specimens from southern populations have the prothorax parallel-sided and the microsculpture on the venter is reduced or absent. The single record from South Canterbury based on a collection made by Roy Crowson contained 2 specimens of  T. ovalis (one was teneral) which must have been a mislabelling of the locality. Information on the holotype of  T. ovalis was given by LESCHEN &amp; GIMMEL (2012). </p>
            <p>Most specimens have been collected by sifting litter and dead wood, and rarely from fungi. They may also be collected from pitfall traps, under bark and rarely from moss.</p>
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	https://treatment.plazi.org/id/3C4D87BDFFD6FFD1FE80FB1D78CFF78F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFD1FFD0FC66FF5E78A6FC2F.text	3C4D87BDFFD1FFD0FC66FF5E78A6FC2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus parallelus Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus parallelus sp. nov.</p>
            <p>(Fig. 16)</p>
            <p>
                 Type material.   HOLOTYPE: ♁ (NZAC), labelled: ‘ New Zealand. NN. Oparara Basin, Oparara Arches Tk. 27 Feb 2007. K. Marske. KM026 Sifting. mixed  Nothofagus /podocarp fores. Sifted wood and leaf litter. 41.09S 172. 11E // NZ Arthropod Collection Private Bag 92170 Auckland  
                <a title="Search Plazi for locations around (long 172.11/lat -41.09)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=172.11&amp;materialsCitation.latitude=-41.09">New Zealand</a>
                 NZAC03019630  ’. PARATYPE (1; NZAC). SOUTH ISLAND: NN: 1, Oparara Basin, Box Canyon/Crazy Paving Caves track and carpark, 27 Feb 2007, mixed  Nothofagus and podocarp forest, sifted wood and leaf litter, K. Marske, 41°8.076′S, 172°11.525′E, KM028, NZAC 04235139. 
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            <p> Diagnosis.  Thortus parallelus sp. nov. is a unicolourous species with a well-developed eye, a narrowed clypeus and parallel abdominal postcoxal lines. It is most similar to  T. bullerensis sp. nov. but the pronotal sides are subparallel and the abdominal postcoxal lines are parallel. </p>
            <p>Description. Length 1.35 mm. Colour of body unicolourous red-brown, with lighter antennal club, mouthparts and legs. Body surfaces glabrous, microsculpture absent, apart from the gular region. Dorsal setae golden, consisting of short scattered suberect straight and subdecumbent curved setae. Ventral surfaces with suberect straight and subdecumbent curved setae; punctation ovate on prosternum, mesoventrite foveolate and moderately impressed, disc of metaventrite with punctures variable, larger on lateral portions, and about as impressed as those on prosternum, punctation of abdominal ventrites 1 to 4 smaller or equal to those on metaventrite, setae not longer at sides, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head not lengthened with a relatively short gena with length subequal to length of antennomere 1; frons constricted, narrowed to a width that is less than about half the length of antennomere 1; supra-antennal ridge with well-developed rim, bead absent; vertex delimited anteriorly by a transverse crenulate line, anterior surface distinctly punctured, posterior surface virtually glabrous and lacking punctures; gular punctation irregular and ovate, diameters larger and more dense than those on sides of head. Eye consisting of approximately 17 facets. Antennomere 2 barrel-shaped, wider than long, antennomere 3 subconical, longer and narrower than 2 and longer and slightly wider than 4, proportions of antennomeres 4–8 same, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse and about the same width as 11, antennomere 11 longer than 10, slightly shorter than 9 and 10 combined. Pronotum transverse, pronotal length/width ratio ~0.91, widest at middle; anterior margin convex; anterior angles obtuse, not projecting and rounded; lateral edges weakly convex, slightly converging anteriorly, gradually converging posteriorly; pronotal disc with punctation subuniform and foveolate, shallow, sparse in central disc (without a distinct median glabrous strip), separated by a distance of up to 1–1.5 times their diameter; median impression absent; transversely depressed at basal 1/5; basomedial macropuncture present, triangulate; posterolateral angles almost right, sharp, not projecting posteriorly; posterior margin sinuate with weak scutellar lobe. Prosternal process with well-developed lateral marginal beads, without longitudinal depression medially, process subparallel-sided and converging posteriorly, slightly expanded posteriorly behind procoxae, apex rounded, width subequal to width of procoxa and 0.5 times as long as prosternum; procoxal cavity with anterolateral notch. Scutellary shield trapezoidal and transverse, 1.9 times as wide as long. Elytra about 1.31 times as long as combined widths, about 1.12 times as wide as width of pronotum, about 1.87 times as long as length of pronotum; setation dual consisting of relatively short suberect and subdecumbent setae; humeral plica weak; parasutural stria present, incomplete, present to middle of elytra; punctation less dense and smaller than pronotum, more impressed at base. Metaventrital process with anterior margin straight, marginal bead weakly indicated at middle. Tarsi moderately slender, 5-5- 5 in female, male unknown; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 shorter than tarsomeres 1–4 combined. First abdominal ventrite with broad rounded intercoxal process, postcoxal lines long, parallel, extending to middle of sclerite.</p>
            <p> Etymology. The specific epithet is the Latin adjective  ‘ parallelus ’, referring to both the shape of the abdominal postcoxal lines and the subparallel sides of the pronotum. </p>
            <p>Distribution. South Island: NN (Fig. 21).</p>
            <p> Remarks.  Thortus parallelus sp. nov. is known from two female specimens collected at Oparara Arches. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFD1FFD0FC66FF5E78A6FC2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFD0FFD0FE8FFC3D7E3BF86F.text	3C4D87BDFFD0FFD0FE8FFC3D7E3BF86F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus simplex Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus simplex sp. nov.</p>
            <p>(Fig. 17)</p>
            <p> Type material.  HOLOTYPE: ♁ (NZAC; card-mounted after dissection), labelled: ‘ Fell Pk. Richmond Ra. 4250′ 13.iii.69 J.C. Watt // Litter // NZ Arthropod Collection Private Bag 92170 Auckland New Zealand NZAC04235076 ’.</p>
            <p> Diagnosis.  Thortus simplex sp. nov. is a unicolourous species with a unifacetted eye. It also lacks well-developed abdominal postcoxal lines which, instead, has a broad bead that extends a short distance onto the disc. It most closely resembles  T. luscus sp. nov. but differs from it by having the pronotum widest in the anterior third and the elytra lacking parasutural striae. </p>
            <p>Description. Length 1.37 mm. Colour of body unicolourous red-brown, with lighter antennal club, mouthparts and legs. Body surfaces glabrous, microsculpture absent. Dorsal setae silver, consisting of short scattered suberect, curved setae. Ventral surfaces with suberect slight curved setae; punctation ovate, prosternum mainly glabrous, mesoventrite foveolate, strongly impressed, disc of metaventrite with shallowly impressed punctures, punctation of abdominal ventrite 1 similar to metaventrite, punctation on ventrites 2 to 4 more denser, setae not longer at sides, but more erect than those on disc, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head not lengthened with a relatively short gena with length subequal to length of antennomere 1; frons moderately constricted, narrowed to a width that is longer than half of the length of antennomere 1; supra-antennal ridge with well-developed rim, bead present; vertex delimited anteriorly by a transverse crenulate line, anterior surface distinctly punctured, posterior surface glabrous, small punctures present along the base; gular punctation irregular and ovate, diameters larger than those on sides of head. Eye consisting of a single facet. Antennomere 2 barrel-shaped, wider than long, antennomere 3 subconical, subequal and narrower than 2 and longer and nearly the same width as 4, proportions of antennomeres 4–8 same, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse and about the same width as 11, antennomere 11 longer than 10, nearly as long as 9 and 10 combined. Pronotum transverse, pronotal length/width ratio ~0.88, widest at apical third; anterior margin convex; anterior angles obtuse, slightly projecting and acute; lateral edges weakly curved and convergent anteriorly, gradually converging posteriorly; pronotal disc with punctation somewhat uniform and foveolate, shallow, absent in central disc with a poorly defined median glabrous strip, separated by a distance of up to 1/2 to 1 times their diameter, though fused and larger in basal half; median impression absent; transversely depressed at basal 1/5; basomedial macropuncture present, triangulate; posterolateral angles obtuse, sharp, not slightly projecting posteriorly; posterior margin sinuate with a scutellar lobe. Prosternal process with well-developed lateral marginal beads, without longitudinal depression medially, process subparallel-sided and slightly converging posteriorly, not expanded posteriorly behind procoxae, apex curved, width subequal to width of procoxa and 0.5 times as long as length of prosternum; procoxal cavity without anterolateral notch. Scutellary shield heart-shaped and weakly transverse, 1.2 times as wide as long. Elytra about 1.44 times as long as combined widths, about 1.08 times as wide as width of pronotum, about 1.77 times as long as length of pronotum; setation uniform consisting of moderately long suberect, curved setae; humeral plica present; parasutural stria absent; punctation at base similar to that on pronotum, then smaller and sparser elsewhere on disc. Metaventrital process with anterior margin straight, marginal bead present at middle. Tarsi moderately slender, female unknown, 5-5- 4 in male, male with pro- and mesotarsomeres 1 and 2 with tenent setae; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 subequal to tarsomeres 1–4 combined. First abdominal ventrite with broad rounded intercoxal process, postcoxal lines absent, but subcoxal bead broad. Aedeagus with relatively long parameres articulated to phallobase, paramere about 2.37 times as long as wide; apices subacute bearing single microsetae, without membranous projections; basipenis about 3.12 times as long as distipenis; distipenis about 1.75 as long as wide, lateral lobes not separated, basal plate absent (?); internal sac with a pair of slender sclerites.</p>
            <p> Etymology. The specific epithet is the Latin adjective  ‘ simplex ’, meaning ‘simple’, referring to its simple, single eye facet. </p>
            <p>Distribution. South Island: MB (Fig. 21).</p>
            <p> Remarks.  Thortus simplex sp. nov. is known by one specimen collected from litter in the Richmond Range. It was completely disarticulated and remounted on a card. The tegmen was lost after imaging. The genitalia were damaged during articulation and the apex of the internal sac was broken off. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFD0FFD0FE8FFC3D7E3BF86F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFEFFFEDFE8DF8D57E9EFCCF.text	3C4D87BDFFEFFFEDFE8DF8D57E9EFCCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus sulcatus Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus sulcatus sp. nov.</p>
            <p>(Fig. 18)</p>
            <p> Type material.   HOLOTYPE: ♁ (NZAC), labelled: ‘ New Zealand SL Slopedown Ra, N of Slopedown Hill 28/1 – 5/2/08 // Pit trap c 580m Steve Rate // Manuka /inaka shrubland E2208228 N5423327 // NZ Arthropod Collection Private Bag 92170 Auckland  New Zealand NZAC04235121  .’ PARATYPES (5; NZAC). SOUTH ISLAND: MC: 1, Port Hills,Ahuriri Scientific Reserve, 7 Apr 2007, washed soil sample broadleaf forest, J. Nunn, </p>
            <p>[43°39.933′S, 172°36.643′E], NZAC 04235236. SL: 1, Owaka, Tautuku Reserve, 19 Jan 1978, S.B. Peck, J. Peck, [46°35.1′S 169°25.133′E], S. &amp; J. Peck Collection, NZAC 04272532; 1, Purakaunui, near Owaka, 31 Dec 1995, J. Nunn Collection, 46°31.181′S, 169°31.667′E, NZAC 04235103; 2, Waionepu Creek, near Tokanui, 13 Feb 1968, litter, J.I. Townsend, 46°36.376′S, 168°58.66′E, NZAC 04235127, NZAC 04235134.</p>
            <p> Diagnosis.  Thortus sulcatus sp. nov. is most similar to  T. amoenus and  T. occidens sp. nov. , which have complete parasutural striae and long genae, but is easily recognized from these and all other species by having a median pronotal impression. </p>
            <p>Description. Length 1.60–1.70 mm. Colour of body dark reddish-brown, mouthparts, legs, and antennal club lighter. Body surfaces semiglabrous, microsculpture absent. Dorsal setae silvery-gold, sparse, dual, consisting of very long decumbent and scattered suberect curved and straight setae. Ventral surfaces with shorter decumbent mostly straight setae; punctation ovate, well-impressed, and uniform, mesoventrite foveolate, central disc of metaventrite with punctures of different sizes, mostly about the same or a little larger than those of prosternum, punctation of abdominal ventrites 1 to 4 generally smaller than those on metaventrite, with those on ventrite 1 generally larger than those on ventrites 2–5, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head lengthened with relatively long gena, about 1.5 times as long as length of antennomere 1; frons constricted, narrowed to width to about half the length of 2/3 rd the length of antennomere 1; supra-antennal ridge with well-developed rim, relatively broad bead present; vertex not delimited anteriorly by a weak transverse crenulate line, surface rugulose with deep irregular punctures (one specimen weakly punctate in the middle); gular punctation deep and ovate, diameters about equal to those on sides of head. Eye consisting of about 20 facets.Antennomere 2 cylindrical, about as wide as long, antennomere 3 subconical, longer and as wide as 2 and longer and slightly wider than 4, proportions of antennomeres 4–8 same, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse and narrower than 11, antennomere 11 longer than 10, shorter than 9 and 10 combined. Pronotum subquadrate, pronotal length/width ratio ~0.96, widest at middle; anterior margin slightly curved; anterior angles obtuse; lateral edges somewhat evenly rounded and gradually converging anteriorly and posteriorly; pronotal disc with punctures regular and uniform, separated by a distance of 0.5–1.0 times their diameter; median impression present, glabrous within; not or weakly transversely depressed at base; basomedial macropuncture absent; posterolateral angles obtuse and rounded, not projecting posteriorly; posterior margin sinuate without scutellar lobe. Prosternal process with well-developed lateral marginal beads, with longitudinal depression medially, process weakly widened posteriorly behind procoxae, apex subacute, width subequal to width of procoxa and 0.47 times narrower than length of prosternum; procoxal cavity with anterolateral notch. Scutellary shield ovoid, 1.45 times wider than long. Elytra about 1.20 times as long as combined widths, about 1.17 times as wide as pronotum at greatest width, about 1.46 times as long as length of pronotum; setation singular, consisting of very long decumbent setae; humeral plica present; parasutural stria present and complete, interspace setose; punctation similar to pronotum near base, but separated by a distance of up to 1–3 times their diameter and less impressed towards apex. Metaventrital process with anterior margin straight, marginal bead completely absent. Tarsi moderately slender, 5-5- 5 in female, 5-5- 4 in male, male protarsomeres 1–2 with tenent setae; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 slightly longer than tarsomeres 1–4 combined. First abdominal ventrite with postcoxal lines convex, extending to mid-length of sclerite; intercoxal process rounded at apex. Aedeagus with relatively long parameres articulated to phallobase, paramere about 3 times as long as wide; apices subacute, asetose (?), and lacking membranous extensions; basipenis about 4.04 times as long as distipenis; distipenis about 1.67 times as long as wide, lateral lobes not separated, basal plate absent; internal sac with a pair of short slender endophallites that are anteriorly widened and spatulate.</p>
            <p> Etymology. The specific epithet is the Latin adjective  ‘ sulcatus ’, meaning ‘furrowed’, in reference to the impression on the pronotum. </p>
            <p>Distribution. South Island: MC, SL (Fig. 21).</p>
            <p> Remarks.  Thortus sulcatus sp. nov. is known from six specimens collected by leaf litter sifting. It is known by only four specimens from Southland, with one specimen collected from the Port Hills. Crowson labelled specimens of this species as ‘ Pithortus ’ n. sp. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFEFFFEDFE8DF8D57E9EFCCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFEDFFEBFC6AFC9D7F27F82C.text	3C4D87BDFFEDFFEBFC6AFC9D7F27F82C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thortus tioripatea Leschen & Chen & Harmer 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Thortus tioripatea sp. nov.</p>
            <p>(Figs 19A, C)</p>
            <p>
                 Type material.   HOLOTYPE: ♁ (NZAC), labelled: ‘ NEW ZEALAND, WD,  
                <a title="Search Plazi for locations around (long 169.30682/lat 44.15698)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=169.30682&amp;materialsCitation.latitude=44.15698">Mt Aspiring</a>
                 NP, Haast Hwy, Cameron Ck Tk K. Marske 15 Jan 2019 nr lookout // Mixed beech, broadleaf forest Sifted leaf and woody litter. 44.15698, 169.30682. 390mm KM359 // Entomology Div. D.S.I.R. New Zealand // NZ Arthropod Collection Private Bag 92170 Auckland New Zealand NZAC04254613.’ 
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            <p> Diagnosis.  Thortus tioripatea sp. nov. is recognized by having an incomplete parasutural stria and the eye composed of about 30 facets. It can be distinguished from  T. sulcatus sp. nov. by the lack of a median pronotal impression, from  T. lobatus sp. nov. by its narrower clypeal constriction, and from  T. amoenus by having more eye facets and acute posterior pronotal angles. </p>
            <p>Description. Length 1.60 mm. Colour of body light reddish-brown, mouthparts, legs, and antennal club lighter, prothorax lighter, the central disc of pronotum darker. Body surface glabrous, microsculpture absent. Dorsal setae silvery-gold, sparse, dual, consisting of short decumbent and scattered suberect setae of about equal lengths. Ventral surfaces with suberect setae of equal lengths to dorsal setae; mesoventrite foveolate, central disc of metaventrite with punctures larger and more impressed than those of prosternum, punctation of abdominal ventrites 1 to 4 weak and indistinct, punctation on ventrite 5 denser with a well-defined patch of posteriorly-directed setae. Head not lengthened with relatively short gena that is equal in length to antennomere 1; frons narrowed between antennae, width narrowed about half the length of antennomere 1; supra-antennal ridge with well-developed rim, bead absent; vertex delimited anteriorly by a transverse crenulate line, anterior and posterior surfaces glabrous; gular punctation deep and ovate, diameters equal to those on sides of head. Eye consisting of about 32 facets. Antennomere 2 cylindrical, slightly wider than long, antennomere 3 subconical, longer and as wide as 2 and longer and wider than 4, proportions of antennomeres 4–8 more or less similar, antennomere 9 wider than 8 and narrower than 10 and 11, antennomere 10 transverse and as wide as 11, antennomere 11 longer than 10, shorter than 9 and 10 combined. Pronotum subquadrate, pronotal length/width ratio ~0.82, widest at middle; anterior margin straight; anterior angles right and not protruding; lateral edges more or less parallel and not gradually converging anteriorly, very weakly curved posteriorly; pronotal disc with punctures subregular and not uniform, separated by a distance of up to 1–5 times their diameter, median glabrous strip present; median impression absent; transversely depressed at base; basomedial macropuncture absent; posterolateral angles obtuse and acute, not projecting posteriorly; posterior margin sinuate with a well-developed scutellar lobe. Prosternal process with well-developed lateral marginal beads, without deep longitudinal depression medially, process nearly parallel-sided not widened posteriorly behind procoxae, apex nearly straight, width less than width of procoxa and 0.36 times shorter than length of prosternum; procoxal cavity without anterolateral notch. Scutellary shield trapezoidal, 1.68 times as wide as long. Elytra about 1.48 times as long as combined widths, about 1.15 times greatest width of pronotum, about 2.06 times length of pronotum; setation dual of equal sizes, consisting of moderately short decumbent setae and very few scattered suberect setae; humeral plica present but reduced; parasutural stria present and short, present at apical 1/3, interspace with few setae; punctation similar to pronotum only at base, otherwise smaller and less dense, separated by a distance of up to 1–7 times their diameter. Metaventrital process with anterior margin straight, marginal bead present at sides, absent at middle. Tarsi moderately slender, female unknown, 5-5- 4 in male, male protarsomeres 1–3 with tenent setae; mesotarsomeres 1–3 of subequal lengths, mesotarsomere 5 equal to combined lengths of tarsomeres 1–4. First abdominal ventrite with postcoxal lines divergent extending just beyond mid-length of sclerite; intercoxal process rounded at apex.</p>
            <p>Etymology. The specific epithet is derived from tiori-patea, meaning in te reo Māori ‘the way ahead is clear’, for Haast Pass which was used as a traditional trade route through the Southern Alps.</p>
            <p>Distribution. South Island: WD (Fig. 21).</p>
            <p> Remarks.  Thortus tioripatea sp. nov. is known by a single male from Haast Pass. It was not dissected. </p>
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	https://treatment.plazi.org/id/3C4D87BDFFEDFFEBFC6AFC9D7F27F82C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
3C4D87BDFFEAFFEAFF01FF5E7FAAFC6F.text	3C4D87BDFFEAFFEAFF01FF5E7FAAFC6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Picrotini	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Key to species of flightless New Zealand  Picrotini</p>
            <p> Remarks. All species covered in this treatment are included below.  Callichrotus gen. nov. runs to couplet 2 with  Picrotus in the key in GIMMEL &amp; LESCHEN (2022). </p>
            <p> 1. Body elongate (Fig. 13A), pronotum with narrow lateral bead, antennal club of 3 antennomeres. ............. .......................................... 2 (  Thortus Broun, 1893 ) </p>
            <p>– Body ovate (Fig. 1B), pronotum with wide lateral bead and a deep sublateral groove, antennal club of 2 antennomeres. ........................................................ 14</p>
            <p>2. Abdominal postcoxal lines absent (Fig. 13B), at most a wide bead may be present. .................................... 3</p>
            <p>– Abdominal postcoxal lines present (Fig. 7B). ......... 4</p>
            <p> 3. Eye well developed and multifacetted; body length over 2.0 mm (Fig. 13A); Auckland Islands. ............... ................................................  T. michauxi sp. nov.</p>
            <p> – Eye unifacetted (Fig. 19D); body length less than 2.0 mm; Northland. ........................  T. helmorei sp. nov.</p>
            <p>4. Eye poorly developed, consisting of 6 or 7 facets or less (Fig. 19D). ........................................................ 5</p>
            <p>– Eye well developed, consisting of 10 or more facets (Fig. 19E). ............................................................... 8</p>
            <p> 5. Body bicoloured, pronotum darker than elytron (Fig. 8A). ...........................................  T. crowsoni sp. nov.</p>
            <p>– Body unicoloured. ................................................... 6</p>
            <p> 6. Head and pronotum coarsely punctured (Fig. 12A). .. ......................................................  T. luscus sp. nov.</p>
            <p>– Head and pronotum weakly punctured (Fig. 17A). … 7</p>
            <p> 7. Eye unifacetted (Fig. 19D); sides of pronotum weakly convex, widest at apical 1/3 (Fig. 17A). .................... ...................................................  T. simplex sp. nov.</p>
            <p> – Eye consisting of 7 facets; sides of pronotum converging anteriorly, widest at basal 1/3 (Fig. 10A). .......... ........................................................  T. latus sp. nov.</p>
            <p> 8. Pronotum with distinct impression at middle (Fig.18A); Southland. ................  T. sulcatus sp. nov.</p>
            <p>– Pronotum lacking impression at middle (Fig. 19A). .. ................................................................................ 9</p>
            <p>9. Sides of pronotum distinctly parallel-sided throughout its length (Fig. 19A); elytron with parasutural stria always present, complete or incomplete. ....... 10</p>
            <p>– Sides of pronotum weakly convex (Fig. 6A) or parallel-sided only at base (Fig. 16A); elytron with parasutural stria incomplete or absent. ......................... 12</p>
            <p> 10. Elytron with parasutural stria incomplete (Fig. 19A); posterior pronotal angles acute. ......  T. tioripatea sp. nov.</p>
            <p>– Elytron with parasutural stria complete (Fig. 6A); posterior pronotal angles rounded. ............................ 11</p>
            <p> 11. Width of frons narrower than width of antennomere 1 (Fig. 6C); metaventrite coarsely punctate; tarsomeres 5-5- 4 in male and 5-5- 5 in female. ............................. .......................................  T. amoenus (Broun, 1912)</p>
            <p> – Width of frons about as wide as width of antennomere 1 (Fig. 11C); metaventrite moderately punctate; tarsomeres 4-4- 4 in both sexes. ...........  T. lobatus sp. nov.</p>
            <p>12. Pronotum widest at middle or at anterior 1/3 (Fig. 7A), anterolateral edge of lateral carina visible in dorsal view; body unicoloured. ............................. 13</p>
            <p> – Pronotum widest in posterior 1/3 (Fig. 14A), anterolateral edge of lateral carina hidden in dorsal view; body typically bicoloured. ......  T. ovalis Broun, 1893</p>
            <p> 13. Pronotum weakly convergent towards base (Fig. 7A), widest at anterior 1/3; scutellar shield strongly transverse, about 2.5 times wider than long; abdominal postcoxal lines weakly divergent. ..................................... ..............................................  T. bullerensis sp. nov.</p>
            <p> – Pronotum subparallel-sided at base (Fig. 16A), widest at middle; scutellar shield about 2.0 times wider than long; abdominal postcoxal lines parallel. ................... ................................................  T. parallelus sp. nov.</p>
            <p>14. Clypeus constricted between antennal insertions (Fig. 3C), eye distant from antennal fossa (Fig. 3C), antennomere 10 transverse, abdominal ventrites free. ... 15</p>
            <p> – Clypeus not constricted between antennal insertions (Fig. 1C), eye contacting antennal fossa (Fig. 1C), abdominal ventrites 1 and 2 connate, antennomere 10 not transverse. .....  Callichrotus gimmeli gen. &amp; sp. nov. </p>
            <p> 15. Pronotal disc with basomedial puncture (Fig. 3A), prosternal process with apex truncate, widely distributed in North and South Island. ................................ .............................  Picrotus thoracicus Sharp, 1886</p>
            <p> – Pronotal disc without basomedial puncture (Fig. 5A), prosternal process with apex rounded; restricted to the Wairarapa. ...............................  P. wairarapa sp. nov.</p>
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	https://treatment.plazi.org/id/3C4D87BDFFEAFFEAFF01FF5E7FAAFC6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Leschen, Richard A. B.;Chen, Yandong;Harmer, Aaron M. T.	Leschen, Richard A. B., Chen, Yandong, Harmer, Aaron M. T. (2024): Revision of flightless New Zealand Picrotini (Coleoptera: Cryptophagidae): phylogeny of Thortus, eye reduction, and rarity. Acta Entomologica Musei Nationalis Pragae 64 (2): 455-500, DOI: 10.37520/aemnp.2024.031, URL: https://doi.org/10.37520/aemnp.2024.031
