taxonID	type	description	language	source
466487E16F43FF831C5F78001EA5FC8A.taxon	description	(9: 680 – 1030 m), fr.: on moist loamy spots of bare soil in low bush-moss- and bushy herbaceous tundra with Betula nana, on fine earth between boulders in rock fields and on edges of snowfields, on soil in snowbed communities [121221]. The species occurs in pure mats or mixed with widespread, mostly acidophilous hepatics (Pseudolophozia sudetica, Lophozia wenzelii, Fuscocephaloziopsis albescens, Cephalozia bicuspidata, Marsupella sprucei, Solenostoma confertissimum, Nardia geoscyphus, Plectocolea hyalina). It was collected twice on carbonate cliffs covered by soil where it occurred with some mostly calciphilous bryophytes: Preissia quadrata, Blepharostoma trichophyllum subsp. brevirete, Mesoptychia heterocolpos, Cyrtomnium hymenophylloides [121176].	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F43FF831C5F7A701E90FB5F.taxon	description	– 2, 6, 8, 11, 12 (8: 633 – 973 m), sp.: tundra belt, on fine earth on sides of sinkholes and in shaded niches on carbonate cliffs and rocks [121150], in pure mats or mixed with Mesoptychia gillmanii, Clevea hyalina, Sauteria alpina, Tritomaria scitula, Blepharostoma trichophyllum subsp. brevirete, Distichium capillaceum. Occurs as well on soil and fine earth in mossy beds and on sides of temporary streams where it grows in pure mats or mixed with Preissia quadrata, Mnium spinosum, Philonotis fontana, Pohlia wahlenbergii. It is an almost circumpolar arctomontane calcicolous species. It has previously been reported for Russia from Caucasus, the Murmansk Province, the mountains of Yakutia and the Far East (Borovichev et al., 2015). Finding the species in Urals in areas with carbonate rocks fills in the huge gap in its distribution.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F43FF831C5F7D85187FF99C.taxon	description	– 2, 19, 21 (4: 380 – 636 m), sp.: on soil in moist mixed Betula-Picea-Pinus sibirica and Betula-Picea tall herbs- Vaccinium myrtillus - green moss forests on steep slopes of mountains [121240] and in wet Picea-Betula forests in valleys of brooks, both in mats without admixture of other bryophytes and mixed with Barbilophozia hatcheri, Lophocolea minor, Lophozia cf. silvicola, Sanionia uncinata. It was once collected in Larix open forest on a carbonate cliff as single stems in mats with Lophocolea minor, Mnium thomsonii, Callicladium haldanianum, Pohlia cruda, Platydictya jungermannioides (72 / 5).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F43FF831C5F7F46182CF8F5.taxon	description	– 1, 2, 4, 8,13, 11, 12, 14, 15, 16, 17, 18, 19, 20 (20: 380 – 840 m), com.: on soil and soil covered rocks in grass- and grass-moss tundras, on soil among herbs, between rocks and on fine earth covered rocks in rock fields, on soil at the bases of trees and on mossy rocks in alder thickets, larch and birch low growth open forests, mixed Betula-Picea and dark coniferous tall herb-moss forests [121118]. The species was gathered several times in sinkholes and on fine earth on ledges on carbonate cliffs in tundras. It occurs both in pure mats and mixed with Barbilophozia barbata, Schljakovia kunzeana, Ptilidium ciliare, Lophozia wenzelii, Lophoziopsis longidens, etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F43FF821C5F7E1F1C52FEDB.taxon	description	– 3, 8, 11, 15, 16, 17, 18, 19, 20, 21 (16: 380 – 850 m), com.: on soil in mixed birch-spruce and dark coniferous forests, in larch and birch low growth open forests, in swampy willow thickets along groundwater seepages, on fine earth covered rocks and on soil in grass and grass-moss tundras [121125]. Often in pure mats, but sometimes mixed with Lophoziopsis longidens, Pleurozium schreberi, Lescuraea secunda, Hylocomiastrum pyrenaicum.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821FFC7BA71CD1FBAB.taxon	description	– 9: sedge- Sphagnum bog at the bottom of quartzite ridge [121187], mixed with Polytrichum strictum or Schistochilopsis incisa, Lophozia wenzelii, Scapania parvifolia, Cephaloziella arctogena. The species was previously known in the Urals from one locality in Polar Urals (Konstantinova & Chernyadjeva, 1995) and one site on Mt. Ner-Oika (Konstantinova & Lapshina, 2014). It is a world wide rare species that has status “ disappeared ” in Central and Western Europe (Schumacker & Matriny, 1995) and listed as rare in the Red Data Book of Russia (Bardunov, 2008). It has recently been found in many sites in Siberia (Konstantinova & Savchenko, 2008).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821FFC7D511FFFFB58.taxon	description	– 1: Puiva village, on soil on road side under Alnus (43 / 2 - 1) mixed with Scapania curta, Pohlia filum. 19: valley of Puiva River [121260], road side, on soil, without admixture of other bryophytes or mixed with Scapania curta, Pohlia filum, Dicranella subulata, Nardia japonica (52 / 1 - 2). Widespread cosmopolitan species.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821FFC7D821F76F9B2.taxon	description	– 11, 19, 20, 21 (5: 433 – 890 m), sp.: on spots of loamy bare soil in Betula nana - grass tundra, mixed with Anthelia juratzkana, Nardia geoscyphus, Scapania curta, Cephalozia bicuspidata, Prasanthus suecica, on fine earth between boulders in rock fields, often with Cephalozia pleniceps, among mosses in dwarf-sedge-moss bogs with Scapania paludicola, Sphenolobus minutus var. grandis, Ptilidium ciliare, Warnstorfia sarmentosa, Aulacomnium turgidum, Loeskypnum badium, in Betula-Picea tall-herb-moss forest, on decaying wood in thin pure mats with admixture of Sciuro-hypnum reflexum and other bryophytes on decaying wood [121266].	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821FFC7F5B1972FEF8.taxon	description	– 2, 4, 6, 8, 9, 12, 17 (11: 630 – 926 m alt.), fr.: on carbonate cliffs in tundras, larch low growth open forests, in alder thickets, on humus and carbonate fine earth on ledges and in niches under rocks on cliffs and in rock fields, on sides of brooks near carbonate rocks [121312]. The species occurs both in pure mats and mixed with Tritomaria scitula, Distichium capillaceum, Mesoptychia heterocolpos, M. gillmanii, M. collaris, Odontoschisma macounii, Saccobasis polita, Preissia quadrata, Clevea hyalina, Cyrtomnium hymenophylloides, Mnium thomsonii, Pohlia cruda, Platydictya jungermannioides. This is the first record of the subspecies for the eastern side of the Urals. The subspecies brevirete has been reported earlier for the western side of the Urals, particularly the Vishera State Nature Reserve (Konstantinova & Bezgodov, 2006). As opposed to the typical subspecies B. trichophyllum subsp. brevirete is restricted to calcareous areas and high latitudes and altitudes. It is much more common than subsp. trichophyllum in the high Arctic and at high altitudes in mountains.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821C0278E2184AFD76.taxon	description	– 4: rock field at the bottom of Zhilnyi brook (64.48296 ° N – 59.65805 ° E; 688 m alt.), on fine earth between rocks, some thalli in mats dominated by Lophozia wenzelii and admixture of Trilophozia quinquedentata, Sanionia uncinata (16 / 5). 9: on moist carbonate fine earth under boulders in rock field [121181, 121186] in pure mats and some thalli among Diplophyllum albicans, D. taxifolium, Pohlia crudoides. C. laxa is an arctomontane species with predominantly Asian – western North American distribution and a single locality in the European Arctic (Konstantinova & Lavrinenko, 2002) and another in the subarctic (Borovichev, 2013), which presumably represent post-glacial invasion (Konstantinova & Mamontov, 2010).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821C027B9C1EB1FC06.taxon	description	– 19, 20, 21 (3: 380 – 433 m), sp.: on decaying wood in coniferous mountain forests [121285] in pure mats or mixed with Fuscocephaloziopsis lunulifolia, F. pleniceps, Blepharostoma trichophyllum.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821C027AEC19FEFB26.taxon	description	– 17: northern spur of Mt. Shatmaga, dwarf shrub-sedge green moss- Sphagnum bog at the bottom of mountain [121267], mixed with Riccardia latifrons, Blepharostoma trichophyllum, Neoorthocaulis binsteadii, Scapania paludicola, Trilophozia quinquedentata, Dicranum laevidens, Warnstorfia sarmentosa. 22: upper part of the forest belt in the middle of Puiva River [121301], at the bottom of mountain, edge of bog, on base of tree mixed with Mylia anomala, Schljakovia kunzeana, Aulacomnium palustre.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821C027DCC1E8CFA0D.taxon	description	– 5: rock field at the edge of snowfield in valley of a brook (64.47855 ° N – 59.63406 ° E; 783 m), on fine earth between boulders (24 / 3), single shoots in mats of Schistochilopsis opacifolia. 9: sedge- Sphagnum bog at the base of quartzite ridge (64.49596 ° N – 59.65054 ° E; 890 m alt.), small admixture in mats of Neoorthocaulis binsteadii, with Lophozia longiflora and Cephaloziella spinigera (30 / 1).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821C027CF7196DF908.taxon	description	– 3, 4, 5, 7, 9, 11, 10, 19, 17 (12: 476 – 980 m), com.: on moist loamy spots in gravel lichen and dwarf-shrub-moss tundras, on fine earth between huge boulders in rock fields, in snowbeds, between mosses in dwarf-shrub-sedge-moss bogs, on sides of brooks [121166]. Usually mixed with Lophozia spp., Fuscocephaloziopsis pleniceps, F. albescens, Scapania curta, Nardia geoscyphus, Plectocolea hyalina, Harpanthus flotovianus, Schistochilopsis opacifolia.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821C027FF218C6F8F6.taxon	description	– 9, 10, 19, 20 (4: 433 – 994 m), sp.: on moist loamy spots in dwarf-sedge-moss tundra, in bryophyte mats in sedge-Sphagnum bog, on decaying wood and at bases of trees in birch-dark coniferous tall herbs-green moss forests [121297]. The species always occurs as small admixture in mats of bryophytes. For the Northern Urals it has previously been reported for Mt. Ner-Oika (Konstantinova & Lapshina, 2014).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF851C027E1C1C7CFEF8.taxon	description	– 8: on carbonate cliffs in tundra [121176], on fine earth between rocks, some stems in mats dominated by Preissia quadrata; 16: on soil in herbaceous tundra on gentle south facing slope (64.49586 ° N – 59.67885 ° E; 810 m alt.), admixture in mats with dominated by Pseudolophozia sudetica (53 / 4 - 2, - 3) or mixed with Lophozia wenzelii and Lophoziopsis excisa. 17: northern spur of Mt. Shatmaga [121265], dwarf shrub-sedge- Sphagnum bog at the bottom of slope, admixture in mats with dominated by Scapania tundrae.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F42FF821FFC78011FD1FD3D.taxon	description	– 4, 5, 7, 8, 11, 16, 19 (12: 676 – 980 m), fr.: on fine earth covered siliceous rocks, between boulders in rock fields on mountain slopes, on soil in snowbed communities, on edges of snowfields, on soil on the side of sinkholes, on road sides [121133]. In pure mats or mixed with Diplophyllum albicans Tetralophozia setiformis, Gymnomitrion concinnatum, Lophozia wenzelii, L. murmanica, Fuscocephaloziopsis albescens, Pohlia crudoides, Kiaeria falcata, Andreaea rupestris, Racomitrium lanuginosum.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851FD678E21F5BFE56.taxon	description	– 18: Betula-Vaccinium myrtillus - green moss open forest [121239] on bare soil without admixture of other bryophytes.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851FD678BC1F4BFD3D.taxon	description	– 9: sedge- Sphagnum bog at the base of quartzite ridge (64.49596 ° N – 59.65054 ° E; 890 m alt.), single shoots in mats dominated by Neoorthocaulis binsteadii mixed with Lophozia longiflora and Cephalozia ambigua (30 / 1). 20: upper part of forest belt in the Middle Puiva River (64.48790 ° N – 59.69986 °; 380 m alt.), seepage at the bottom of mountain at the edge of a bog, on base of tree in mats of Dicranum sp. (66 / 2).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851FD67BA71FFBFC06.taxon	description	– 10: rock field impounded by water from lake (64.50243 ° N – 59.63124 ° E; 990 m alt.), on fine earth as admixture in mats with Lophozia murmanica, Gymnomitrion concinnatum, Sanionia uncinata, Oncophorus wahlenbergii, Hylocomiastrum pyrenaicum (41 / 1 - 1). The species has previously been reported for the Urals from Mt. Ner-Oika. (Konstantinova & Lapshina, 2014).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851FD67AEC1F99FBED.taxon	description	– 6: mossy rocky bed of brook [121228], on rocks covered by fine earth, mixed with Marchantia polymorpha ssp. montivagans, Pohlia wahlenbergii; mossy rocky bed of temporary brook, on fine earth on rocks in mats dominated by Mesoptychia gillmanii and admixture of Jungermannia atrovirens, Bryoerythrophyllum recurvirostre, Blepharostoma trichophyllum subsp. brevirete, Marchantia polymorpha subsp. montivagans (46 / 4 - 1).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851FD67D171FCCFA78.taxon	description	– 9: under rocks in rock field (64.49653 ° N – 59.64451 ° E; 973 m alt.), on carbonate fine earth (29 / 4 - 3), some thalli mixed with Blepharostoma trichophyllum subsp. brevirete, Tritomaria scitula, Mnium thomsonii. 12: carbonate cliff on left bank of Puiva River [121219], on fine earth in shaded niche at the bottom of cliff, mixed with Mesoptychia gillmanii, Asterella lindenbergiana, Blepharostoma trichophyllum subsp. brevirete, Orthothecium strictum, Distichium capillaceum, Isopterygiopsis pulchella, Encalypta cf. streptocarpa. It is a widespread arctomontane species restricted to areas with calcareous bedrocks. This species was previously known in the North Urals from the western side in the Republic of Komi (Bakalin et al., 2001; Dulin, 2007).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851FD67F621CE4F908.taxon	description	– 19: on side of brook, on soil among tall herbaceous vegetation, in pure mats or mixed with Pellia neesiana, Plagiothecium sp. [121259]. It is mostly boreomontane species, but it occurs sporadically at high altitudes and latitudes.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851FD67FF21CA2F828.taxon	description	– 4, 9, 17 (8: 888 – 1030 m), sp.: on fine earth between boulders in silicate rock fields [121277]. Occurs both in pure mats and mixed with Pseudolophozia sudetica, Gymnomitrion concinnatum, Tetralophozia setiformis, Lophozia murmanica, L. longiflora, Fuscocephaloziopsis albescens, Pohlia crudoides, Racomitrium lanuginosum, Andreaea rupestris. It is a widespread arctomontane species, but it has previously only been reported from a few sites in the Urals (Zinovjeva, 1973; Konstantinova & Lapshina, 2014)	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851FD67ED21973FE56.taxon	description	– 3, 9, 17 (5: 644 – 1030 m), sp.: on fine earth between boulders in rock fields, on loamy fine earth in rocky dwarf shrubs-lichen tundra, on bank of a temporary brook under a snowfield [121271]. Usually mixed with other hepatics: Marsupella sprucei, Gymnomitrion concinnatum, Anthelia juratzkana, Prasanthus suecica, Isopaches bicrenatus. The species has previously been recorded in the Urals from a single locality in the Vishera State Nature Reserve (Konstantinova & Bezgodov, 2006) and a single locality in the vicinity of Mt. Ner-Oika (Konstantinova & Lapshina, 2014).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851C5F78BC1EA3FD3D.taxon	description	– 3, 7, 9, 11, 17 (6: 644 – 946 m), sp.: on rocks and on fine earth between rocks in rock fields, on soil in snowbed along the temporary brook under snowfield [121180]. The species occurs both in pure mats and mixed with Pseudolophozia sudetica, Diplophyllum albicans, Fuscocephaloziopsis albescens, Lophozia wenzelii, L. murmanica, Gymnomitrion concinnatum, Pohlia cruda, P. crudoides.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851C5F7BA7186DFCE2.taxon	description	– 18: on road between tracks [121233], on sandy soil, in mats dominated by Nardia japonica and admixture of Marsupella sprucei, Scapania obcordata, Isopaches bicrenatus. Mountain circumboreal species restricted to bare soil.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851C5F7A081EA2FBED.taxon	description	– 3, 5, 7, 9, 10, 17 (6: 680 – 994 m), fr.: between rocks at edges of snowfields, in moist shaded niches in rock fields, in snowbeds, sometimes on spots of loamy bare soil in dwarf-shrub-sedge-moss tundra [121159], usually mixed with Anthelia juratzkana, Cephalozia bicuspidata, Lophozia murmanica, Diplophyllum albicans, D. taxifolium, Gymnomitrion concinnatum, Pohlia drummondii.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851C5F7D171939FA92.taxon	description	– 19, 20 (3: 380 – 433 m), sp.: on decaying wood in coniferous forests [121247], on base of a tree at the edge of a bog, in pure mats or mixed with Lepidozia reptans, Mylia anomala, Dicranum fuscescens, Tetraphis pellucida, etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851C5F7C781EA3F9EB.taxon	description	– 2, 3, 4, 5, 11,19 (7: 433 – 830 m), fr.: tundra belt, on fine earth in niches on carbonate cliffs mixed with Asterella lindenbergiana, Pohlia cruda, Bartramia ithyphylla, on fine earth between boulders in rock fields, among mosses in sedge-moss bog, on bare soil in snowbeds and along temporary brooks under snowfields, as well on decaying wood in mountain coniferous forests [121150]. Occurs both in pure mats and mixed with Blepharostoma trichophyllum, Cephalozia bicuspidata, Lophozia murmanica, Schistochilopsis opacifolia, Mesoptychia gillmanii.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF851C5F7F111EC8F8D2.taxon	description	– 5, 7, 9 (3: 783 – 945 m), sp.: on fine earth between rocks in rock fields at the edge of snowfield, on soil in dwarf shrubs-sedge-green moss- Sphagnum bog and in pools in sedge- Sphagnum bogs mixed with Schljakovia kunzeana, Scapania irrigua, S. irrigua subsp. rufescens, S. paludicola, Odontoschisma elongata, Pseudolophozia sudetica, Warnstorfia sarmentosa [121190].	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F45FF841C5F7E381F5BFEDB.taxon	description	– 9, 10, 17 (6: 840 – 1030 m), sp.: on fine earth-loamy cryogenic spots of bare soil in dwarf shrubs-lichens rocky tundra, on rocks in rock fields, on fine earth on silicate cliffs [121179]. Occurs in pure mats but more often mixed with Diplophyllum albicans, D. taxifolium, Tetralophozia setiformis, Marsupella sprucei, Gymnomitrion coralloides, Sphenolobus minutus, S minutus var. grandis, Anthelia juratzkana, Pseudolophozia sudetica, Lophozia longiflora, L. murmanica, Pohlia crudoides, Andreaea rupestris.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841FFC78011E5AFE6B.taxon	description	– 8, 13, 17 (3: 788 – 980 m), sp.: on fine earth on ledges of carbonate cliffs and between boulders and on loamy spots in dwarf shrub-lichens rocky tundra [121123], mixed with Sphenolobus minutus, Trilophozia quinquedentata, Gymnomitrion concinnatum, etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841FFC78911FC6FD18.taxon	description	– 5, 19 (3: 476 – 783 m), sp.: on soil and fine earth on sides of streams, in rock fields at the base of snowfields [121156], mixed with Pellia neesiana, Scapania paludosa, Lophozia longiflora, Cephalozia bicuspidata, Rhizomnium magnifolium, Sanionia uncinata, Pohlia drummondii, Bartramia ithyphylla, etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841FFC7BC21F7BFC07.taxon	description	– 17, 18 (3: 488 – 980 m), sp.: on sandy soil on roads between tracks, on fine earth-loamy cryogenic spots in dwarf-shrubmoss-lichen and dwarf-shrubs-lichen rocky tundras [121271], mixed with Anthelia juratzkana, Diplophyllum obtusifolium, Gymnomitrion concinnatum, Endogemma caespiticia, Marsupella sprucei, Nardia japonica, Scapania obcordata, S. parvifolia, Prasanthus suecica.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841FFC7AEC1FA9FB02.taxon	description	– 2, 6 (4: 633 – 634 m), sp.: on sides and in beds of mossy dry brooks [121321], on fine earth on rocks, in pure mats or mixed with Chiloscyphus polyanthos, Schistidium riparium, Hygrohypnella ochracea, Pohlia wahlenbergii, Dichodontium pellucidum, Philonotis fontana, Mesoptychia gillmanii, Bryoerythrophyllum recurvirostre. The species was collected once on the vertical wall of a carbonate cliff in larch light forest [121336], mixed with Mesoptychia heterocolpos.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841FFC7DE81C68FB7D.taxon	description	– 2: in the mossy bed of a temporary brook, in pure mats, on moist rocks [121318].	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841FFC7C671FA2FA22.taxon	description	– 11: herbaceous tundra (64.50132 ° N – 59.66006 ° E; 830 m alt.), on sides of a karst cave, on fine earth, single shoots mixed with Mesoptychia gillmanii Pohlia wahlenbergii, Philonotis sp., Asterella lindenbergiana (09 / 3). Widespread arctomontane basiphilic or calcicolous hepatic restricted to high altitudes or high Arctic.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841FFC7CC81FF7F908.taxon	description	– 13: Dwarf shrub (Betula nana, Vaccinium uliginosum) - herb-green moss tundra on steep south-east facing slope of mountain [121121], on bare soil, single shoots in mats dominated by Preissia quadrata and admixture of Lophocolea minor, Scapania cuspiduligera, Tritomaria scitula. 2: bed of mossy partly dry brook [121326], on moist vertical side of rock in pure mats or mixed with Pohlia wahlenbergii, Dichodontium pellucidum.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841FFC7FF21CB7F828.taxon	description	– 21: Betula-Picea-Pinus sibirica fern- Vaccinium myrtillus - green moss forest in the upper part of the forest belt [121284], on decaying wood in mats dominated by Dicranum sp. and admixture of Lophozia cf. silvicola or Sphenolobus minutus, Pohlia nutans, as well as on soil as admixture to Barbilophozia barbata. 19: birch-spruce tall herbs-green moss forest on south facing steep slope [121247], on decaying wood in mats dominated by Fuscocephaloziopsis lunulifolia and admixture of Tetraphis pellucida, Plagiothecium sp.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841FFC7ED2194EFE32.taxon	description	– 6: mossy rocky bed of dry brook [121286], on soil. 19: birch-spruce tall herbs-green moss forest on south faced steep slope [121244], on decaying wood in pure mats and mixed with Lophoziopsis longidens, Ptilidium pulcherrimum, Brachythecium salebrosum. 21: Betula – Picea – Pinus sibirica - fern-Vaccinium myrtillus - green moss forest in upper part of forest belt [121282], on decaying wood, mixed with Sciuro-hypnum reflexum, Stereodon plicatulus, Lophocolea minor, Sanionia uncinata.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841C0278D81EB1FD76.taxon	description	– 2, 6,13, 19, 21 (5: 380 – 725 m), sp.: on spots of bare soil in dwarf shrubs-herbs-green-moss tundra, on mossy carbonate cliffs in larch light forest [121120], mixed with Preissia quadrata, Tritomaria scitula, Scapania cuspiduligera, S. gymnostomophila, Mnium thomsonii, Pohlia cruda, Platydictya jungermannioides, sometimes on fine earth along temperate brooks. In dark coniferous and in mixed tall herbs-green moss forests the species occurs mostly on decaying wood and at base of trees [121240], mixed with Lophocolea heterophylla, Barbilophozia barbata, Ptilidium pulcherrimum.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841C027B9C19D3FC1B.taxon	description	– 20: sedge- Sphagnum bog with single trees of birch and spruce [121292], on decaying stump of wood, in pure mats. This species was previously recorded in the Urals from the western side in the Vishera State Nature Reserve (Konstantinova & Bezgodov, 2006).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841C027AC11EA4FB7D.taxon	description	– 3, 4, 9, 10, 11,13, 17, 18, 19, 20 (16: 390 – 994 m), com.: on sides of karst hollows [121130], on spots of bare loamy soil, on carbonate cliffs, in sedge-Sphagnum and dwarf shrub-moss bogs [121189], between boulders in rock fields, on rocks in birch - Vaccinium - green moss low growth open forests, on banks of brooks. In pure mats or mixed with Neoorthocaulis binsteadii, Schistochilopsis incisa, Fuscocephaloziopsis albescens, Diplophyllum albicans, Gymnomitrion concinnatum, Cephalozia bicuspidata, Lophozia murmanica, Harpanthus flotovianus. Cephalozia pleniceps.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841C027C671864F99D.taxon	description	– 4, 5, 10, 11, 17, 19 (10: 476 – 980 m), fr.: on sides of karst hollow in herbaceous tundra, on fine earth between boulders in rock fields, under snowfield in sedge-moss bog at the bottom of mountain, on side of brook [121129], in pure mats or mixed with Pseudolophozia sudetica, Lophozia longiflora, Schistochilopsis opacifolia Harpanthus flotovianus, Fuscocephaloziopsis albescens, Gymnomitrion concinnatum, Tetralophozia setiformis, Sanionia uncinata, Pohlia crudoides, P. drummondii.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841C027F471E90F898.taxon	description	– 3, 19, 20, 21 (6: 380 – 680 m), sp.: on bases of trees and decaying wood in mixed and dark coniferous forests, in sedge- Sphagnum with birch and spruce bog [121291], on soil in subalpine short grassgreen moss-liverwort meadow. Often in pure mats or mixed with Sciuro-hypnum reflexum, Sanionia uncinata, Dicranum fuscescens, D. montanum, Pohlia nutans, Plagiothecium spp., Lophocolea heterophylla, Lepidozia reptans, Ptilidium pulcherrimum.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F44FF841C027E421EFAF862.taxon	description	– 5, 14, 15, 19 (5: 610 – 783 m alt.), sp.: on fine earth between boulders in rock fields, on sides of brooks [121194], mixed with other bryophytes, on mossy rocks in birch and larch low growth open forests, in pure mats or mixed with Plectocolea hyalina, Barbilophozia hatcheri, Sciuro-hypnum reflexum.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871C5F7BE1182DFC72.taxon	description	– 2, 3, 6, 11, 12 (7: 633 – 855 m), sp.: the species is not rare on sides and in niches in karst hollows in tundra belt (usually in herbaceous and Dryas - herbaceous communities), it occurs as well on banks and mossy rocks in beds of temporary brooks [121136], both on soil and fine earth, in pure mats or mixed with Blepharostoma trichophyllum subsp. brevirete, Mesoptychia heterocolpos, M. badensis, Fuscocephaloziopsis pleniceps, Chiloscyphus polyanthos, Asterella lindenbergiana, Preissia quadrata, Bryoerythrophyllum recurvirostre, Distichium capillaceum, Platydictya jungermannioides, Pohlia wahlenbergii, P. cruda, Bartramia ithyphylla, Dichodontium pellucidum, etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871FD67B721CB0FDFB.taxon	description	– 5: on rocks in the bed and on sides of brook [121157]. This variety was described from Caucasus (Bakalin, 2005) and was until now known from Caucasus only.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871FD679B11FE5FD88.taxon	description	– 1, 3, 4, 5, 7, 9, 14, 10, 16 (11: 632 – 946 m), fr.: on soil and spots of bare loamy soil in tundras, in snowbed communities, between boulders in rock fields, in dwarf-shrub-sedge- Sphagnum bogs, in seepages, on banks of brooks and on road sides [121158], in pure mats or mixed with Pseudolophozia sudetica, Barbilophozia hatcheri, Ptilidium ciliare, Neoorthocaulis floerkei, Fuscocephaloziopsis albescens, Anthelia juratzkana, Schljakovia kunzeana, Scapania irrigua, S. curta, Plectocolea subelliptica, Cephalozia bicuspidata, Polytrichastrum sexangulare, Sanionia uncinata, Kiaeria starkei.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871FD67BE11CE4FCCD.taxon	description	– 1, 11, 15, 16, 18, 20 (6: 390 – 890 m), sp.: on soil among mosses, on spots of bare soil in herbaceous tundras, on road sides [121237], always mixed with other bryophytes: Pseudolophozia sudetica, Lophozia wenzelii, L. longiflora, Cephaloziella divaricata, Scapania mucronata, Schistochilopsis opacifolia.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871FD67A371F00FBED.taxon	description	– 15, 19, 20, 21 (6: 390 – 610 m), fr.: on decaying wood and on bases of trees in mixed birch - spruce and birch dark coniferous forests, in mixed tall herbaceous-green moss and tall herbaceous- Vaccinium myrtillus - green moss forests and birch low growth open forests [121251], occurs both in pure mats or mixed with Barbilophozia lycopodioides, B. hatcheri, Lophocolea heterophylla, Lophozia cf. ventricosa, Tritomaria exsectiformis, Ptilidium pulcherrimum, Sanionia uncinata, Dicranum spp., Brachythecium salebrosum.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871FD67CF71C31F9EB.taxon	description	– 3, 5, 6, 8, 11, 20 (9: 380 – 890 m), fr.: on soil on sides of karst hollows in grass tundras, in seepages and in snowbeds, on coarse-grained fluvial sediments on banks of streams [121151], often in pure mats or mixed with Chiloscyphus polyanthos, Jungermannia atrovirens, Mesoptychia gillmanii, Pohlia wahlenbergii, Philonotis fontana.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871FD67F111DADF898.taxon	description	– 9: rock field on quartzite ridge, on fine earth between boulders [121179], mixed with Fuscocephaloziopsis albescens, Gymnomitrion concinnatum, Diplophyllum taxifolium. This species was previously reported for the Urals by Zinovjeva (1973).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871FD67E421FE4F862.taxon	description	– 17, 18 (4: 488 – 1030 m), sp.: on spots of fine-earth-loamy soil in rocky dwarf shrublichen tundra surrounded by rock fields [121270], mixed with Gymnomitrion concinnatum, Diplophyllum obtusifolium, Prasanthus suecicus, Isopaches bicrenatus. This species was collected once on a road on sandy soil between tracks mixed with Endogemma caespiticia, Nardia japonica (47 / 3).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871C5F79B1184CFEC6.taxon	description	– 11: karst ledge in Dryas - herbaceous tundra, on fine earth in shaded moist niches [121136], mixed with Leiocolea gillmanii.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871C5F782C1859FDFB.taxon	description	– 2, 5, 8, 9, 12 (6: 634 – 926 m), sp.: carbonate rock outcrops in tundra, on fine earth at the bottom and between rocks, on rocks in the bed and on banks of dry mossy brook [121172], always mixed with other bryophytes, particularly Blepharostoma trichophyllum subsp. brevirete, Tritomaria scitula, Odontoschisma macounii, Saccobasis polymorpha, Preissia quadrata, Syntrichia norvegica, Distichium capillaceum. It was collected once in a herbs-moss-liverwort community in a snowbed, mixed with Neoorthocaulis floerkei and Lophozia wenzelii.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871C5F7A9B19A8FAE8.taxon	description	– 2, 3, 6, 8, 9, 11, 12 (10: 633 – 973 m), fr.: on carbonate outcrops, on sides of karst hollows in herbaceous tundras, on fine carbonate earth in shaded niches between boulders in rock fields, on banks and on mossy rocks in beds of temporary streams [121172]. Always mixed with other bryophytes: Mesoptychia gillmanii, M. collaris, Blepharostoma trichophyllum subsp. brevirete, Tritomaria scitula, Preissia quadrata, Jungermannia atrovirens, Distichium capillaceum, Drepanium recurvatum, Myurella tenerrima, Platydictya jungermannioides, Cyrtomnium hymenophylloides, Mnium thomsonii, Encalypta ssp., etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871C5F7C12181DFA5B.taxon	description	– 20: in seepage at the bottom of the slope on edge of a bog [121301], on base of tree, with admixture of Calypogeia sphagnicola, Cephalozia spp., Schljakovia kunzeana, Aulacomnium palustre.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871C5F7C811832F92D.taxon	description	– 1, 3, 10, 11, 17 (5: 644 – 994 m alt.), sp.: in tundra belt, on loamy cryogenic spots, on fine earth on bank of stream from snowfield, on soil on road sides [121221]. Always mixed with other bryophytes: Solenostoma confertissimum, Plectocolea hyalina, P. subelliptica, Anthelia juratzkana, Scapania curta, Cephalozia bicuspidata, Psilopilum laevigatum, Pohlia drummondii.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871C5F7FD7183FF8BD.taxon	description	– 18: on road between tracks, on sandy soil [121233], with admixture of Isopaches bicrenatus, Gymnocolea borealis, Endogemma caespiticia, Scapania obcordata. 19: on road side [121261], mixed with Scapania curta, Solenostoma sphaerocarpum, Plectocolea hyalina.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF861C5F7E271C27FEDB.taxon	description	– 9: sedge- Sphagnum bog at the base of quartzite ridge [121188], in mats with small admixture of Lophozia longiflora, Cephalozia ambigua, Cephaloziella spinigera. 17: dwarf shrubs-sedge-Hypnum-Sphagnum bog at the bottom of mountain [121263] in mat dominated by Dicranum elongatum and admixture of Lophozia longiflora, Aulacomnium turgidum. or mixed with Sphenolobus minutus, Riccardia latifrons, Cephalozia bicuspidata, Blepharostoma trichophyllum subsp. brevirete, Scapania tundra.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F47FF871FD67D171F6BFA0D.taxon	description	– 2 carbonate mossy cliffs in Larix sibirica light forest, on ledges of rocks between rocks [121334], admixture in mats dominated by Mesoptychia heterocolpos, mixed with Tritomaria scitula, Scapania gymnostomophila, Blepharostoma trichophyllum subsp. brevirete, Cyrtomnium hymenophylloides, Brachythecium cirrosum. This poorly known arctic species was previously reported in the Urals from single localities on the western side of mountains (Bezgodov et al., 2003, Konstantinova & Bezgodov, 2006).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861FFC78011F8AFD88.taxon	description	– 5, 7, 11, 14 (4: 775 – 946 m), sp.: on soil in herbaceous tundra on sides of karst hollow, in snowbed communities, in spring fen [121134]. Often in pure mats or mixed with Lophozia wenzelii and Barbilophozia hatcheri.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861FFC7B721FCAFDC2.taxon	description	– 19: on bank of brook, on soil [121255], mixed with Sanionia uncinata.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861FFC7B281F2AFD3D.taxon	description	– 7: dwarf-shrubsedge-green moss- Sphagnum bog on slope [121167], in small hollows, mixed with Gymnocolea inflata Scapania irrigua subsp. rufescens, Warnstorfia sarmentosa.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861FFC7BA71F24FC06.taxon	description	– 8: on carbonate cliffs in tundra [121174], on fine earth between rocks, mixed with Mesoptychia heterocolpos, M. collaris, Blepharostoma trichophyllum subsp. brevirete, Tritomaria scitula, Schistochilopsis opacifolia, Distichium capillaceum, Mnium thomsonii, Encalypta affinis. This species has previously been recorded for the North Urals by Zinovjeva (1973).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861FFC7AEC1CC6FBAB.taxon	description	– 1, 3, 5, 6, 19, 20 (9: 390 – 787 m), fr.: on banks of streams, in seepages, in hollows in swampy forests [121156], in pure mats or mixed with Plectocolea hyalina, P. obovata, Scapania subalpina, S. irrigua, S. paludosa, Pohlia wahlenbergii, P. filum, Calliergonella lindbergii.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861FFC7D511F60FB7D.taxon	description	– 4, 5, 9, 10, 19 (6: 476 – 973 m alt.), fr.: on bare soil on banks of streams and on road sides, on moist loamy spots in dwarf-shrub-sedge-moss tundra [121221]. Always mixed with other hepatics: Nardia geoscyphus, N. japonica, Scapania curta, Solenostoma sphaerocarpum, S. confertissimum, Pellia neesiana, etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861FFC7C671E5DFA5B.taxon	description	– 5: willow (Salix lanata) thickets along stream, on fine earth covered stems of willow [121153], in pure mats (22 / 1 - 1) and mixed with Scapania irrigua, S. paludosa, Pellia neesiana, Calliergonella lindbergii, Sciuro-hypnum latifolium, Warnstorfia exannulata. 19: on bank of brook (64.45512 ° N – 59.75020 ° E; 476 m alt.), on soil, mixed with Scapania subalpina, Philonotis fontana, Chiloscyphus polyanthos (51 / 7 - 2).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861FFC7C811F0EF898.taxon	description	– 1: under alder shrubs along road side: (64.48391 ° N – 59.66510 ° E; 632 m alt.), on soil, in pure mats (43 / 1 - 1) and mixed with Scapania curta, Nardia geoscyphus (43 / 1 - 3), Lophozia wenzelii (43 / 2 - 2). 2: mossy dry brook (64.48383 ° N – 59.66943 ° E; 634 m alt.), on rocks in the bed and on fine earth on bank (71 / 15). This species has previously been reported for Polar Urals as Plectocolea obovata var. minor (Konstantinova & Czernjadieva, 1995).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861FFC7E421E53F862.taxon	description	– 11, 17 (3: 840 – 980), sp.: on loamy spots of bare soil in Betula nana - herbaceous tundra, dwarf shrub-moss-lichen, dwarf shrub – lichen tundra [121271], mixed with Anthelia juratzkana, Cephalozia bicuspidata, Blepharostoma trichophyllum, Nardia geoscyphus, Marsupella sprucei, Gymnomitrion concinnatum, Isopaches bicrenatus, Scapania parvifolia, Diplophyllum obtusifolium.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861C0279B11EECFE6B.taxon	description	– 2, 6, 8, 13 (6: 630 – 926 m), sp.: on fine earth and humus between boulders on carbonate rock outcrops in tundra, on soil on sides of karst hollows, on spots of bare soil in herbaceous tundra, on banks of mossy brooks, on fine earth between rocks in alder thickets [121121]. Always mixed with other bryophytes, particularly with Blepharostoma trichophyllum subsp. brevirete, Mesoptychia heterocolpos, M. gillmanii, M. collaris, Lophocolea minor, Scapania cuspiduligera, Asterella lindenbergiana.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861C0278911821FC8B.taxon	description	– 2: on fine earth on bank of mossy dry brook [121327], in mats dominated by Mesoptychia gillmanii, Scapania curta and admixture Schistochilopsis opacifolia, Pohlia cruda, Bartramia ithyphylla. It is a rare hepatic that always occurs as admixture in mats of other hepatics. It is listed as rare in the Red Data Book of Russia (Bardunov, 2008) and in the Red Data Book of Europe (Schumacker & Martiny, 1995). The species has previously been recorded in the Urals from one locality in the Vishera State Nature reserve (Konstantinova & Bezgodov, 2006).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861C027A711804FBC8.taxon	description	– 7, 9, 10, 11, 13, 14, 15, 16, 17 (13: 790 – 973 m), com.: on soil in Dryas - herbaceous and dwarf shrubs-green moss tundras, in dwarf shrub-sedge-moss bogs, sometimes on fine earth between rocks in rock fields [121137]. In pure mats or mixed with Barbilophozia hatcheri, Schljakovia kunzeana, Lophozia wenzelii. P. pulcherrimum (Weber) Vain. (per.): – 19, 62 (380 – 433 m), sp.: on bases of trees and on decaying wood in mixed coniferous forests [121254], in pure mats or mixed with Barbilophozia barbata, Lophocolea heterophylla, Lophozia silvicola, Lophoziopsis longidens, Brachythecium salebrosum, etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861C027D3219F3FB7D.taxon	description	– 17: dwarf shrub-sedge-Hypnum-Sphagnum bog at the bottom of slope [121263], mixed with Sphenolobus minutus, Neoorthocaulis binsteadii, Cephalozia bicuspidata, Blepharostoma trichophyllum subsp. brevirete, Sphenolobus minutus var. grandis, Scapania tundrae, Loeskypnum badium.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861C027C67199AFA46.taxon	description	– 11: on sides of karst hollow in Dryas - herbaceous tundra [121138], on soil covered fine earth, mixed with Mesoptychia gillmanii, Asterella lindenbergiana. 2: on mossy rocks on bank and in the bed of a dry brook [121320], on fine earth, mixed with Mesoptychia gillmanii, M. heterocolpos, Tritomaria scitula, Blepharostoma trichophyllum subsp. brevirete.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861C027CAC19D8F92D.taxon	description	– 2: on rocks in bed of dry brook [121322] and on fine earth on banks of a brook [121323], mixed with Mesoptychia collaris, M. gillmanii, Scapania cuspiduligera, Tritomaria scitula, Blepharostoma trichophyllum subsp. brevirete, Distichium capillaceum, Preissia quadrata. This species has previously been reported in the Urals from one locality on Mt. Ner-Oika (Konstantinova & Lapshina, 2014).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F46FF861C027FD71942F862.taxon	description	– 2: at the base of carbonate boulders in a subalpine meadow [121313], in shaded niches, on fine earth, mixed with Blepharostoma trichophyllum subsp. brevirete, Tritomaria scitula, Distichium capillaceum, Mesoptychia spp., Encalypta spp. 9: on carbonate fine earth under boulders in rock field (64.49653 N ° – 59.64451 ° E; 973 m alt.), mixed with Blepharostoma trichophyllum subsp. brevirete, Tritomaria scitula, Mnium thomsonii (29 / 4 - 3). The species was previously known from the western side of the Urals, in the Pechoro-Ilych Strict Nature Reserve (Bezgodov et al., 2003).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891FD679B11CC7FE1D.taxon	description	– 1, 2, 11, 13, 19, 21 (7: 476 – 890 m), fr.: on bare soil on banks of streams, on road sides, on spots of moist clay spots in tundra [121123]. Sometimes in pure mats, but more often mixed with Nardia geoscyphus, Plectocolea subelliptica, P. hyalina, Blasia pusilla, Nardia japonica, Scapania obcordata, Pohlia filum, Cephalozia bicuspidata.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891FD678C71C8CFDFB.taxon	description	– 2, 12, 13 (4: 634 – 840 m), sp.: on ledges and under rocks on carbonate rock outcrops, at the bottom of carbonate cliffs, on bank of dry brook, on bare soil in dwarf shrub-herbaceous-green moss tundra near carbonate outcrops [121121]. In pure mats or mixed with Mesoptychia gillmanii, Tritomaria scitula, Saccobasis polymorpha, Scapania gymnostomophila, Lophocolea minor, Preissia quadrata.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891FD67BE11F69FC38.taxon	description	– 13: dwarf-shrub-herbsgreen-moss tundra on steep slope of south-east facing slope [121121], on bare soil, some shoots in mats dominated by Preissia quadrata mixed with Scapania cuspiduligera, Lophocolea minor, Jungermannia pumila. 2: carbonate mossy rock outcrops in larch light forest, on fine earth, mixed with Scapania cuspiduligera [121335] and some shoots in mats dominated by Mesoptychia heterocolpos and admixture of Lophoziopsis pellucida, Tritomaria scitula, Blepharostoma trichophyllum subsp. brevirete, Cyrtomnium hymenophylloides, Brachythecium cirrosum [121334].	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891FD67AA21F2EFB96.taxon	description	– 5: rock field at the bottom of snowfield [121163], on fine earth between boulders, mixed with Anthelia juratzkana, Pseudolophozia sudetica.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891FD67D7C1CCAFB58.taxon	description	– 1, 4, 5, 14, 16 (8: 295 – 810 m), fr.: in boggy willow stands in seepages, on banks of streams, between and under rocks in rock fields at edges of snowfields [121117]. In pure mats or mixed with Schljakovia kunzeana, Lophozia wenzelii, Gymnocolea inflata, Straminergon stramineum, Sanionia uncinata, Aulacomnium palustre, Philonotis fontana.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891FD67D821CDEF9B2.taxon	description	– 4, 7 (4: 713 – 946 m), sp.: in boggy willow thickets in seepages, on banks of streams, between and under rocks in rock fields at edges of snowfields. It has been collected in the same habitats as the typical subspecies [121147]. It has also been collected in hollows in dwarf shrub-sedge-moss bogs mixed with Gymnocolea inflata, Odontoschisma elongata, Warnstorfia sarmentosa (26 / 1) and in snowbeds mixed with Lophozia murmanica, Cephalozia bicuspidata (28 / 3 - 1). This subspecies differs from the typical subspecies in obtuse to rounded lobes and red pigmentation.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891FD67F581C0EF908.taxon	description	– 1: on mossy road side [121119], on soil, mixed with Schistochilopsis opacifolia, Lophoziopsis excisa (03 / 1).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891FD67FF21F8DF898.taxon	description	– 1, 13, 11 (3: 295 – 788 m), sp.: on bare soil on road sides and between tracks, on fine earth on ledge of carbonate outcrops in tundra [121233]. Usually mixed with Scapania curta, Nardia japonica, Gymnocolea borealis, Isopaches bicrenatus, Solenostoma spp.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891FD67E421F43F84D.taxon	description	– 7, 9, 14, 17, 19, 20 (6: 390 – 890 m), sp.: in hollows in sedge- Sphagnum and dwarf-sedge-moss bogs, in seepages in dwarf shrubgreen-moss tundra, on banks of brooks [121190]. In pure mats or mixed with Gymnocolea inflata, Warnstorfia exannulata, W. sarmentosa, Loeskypnum badium.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891C5F79B119D6FE1D.taxon	description	– 5, 6, 7, 14, 19, 20 (7: 380 – 874 m), fr.: on banks of streams, in boggy willow thickets, near seepages [121302]. Usually in pure mats or mixed with Scapania irrigua, Harpanthus flotovianus, Pellia neesiana, Cephalozia bicuspidata, Pohlia wahlenbergii, Philonotis fontana, Rhizomnium magnifolium, Calliergonella lindbergii, Warnstorfia exannulata.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891C5F78C71EE6FDFB.taxon	description	– 4, 6, 9, 10, 17 (6: 688 – 930 m), sp.: between rocks in rock fields, on bare soil on banks of brooks, on spots of bare soil in tundras [121145]. Usually mixed with other hepatics: Lophoziopsis longidens, Cephalozia bicuspidata, Pseudolophozia sudetica, Diplophyllum albicans, Sphenolobus minutus, Mesoptychia spp., etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891C5F7A7119FFFC1B.taxon	description	– 4, 5, 6, 19 (6: 410 – 784 m), sp.: on soil and fine earth on banks and in beds of streams [121257]. Usually mixed with Pellia neesiana, Plectocolea obovata, Pohlia wahlenbergii, Calliergonella lindbergii, Rhizomnium magnifolium.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891C5F7AC11847FAB6.taxon	description	– 3: short grass green-moss subalpine meadow [121310], on soil, mixed with Kiaeria starkei, Sanionia uncinata, Pohlia nutans. This is a mostly arctic species known from most sectors of the Arctic as well as from South Siberia (Damsholt, 2002; Konstantinova, Bakalin et al., 2009). This taxon is not recognized on the species level by some authors (Potemkin, 1999). Finding this species in Northern Ural fills up the gape in distribution in Eurasia and supports its circumpolar distribution. 17: dwarf shrub-sedge- Hypnum-Sphagnum bog at the bottom of slope [121265], mixed with Warnstorfia sarmentosa, Loeskypnum badium, Neoorthocaulis binsteadii, Sphenolobus minutus, Riccardia latifrons, Blepharostoma trichophyllum subsp. brevirete, etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891C5F7CF71928F9D6.taxon	description	– 2, 9, 19, 21 (5: 380 – 980), sp.: on decaying wood in mixed birch-dark coniferous tall herbaceous-green moss forests [121245], in sedge- Sphagnum bogs, on bank of dry brook, on fine earth covered rocks, in pure mats or mixed with Tritomaria exsectiformis, Ptilium cristacastrensis, Sanionia uncinata, Dicranum spp., Lophozia spp., etc.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891C5F7F3C190BF828.taxon	description	– 1, 2, 3, 4, 5, 8 (8: 634 – 926 m), fr.: on soil between rocks in rock fields, on fine earth between rocks on banks of streams under snowfields [121145]. In pure mats or mixed with Lophozia spp., Cephalozia bicuspidata, C. ambigua, Fuscocephaloziopsis pleniceps, Lophoziopsis excisa, Scapania mucronata, Harpanthus flotovianus, Bartramia ithyphylla, Pohlia drummondii. It was once collected on fine earth between rocks on carbonate cliff [121174], mixed with Odontoschisma macounii, Blepharostoma trichophyllum subsp. brevirete, Mesoptychia heterocolpos, Distichium capillaceum, Mnium thomsonii, Encalypta affinis.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891C5F7BE119BDFC8B.taxon	description	– 11: herbaceous tundra, on side of karst hollow (64.50132 ° N – 59.66006 ° E; 830 m alt.), on fine earth in niches, admixture in mats dominated by Mesoptychia gillmanii, M. heterocolpos (09 / 4).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F49FF891C5F7C5C1EF3FA0D.taxon	description	– 19: on fine earth in pool on bank of brook [121256], in pure mats and mixed with Warnstorfia exannulata.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC79B11E53FE6B.taxon	description	– 1, 4, 5, 14, 16, 17, 20 (7: 295 – 840 m), fr.: between boulders in rock fields at the edge of snowfields, among mosses on soil in grass tundras and boggy willow in seepages, on sides of bogs and on banks of streams [121117]. Occurs in pure mats but more often mixed with other bryophytes, more often with Barbilophozia hatcheri, B. lycopodioides, Lophozia wenzelii, Scapania irrigua, Pohlia drummondii, Aulacomnium palustre, Paludella squarrosa, Rhizomnium pseudopunctatum, Sanionia uncinata, Sciuro-hypnum latifolium.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC78911FA4FD18.taxon	description	– 5: rock field near edge of snowfield [121159], on fine earth between boulders mixed with Fuscocephaloziopsis albescens, Sanionia uncinata, Pohliadrummondii. 11: karst hollow in grass tundra [121215], on soil, mixed with Tritomaria scitula, Timmia austriaca, Distichium capillaceum.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC7BC21C8CFC8B.taxon	description	– 10: dwarf shrub-sedge-moss tundra on bank of lake [121221], on moist loamy spots of bare soil, mixed with Nardia geoscyphus, Plectocolea hyalina.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC7A711C3CFC5D.taxon	description	– 17: on fine-earth-loamy spots in dwarf-moss-lichen tundra on slope of mountain on flat area in rock field [121270], mixed with Marsupella sprucei, Prasanthus suecica, Scapania parvifolia. This very small and poorly known species has previously been reported for Polar Urals (Konstantinova & Czernjadieva, 1995).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC7A871CF4FACB.taxon	description	– 10: dwarf-sedge-moss tundra on bank of lake [121227], on moist loamy spots of bare soil in pure mats or mixed with Solenostoma confertissimum, Nardia geoscyphus. 19: on road side, on soil (64.45512 N ° – 59.75020 ° E; 476 m alt.), several shoots among Scapania curta, Plectocolea hyalina, Nardia japonica (52 / 2 - 3).	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC7A871CF4FACB.taxon	discussion	Sphenolobus minutus (Schreb.) Berggr. (gem., per.) – 4, 8, 9, 15, 17, 21 (8: 380 – 926 m), fr.: on fine earth covered boulders in silicate rock fields, on ledges and in niches of carbonate cliffs, on soil in dwarf-shrub-sedge-moss- and dwarfshrubs- Sphagnum bogs, on boulders, on soil and on decaying wood in mixed forests. Occurs in pure mats or mixed with other bryophytes.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC7C311FFFFA46.taxon	description	– 8, 17 (790 – 930 m), sp.: on carbonate cliffs, in rock fields, in dwarf-shrub- Hypnum-Sphagnum bog, [121267], usually mixed with other bryophytes.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC7CAC1F93F9D6.taxon	description	– 4: in rock field at the bottom of mountain slope [121142], on fine earth on huge boulders mixed with Sphenolobus minutus, Sanionia uncinata, Pleurozium schreberi, Dicranum sp. 17: rock field on steep mountain slope [121268], on fine earth between rocks, in pure mats.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC7F3C1F84F967.taxon	description	– 4, 9, 14, 17 (8: 930 – 1030 m), fr.: on siliceous rocks (quartzite, granitoid) in rock field [121182]. It often occurs in pure mats, more seldom mixed with Andreaea rupestris, Pseudolophozia sudetica, Diplophyllum albicans, Kiaeria falcata.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC7E1C1810FEA2.taxon	description	– 3, 4, 5, 8, 9, 10, 12, 13, 17 (11: 644 – 1030 m), fr.: between boulders in rock fields, on fine earth and soil both moist, shaded niches and illuminated sites on carbonate cliffs, in dwarf shrubs-sedge-moss bogs, in willow thickets along temporary brooks [121181]. It occurs in pure mats or mixed with other bryophytes.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881FFC7F8C1F01F8F6.taxon	description	– 19: Birch-spruce tall herbaceous-green moss forest on steep south facing slope [121245], on decaying wood, mixed with Sanionia uncinata, Plagiothecium laetum, Stereodon plicatulus, Dicranum montanum, Tetraphis pellucida.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
466487E16F48FF881C02784B1913FDE7.taxon	description	– 2, 8, 9, 11, 12, 13 (8: 636 – 926 m alt.), fr.: on fine earth and between rocks on ledges on mossy carbonate cliffs in tundra and low growth open forests on soil on sides of karst hollows in herbaceous tundra, along mossy brook near carbonate cliffs [121171]. It sometimes occurs in pure mats but more often mixed with Blepharostoma trichophyllum subsp. brevirete, Mesoptychia heterocolpos, M. gillmanii, Scapania cuspiduligera, Tritomaria quinquedentata, Distichium capillaceum, Mnium thomsonii, Cyrtomnium hymenophylloides, Platydictya jungermannioides, Sanionia uncinata.	en	Konstantinova, Nadezhda A., Lapshina, Elena D. (2017): The hepatics of the Upper Puiva River (Sub-Polar Ural, Khanty-Mansi Autonomous District). Arctoa 26 (1): 35-46, DOI: 10.15298/arctoa.26.03, URL: https://doi.org/10.15298/arctoa.26.03
