taxonID	type	description	language	source
565565B5F7F55D819E1CD1A9862E9E18.taxon	diagnosis	Diagnosis. The only species of the subgenus Holoconops from the Oriental realm that is diagnosed by the following combination of characters: Adult male (Fig. 1 A – I). Gonocoxite wider at base to sub-basal part and narrow at anterior part; gonostylus short and stout, curved at middle and apex with two spines; parameres jointed at middle; posterior portion roughly rectangular, apex of paramere bearing an outwardly directed small projection; aedeagus elongate, rod-shaped, little swollen at apex. Adult female (Fig. 1 J – R). Eyes bare; mandible with 12 – 13 teeth, palpomere III with 4 – 5 well developed protruded capitate sensilla; two unequal spermathecae without any hyaline punctation.	en	Ghosh, Lipika, Pal, Gouri Sankar, Hazra, Niladri (2025): On a new haematophagous species of the genus Leptoconops (Holoconops) Kieffer (Diptera, Ceratopogonidae) from the coastal region of West Bengal, India. Evolutionary Systematics 9 (1): 33-38, DOI: 10.3897/evolsyst.9.138575
565565B5F7F55D819E1CD1A9862E9E18.taxon	description	Description. Male (n = 6). Head. Head dark brown, eyes bare and separated about as wide as 10 facets. All flagellomeres (I – XII) small and globular except terminal flagellomere (XIII); proportion of length of flagellomeres (I – XIII): 18 – 20 (19.2): 09 – 10 (9.66): 08 – 11 (9.6): 08 – 11 (9.83): 08 – 11 (9.66): 08 – 10 (9.83): 07 – 10 (9.5): 08 – 11 (9.83): 08 – 12 (10): 10 – 15 (11.83): 10 – 15 (11.83): 11 – 20 (14.66): 36 – 45 (39.5); AR: 0.93 – 1.05 (0.98) (Fig. 1 A). Maxillary palp with four palpomeres (terminal two fused); palpomere III (Fig. 1 B) longest with a round, shallow sensory pit at apical end, with 4 – 5 protruding capitate sensilla; proportion of length of palpomeres (I – IV): 10 – 12 (11.3): 07 – 10 (8.6): 17 – 22 (20.5): 16 – 20 (18); PR 3.00 – 3.33 (3.1); P / H 0.62 – 0.64 (0.63). Thorax. Dark brown. Scutellum with two bristles. Wing (Fig. 1 C). Covered with numerous microtrichia. WL 0.75 – 0.90 (0.82), WW 0.30 – 0.35 (0.32), CR 0.40 – 0.47 (0.45); stigma pale brown. Legs. Uniformly brown to dark brown, without any banding pattern (Fig. 1 D). HTC (Fig. 1 E) with four spines and a spur; first one smallest, near spur and second one longest; first to fourth ones starting from spur measuring 20 – 25 (22.5), 42.5 – 45 (44.2), 30 – 37 (34), 25 – 32.5 (29.2) respectively. TRI 1.50 – 1.62 (1.56), TRII 1.38 – 1.51 (1.47), TRIII 1.25 – 1.40 (1.34). Abdomen. Dark Brown without any scale. Sternite IX 70 – 82 (75) long and 107 – 112 (109.6) wide, 1.4 times wider than length. Tergite IX 130 – 150 (140.5) long, 110 – 122 (117) wide, almost 1.2 times longer than wide and posterior margin triangular extending up to apex of Gonocoxite; basal margin of tergite IX bearing a outwardly directed, crescent-shaped greatly sclerotised structure with a sharply pointed inwardly directed spine at middle. Genitalia (Figs 1 F – I). Apicolateral process small with 8 – 10 setae. Gonocoxite dark brown (Fig. 1 F); wider at base and narrower in apex, 112 – 127 (119) long, almost twice its greatest breadth, 62 – 67 (63) with stout setae on inner part of gonocoxite. Gonostylus brown (Fig. 1 G), 40 – 52 (46.5) long and 15 – 20 (18.33) wide, usually 0.4 times gonocoxal length, apex of gonostylus with two well-sclerotised equal-sized spines. Parameres (Fig. 1 I) jointed at middle, basal part of each paramere roughly trapezoid in shape, apex of paramere with outwardly directed small projection, distal part each paramere boot-shaped. Aedeagus elongated, rod-shaped, little blunt at apex (Fig. 1 H). Female (n = 3). Overall, they have similar body colouration, and distribution of setae on body (Fig. 1 J); similar shape of maxillary palp with males, hind tibial comb but some usual sexual and other differences are stated below. Head. All antennal flagellomeres rounded except terminal one (elongated) (Fig. 1 K), pale brown to brown, bearing sensilla chaetica (I – X); terminal flagellomere (XI) with sensilla trichodea; proportion of length of flagellomeres (I – XI): 7 – 10 (8.33): 7 – 8 (7.33): 5 – 8 (6.33): 7 – 8 (7.33): 7 – 8 (7.33): 8 – 10 (8.66): 7 – 8 (7.33): 6 – 7 (6.66): 7 – 10 (8): 7 – 10 (8): 19 – 25 (21.33); AR 0.62 – 0.67 (0.64). Mandible armed, with 12 – 13 mandibular teeth (Fig. 1 L). Labrum with five serrations. Clypeus with four setae arranged in a row (Fig. 1 M). Proportion of length of palpomeres (Fig. 1 N) (I – III, IV + V): 8 – 10 (9): 8 – 10 (9): 16 – 25 (21): 10 – 19 (14.66); PR 1.7 – 2.1 (1.9) and P / H 0.80 – 0.88 (0.83). Thorax. Scutum and scutellum dark brown; scutellum with four bristles. Wings (Fig. 1 O). WL 0.77 – 0.90 (0.84), WW 0.36 – 0.42 (0.38) and CR 0.44 – 0.49 (0.46). Legs. TRI 1.71 – 1.82 (1.77), TRII 1.33 – 1.51 (1.41), TRIII 1.40 – 1.48 (1.44); HTC with four spines and a spur (Fig. 1 P); first to fourth spines from spur measuring 20 – 22.5 (21), 37.5 – 42.5 (39), 35 – 38 (36.5), 24 – 26 (25), respectively. Abdomen (Figs 1 Q – R). Dark brown without any scales, with setae sparsely distributed. Genital lamellae pale brown (Fig. 1 Q); little shorter, flat at base, and gradually narrow in apical part, 1.5 times longer than wide and roughly 2.5 times longer than sternite IX; sternite IX 55 – 63 (60) long and 162 – 175 (168) wide, well excavated. Spermathecae (Fig. 1 R) two, oval, almost equal, well sclerotised, without any short neck and hyaline punctation; measuring about 40 – 47 (44.3) × 25 – 30 (27.3) and 36 – 44 (40.3) × 25 – 32 (28.3).	en	Ghosh, Lipika, Pal, Gouri Sankar, Hazra, Niladri (2025): On a new haematophagous species of the genus Leptoconops (Holoconops) Kieffer (Diptera, Ceratopogonidae) from the coastal region of West Bengal, India. Evolutionary Systematics 9 (1): 33-38, DOI: 10.3897/evolsyst.9.138575
565565B5F7F55D819E1CD1A9862E9E18.taxon	etymology	Etymology. The specific name of the new species “ beccus ” derives from the beak-like projection of the apex of parameres.	en	Ghosh, Lipika, Pal, Gouri Sankar, Hazra, Niladri (2025): On a new haematophagous species of the genus Leptoconops (Holoconops) Kieffer (Diptera, Ceratopogonidae) from the coastal region of West Bengal, India. Evolutionary Systematics 9 (1): 33-38, DOI: 10.3897/evolsyst.9.138575
565565B5F7F55D819E1CD1A9862E9E18.taxon	distribution	Distribution. India (West Bengal).	en	Ghosh, Lipika, Pal, Gouri Sankar, Hazra, Niladri (2025): On a new haematophagous species of the genus Leptoconops (Holoconops) Kieffer (Diptera, Ceratopogonidae) from the coastal region of West Bengal, India. Evolutionary Systematics 9 (1): 33-38, DOI: 10.3897/evolsyst.9.138575
