identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8AC11699435157C5B8AA138EB7CD128B.text	8AC11699435157C5B8AA138EB7CD128B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lipolexis depressiceps S. Kim & H. Kim 2025	<div><p>Lipolexis depressiceps S. Kim &amp; H. Kim sp. nov.</p><p>Figs 2 A – M, 6 E</p><p>Type material.</p><p>Holotype: South Korea • ♀; 1549, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.6&amp;materialsCitation.latitude=35.03" title="Search Plazi for locations around (long 126.6/lat 35.03)">Chusan-ri</a>, Ongnyong-myeon, Gwangyang-si, Jeollanam-do; 35°01.8'N, 126°36'E; 10–24 Sep. 2019 by Malaise trap; leg. S. Kim; GenBank: PV 165858 (Ap 23019).</p><p>Diagnosis.</p><p>In some morphological characters (number of antennal segments, maxillary and labial palpomeres, central carinae of dorsal petiole), Lipolexis depressiceps sp. nov. is similar to the other three gracilis group species, L. gracilis, L. pelopsi, and L. sulcata sp. nov .. However, it clearly differs from them in having deeply depressed and anteriorly narrowed occipital carina (partially obscured in dorsal aspect, while distinctly visible in other species), distinctly elongated F 1 and F 2 (length / width ratio = 4.7 and 4.3, while &lt;4.0 in other species), shorter pterostigma / R 1 ratio (0.8, while 1.0– 1.1 in L. gracilis, 0.9–1.1 in L. pelopsi and 1.1 in L. sulcata sp. nov.), and more elongated hind femur (length / width = 5.2, while &lt;4.5 in other species).</p><p>Description.</p><p>Female. Length of body about 3.0 mm (Fig. 2 A). Length of forewing 1.5 mm (Fig. 2 J).</p><p>Head. Width of head 1.5 × its maximum length in dorsal aspect; occipital carina narrow and deeply compressed anteriorly; head 1.3 × wider than mesosoma, with sparse long setae; eye 1.7 × as long as temple in dorsal aspect; ocello-ocular line (OOL) 3.1 × as long as posterior (= lateral) ocellus diameter; ratio of OOL: antero-posterior ocellar line (AOL): postero-ocellar line (POL) = 25: 10: 14 (Fig. 2 G). Eyes oval, sparsely setose; face with densely long setae; width / height ratio = 1.1; tentorial index = 0.4; clypeus oval with 12 long setae; malar space 0.1 × as long as longitudinal eye diameter (Fig. 2 D). Antenna 12 - segmented; F 1 equal to F 2; F 1 and F 2 4.7 × and 4.3 × as long as their width at the middle, respectively; F 1 with three longitudinal placodes and F 2 with four longitudinal placodes (Fig. 2 B, D). Maxillary palp with four palpomeres, labial palp with one palpomere.</p><p>Mesosoma. Mesoscutum with notaulices on anterolateral margin, effaced dorsally (Figs 2 E, 6 E). Dorsal surface smooth, with two rows of 9–10 long setae along the dorsolateral part of mesoscutum; scutellum nearly triangular, with 2 long setae on each side (Fig. 2 E). The precoxal sulcus is weakly impressed on the anterior part of the mesopleuron (Fig. 6 E). Propodeum areolated, areola length / width ratio = 1.1 (Fig. 2 F). Pterostigma 2.4 × as long as width; pterostigma 0.8 × as long as vein R 1 (= metacarpus); r and RS vein extended (Fig. 2 J).</p><p>Leg. Hind femur slender, length / width ratio = 5.2; hind tibia length / width ratio = 11.6; hind femur: hind tibia: hind tarsus = 1: 1.6: 1.8 (Fig. 2 L, K).</p><p>Metasoma. Petiole elongated, 3.1 × as long as wide at spiracles; distinctly prominent central carinae, situated along the dorsal surface of the petiole, start at the anterior part and bifurcate near spiracles; dorsolateral part of petiole posterior of spiracles concave on each side (Fig. 2 H, I). Ovipositor sheath slender and long, wide at base, curved downwards; ratio of ovipositor sheath length / width 3.1 at base and 8.8 at tip (Fig. 2 K).</p><p>Colour. Antenna brown; scape and pedicel yellowish brown, F 1 at least partly yellowish brown. Head and face black, clypeus with mouthparts yellowish brown. Mesosoma and metasoma brown; mesoscutum black; petiole brown. Legs pale brown with dark apices.</p><p>Distribution.</p><p>South Korea.</p><p>Etymology.</p><p>Lipolexis depressiceps sp. nov. is derived from the Latin words depressus (meaning pressed down or flattened) and ceps (meaning ‘ head’), referring to its distinctly anteriorly depressed vertex in dorsal aspect.</p></div>	https://treatment.plazi.org/id/8AC11699435157C5B8AA138EB7CD128B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kim, Sangjin;Sohn, JuHyeong;Kim, Hyojoong	Kim, Sangjin, Sohn, JuHyeong, Kim, Hyojoong (2025): Three new species and a new record of the genus Lipolexis (Hymenoptera, Braconidae, Aphidiinae) from South Korea. ZooKeys 1245: 323-342, DOI: 10.3897/zookeys.1245.138802
BCD4A88758CA5CFFBB042FAB85C26361.text	BCD4A88758CA5CFFBB042FAB85C26361.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lipolexis Forster 1863	<div><p>Genus Lipolexis Förster, 1863</p><p>Gynocryptus Quilis, 1931: 25–30 .</p><p>Type species.</p><p>Lipolexis gracilis Förster, 1863, type locality Germany.</p></div>	https://treatment.plazi.org/id/BCD4A88758CA5CFFBB042FAB85C26361	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kim, Sangjin;Sohn, JuHyeong;Kim, Hyojoong	Kim, Sangjin, Sohn, JuHyeong, Kim, Hyojoong (2025): Three new species and a new record of the genus Lipolexis (Hymenoptera, Braconidae, Aphidiinae) from South Korea. ZooKeys 1245: 323-342, DOI: 10.3897/zookeys.1245.138802
9A82AD3BDA0458AA9342C7D09BF4E667.text	9A82AD3BDA0458AA9342C7D09BF4E667.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lipolexis longipetiolata S. Kim & H. Kim 2025	<div><p>Lipolexis longipetiolata S. Kim &amp; H. Kim sp. nov.</p><p>Figs 4 A – M, 6 G</p><p>Type material.</p><p>Holotype: South Korea • ♀; 434, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.566666&amp;materialsCitation.latitude=35.383335" title="Search Plazi for locations around (long 127.566666/lat 35.383335)">Buun-ri</a>, Sannae-myeon, Namwon-si, Jeonbuk-do; 35°23'N, 127°34'E; 09 Jun. 2022; leg. S. Kim; GenBank: PV 165863 (Ap 220054) . Paratype: South Korea • 1 ♀; 1549, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.6&amp;materialsCitation.latitude=35.03" title="Search Plazi for locations around (long 126.6/lat 35.03)">Chusan-ri</a>, Ongnyong-myeon, Gwangyang-si, Jeollanam-do; 35°01.8'N, 126°36'E; 24 Sep. – 08 Oct. 2019 by malaise trap; leg. S. Kim; GenBank: PV 165862 (AP 23020) • 1 ♀; 716, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.5&amp;materialsCitation.latitude=37.231667" title="Search Plazi for locations around (long 128.5/lat 37.231667)">Donggang-ro</a>, Yeongwol-eup, Yeongwol-gun, Gangwon-do; 37°13.9'N, 128°30'E; 04 – 23 Jun. 2014 by malaise trap; leg. S. Kim; GenBank: PV 165861 (Ap 23008) .</p><p>Diagnosis.</p><p>In some morphological characters (number of antennal segments, maxillary and labial palpomeres, central dorsal carinae of petiole), Lipolexis longipetiolata sp. nov. is similar to other three gracilis group species, L. peregrina, L. takadai, and L. pakistanica . However, it clearly differs from them in having elongated petiole (petiole length / width ratio = 3.4–3.6, while 3.1–3.3 in L. peregrina, 2.8 in L. takadai, and 2.7 in L. pakistanica), elongated F 1 than L. peregrina (length / width ratio = 4.3, while 3.1–3.8 in L. peregrina), stouter tibia (length / maximum width ratio = 8.9, while 11.0 in L. peregrina), and smaller ocellus (OOL / posterior ocellus diameter ratio = 5.0, while 3.0 in L. peregrina).</p><p>Description.</p><p>Female. Length of body about 2.1 mm (Fig. 4 A). Length of forewing 1.2 mm (Fig. 4 J).</p><p>Head. Width of head 1.4 × its maximum length in dorsal aspect; occipital carina gently concave; head 1.3 × wider that mesosoma, with sparse long setae; eye 1.5 × as long as temple in dorsal aspect; OOL 5.0 × as long as posterior ocellus diameter; ratio of OOL: AOL: POL = 29: 10: 10 (Fig. 4 G). Eyes oval, sparsely setose; face with sparse long setae; width / height ratio = 1.1; tentorial index = 0.4; clypeus oval with 8 long setae; malar space 0.2 × as long as longitudinal eye diameter (Fig. 4 D). Antenna 12 - segmented; F 1 equal or longer than F 2 (1.0–1.1); F 1 and F 2 4.3 × and 3.8–4.2 × as long as their width at the middle, respectively; F 1 and F 2 with two longitudinal placodes respectively (Fig. 4 B, C). Maxillary palp with three palpomeres, labial palp with one palpomere.</p><p>Mesosoma. Mesoscutum with notaulices on anterolateral margin, effaced dorsally (Figs 4 E, 6 G). Dorsal surface smooth, with two rows of six long setae along the dorsolateral part of mesoscutum; scutellum nearly triangular, with three long setae on each (Fig. 4 E). The precoxal sulcus is weakly impressed on the anterior part of the mesopleuron (Fig. 6 G). Propodeum areolated, areola length / width ratio = 1.1 (Fig. 4 F). Pterostigma 2.6–3.0 × as long as width; pterostigma 0.8 × as long as vein R 1 (= metacarpus); r and RS vein extended (Fig. 4 J).</p><p>Leg. Hind femur length / width ratio = 4.5; hind tibia length / width ratio = 8.6; hind femur: hind tibia: hind tarsus = 1: 1.5: 1.8 (Fig. 4 M, L).</p><p>Metasoma. Petiole long and slender, wide at base, slightly narrowing towards the apex. 3.4–3.6 × as long as wide at spiracles; distinctly prominent central carinae, situated along the dorsal surface of the petiole, start at the anterior part and bifurcate near spiracles; dorsolateral part of petiole posterior of spiracles concave on each side (Fig. 4 H, I). Ovipositor sheath slender and long, wide at base, curved downwards; ratio of ovipositor sheath width / length = 3.5–4.0 at base and 8.3–8.4 at tip (Fig. 4 K).</p><p>Colour. Antenna brown; scape and pedicel pale brown, F 1 pale brown at least basal 1 / 3 partly yellowish brown, gradually pale brown to apex. Head and face dark brown, clypeus with mouthparts pale brown. Mesosoma dark brown and metasoma brown; mesoscutum dark brown; petiole pale brown. Legs pale brown with dark apices.</p><p>Distribution.</p><p>South Korea.</p><p>Etymology.</p><p>Lipolexis longipetiolata sp. nov. derived from the Latin words longus (meaning long) and petioles (meaning petiole), referring to its distinctly elongated petiole.</p></div>	https://treatment.plazi.org/id/9A82AD3BDA0458AA9342C7D09BF4E667	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kim, Sangjin;Sohn, JuHyeong;Kim, Hyojoong	Kim, Sangjin, Sohn, JuHyeong, Kim, Hyojoong (2025): Three new species and a new record of the genus Lipolexis (Hymenoptera, Braconidae, Aphidiinae) from South Korea. ZooKeys 1245: 323-342, DOI: 10.3897/zookeys.1245.138802
8228C8F27947581AB32E172B0A60521B.text	8228C8F27947581AB32E172B0A60521B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lipolexis peregrina , Tomanovic & Kocic 2020	<div><p>Lipolexis peregrina Tomanović &amp; Kocić, 2020</p><p>Figs 5 A – M, 6 H</p><p>Lipolexis peregrinus Tomanović &amp; Kocić, 2020: 667.</p><p>Specimen examined.</p><p>South Korea • 1 ♀; 666-53, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.65&amp;materialsCitation.latitude=36.616665" title="Search Plazi for locations around (long 127.65/lat 36.616665)">Miwon-ri</a>, Miwon-myeon, Sangdang-gu, Cheongju-si, Chungcheongbuk-do; 36°37'N, 127°39'E; 30 Jun. – 15 Jul. 2015. by malais trap; leg. S. Kim; GenBank: PV 165855 (Ap 23009) • 1 ♀; 852, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.2&amp;materialsCitation.latitude=35.581665" title="Search Plazi for locations around (long 127.2/lat 35.581665)">Hyanggyo-ri</a>, Cheongung-myeon, Imsil-gun, Jeonbuk-do; 35°34.9'N, 127°12'E; 07 Jun. 2019; leg. S. Kim; GenBank: PV 165857 (AP 23017) • 1 ♀; 1549, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.6&amp;materialsCitation.latitude=35.03" title="Search Plazi for locations around (long 126.6/lat 35.03)">Chusan-ri</a>, Ongnyong-myeon, Gwangyang-si, Jeollanam-do; 35°01.8'N, 126°36'E; 24 Sep. – 08 Oct. 2019 by malais trap; leg. S. Kim; GenBank: PV 165856 (Ap 23021) .</p><p>Redescription.</p><p>Female. Length of body about 2.1 mm (Fig. 5 A). Length of forewing 1.3 mm (Fig. 5 J).</p><p>Head. Width of head 1.5 × its maximum length in dorsal aspect; occipital carina broad and moderately concave anteriorly; maximum head length 1.3 × as long as its minimum length; head 1.2 × wider that mesosoma, with sparse long setae; eye 1.7 × as long as temple in dorsal aspect; OOL 3.0 × as long as posterior ocellus diameter; ratio of OOL: AOL: POL = 17: 10: 7 (Fig. 5 G). Eyes oval, sparsely setose; face densely long setose, width / height ratio = 1.3; tentorial index = 0.32; clypeus oval and with 8 setae; malar space 0.2 × as long as longitudinal eye diameter (Fig. 5 D). Antenna 12 - segmented; F 1 subequal to F 2 (F 2 0.9 to 1.1 × as long as F 1); F 1 and F 2 3.1–3.8 × and 3.4–3.6 × as long as their width at the middle, respectively; F 1 and F 2 with two and three longitudinal placodes, respectively (Fig. 5 B, C). Maxillary palp with three palpomeres, labial palp with one palpomere.</p><p>Mesosoma. Mesoscutum with notauli on anterolateral margin, effaced dorsally; The precoxal sulcus is weakly impressed on the anterior part of the mesopleuron (Figs 5 E, 6 H). Dorsal surface smooth, with two rows of four or five long setae along the dorsolateral part of mesoscutum; scutellum nearly triangular, with two long setae on each side (Fig. 5 E). Propodeum areolated, areola length / width ratio = 1.2 (Fig. 5 F). Pterostigma 2.5–2.7 × as long as width; pterostigma 0.8–0.9 × as long as vein R 1 (= metacarpus); r and RS vein extended (Fig. 5 J).</p><p>Leg. Hind femur length / width ratio = 5.0; hind tibia length / width ratio = 11.0; hind femur: hind tibia: hind tarsus = 1: 1.6: 1.7 (Fig. 5 L, M).</p><p>Metasoma. Petiole elongated, 3.1–3.3 × as long as wide at spiracles; distinctly prominent central carinae, situated along the dorsal surface of the petiole, start at the anterior part and bifurcate near spiracles; dorsolateral part of posterior spiracles concave on each side (Fig. 5 H, I). Ovipositor sheath slender and long, wide at base, curved downwards; ratio of ovipositor sheath width / length 3.0 (2.9–3.1) at base and 8.5 (7.9–9.3) at tip (Fig. 5 K).</p><p>Colour. Antenna brown; scape and pedicel brown, F 1 entirely or at least partly pale brown. Head and face dark brown, clypeus with mouthparts pale brown. Mesosoma and metasoma brown; mesoscutum dark brown; petiole pale brown. Legs pale brown with dark apices.</p><p>Distribution.</p><p>Europe (Spain and Slovenia) and Oriental region (China, Japan [from GenBank], and South Korea).</p></div>	https://treatment.plazi.org/id/8228C8F27947581AB32E172B0A60521B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kim, Sangjin;Sohn, JuHyeong;Kim, Hyojoong	Kim, Sangjin, Sohn, JuHyeong, Kim, Hyojoong (2025): Three new species and a new record of the genus Lipolexis (Hymenoptera, Braconidae, Aphidiinae) from South Korea. ZooKeys 1245: 323-342, DOI: 10.3897/zookeys.1245.138802
DD04DE8B91145479818FECA4AFFBB295.text	DD04DE8B91145479818FECA4AFFBB295.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lipolexis sulcata S. Kim & H. Kim 2025	<div><p>Lipolexis sulcata S. Kim &amp; H. Kim sp. nov.</p><p>Figs 3 A – M, 6 F</p><p>Type material.</p><p>Holotype: South Korea • ♀; 1549, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.6&amp;materialsCitation.latitude=35.03" title="Search Plazi for locations around (long 126.6/lat 35.03)">Chusan-ri</a>, Ongnyong-myeon, Gwangyang-si, Jeollanam-do; 35°01.8'N, 126°36'E; 12–27 Oct. 2019 by malais trap; leg. S. Kim; GenBank: PV 165859 (Ap 23018) . Paratype: South korea • 1 ♀; same location as for holotype; 24 Sep. – 08 Oct. 2019 by Malaise trap; leg. S. Kim; GenBank: PV 165860 (Ap 23022) .</p><p>Diagnosis.</p><p>In some morphological characters (number of antennal segments, maxillary palps, and labial palps, central carinae of dorsal petiole), Lipolexis sternaulata sp. nov. is similar to the other three gracilis group species, L. gracilis, L. pelopsi, and L. depressiceps sp. nov. However, it clearly differs from them in having distinctly developed sternaulus (= precoxal sulcus) on the anterior part of the mesopleuron, not reaching the metapleuron (while weakly impressed in other species), shorter tibia than L. gracilis and L. pelopsi (length / maximum length ratio = 8.6, while 14.0 and 17.4 in L. gracilis &amp; L. pelopsi), elongated ovipositor sheath (length / width ratio = 3.2–4.0 at base and 9.7–10.6 at tip, while 3.1 at base and 9.1 at tip in L. pelopsi, 2.9 at base and 9.6 at tip in L. gracilis and 3.1 at base and 8.8 at tip in L. depressiceps sp. nov.), and elongated pterostigma (length / width ratio = 2.6–2.7, while 2.3–2.6 in other species).</p><p>Description.</p><p>Female. Length of body about 2.4 mm (Fig. 3 A). Length of forewing 1.3 mm (Fig. 3 K).</p><p>Head. Width of head 1.5 × its maximum length in dorsal aspect; occipital carina gently concave; head 1.3 × wider than mesosoma, with sparse long setae; eye 1.6 × as long as temple in dorsal aspect; OOL 3.1 × as long as posterior ocellus diameter; ratio of OOL: AOL: POL = 26: 10: 10 (Fig. 3 G). Eyes oval, sparsely setose; face with sparse long setae; width / height ratio = 1.2; densely long setae around antennal socket; tentorial index = 0.3; clypeus oval and with 7 long setae; malar space 0.2 × as long as longitudinal eye diameter (Fig. 3 D). Antenna 12 - segmented; F 1 equal to F 2; F 1 and F 2 3.4–3.6 × and 3.5–3.8 × as long as their width at the middle, respectively; F 1 and F 2 with three longitudinal placodes (Fig. 3 B, C). Maxillary palp with four palpomeres, labial palp with one palpomere.</p><p>Mesosoma. Mesoscutum with notaulices on anterolateral margin, effaced dorsally (Figs 3 E, 6 F). Dorsal surface smooth, with two rows of 6–8 long setae along the dorsolateral part of mesoscutum, respectively; scutellum nearly triangular, with three and four long setae on each side (Fig. 3 E). The precoxal sulcus is present on the anterior part of the mesopleuron, not reaching the metapleuron (Fig. 6 F). Propodeum areolated, areola length / width ratio = 1.3 (Fig. 3 F). Pterostigma 2.6–2.7 × as long as width; Pterostigma 1.1 × as long as vein R 1 (= metacarpus); r and RS vein extended (Fig. 3 J).</p><p>Leg. Hind femur length / width ratio = 4.3; hind tibia length / width ratio = 11.3; hind femur: hind tibia: hind tarsus = 1: 1.6: 1.7 (Fig. 3 L, M).</p><p>Metasoma. Petiole elongated, wide at base, slightly narrowing towards the apex; 3.1–3.3 × as long as wide at spiracles; distinctly prominent central carinae, situated along the dorsal surface of the petiole, and bifurcate near of spiracles; dorsolateral part of posterior spiracles is concave on each side (Fig. 3 H, I). Ovipositor sheath elongated, wide at base, curved downwards; ratio of ovipositor sheath length / width = 3.2–4.0 at base and 9.7–10.6 at tip (Fig. 3 K).</p><p>Colour. Antenna brown; scape and pedicel yellowish brown, F 1 yellowish brown at least basal 1 / 3 partly yellowish brown, gradually brown to apex. Head, face and clypeus with mouthparts pale brown. Mesosoma pale brown and metasoma brown; mesoscutum brown; petiole pale brown. Legs pale brown with dark apices.</p><p>Distribution.</p><p>South Korea.</p><p>Etymology.</p><p>Lipolexis sternaulata sp. nov. is derived from the Latinised form of the precoxal sulcus, referring to its distinctly developed precoxal sulcus on the mesopleuron.</p></div>	https://treatment.plazi.org/id/DD04DE8B91145479818FECA4AFFBB295	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kim, Sangjin;Sohn, JuHyeong;Kim, Hyojoong	Kim, Sangjin, Sohn, JuHyeong, Kim, Hyojoong (2025): Three new species and a new record of the genus Lipolexis (Hymenoptera, Braconidae, Aphidiinae) from South Korea. ZooKeys 1245: 323-342, DOI: 10.3897/zookeys.1245.138802
