identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
13FBBC52F14952379F422F6DA57AC31E.text	13FBBC52F14952379F422F6DA57AC31E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metahelotella alatonotata Huang & Gu 2025	<div><p>Metahelotella alatonotata sp. nov.</p><p>Figs 1 A – G, 2 A – H</p><p>Type materials.</p><p>Holotype: China • ♂; Yunnan, Dehong, Yingjiang County, Mangyun Village; 2023. V – VI; local collector leg.; SYSU . Paratype: China • 1 ♀; Yunnan, Gaoligong Mountains, Baihualing; 2019. V. 22; local collector leg.; SYSU .</p><p>Differential diagnosis.</p><p>This species is similar to Metahelotella schawalleri Lee, 2009, but can be easily distinguished by the combination of the following characters: elytral markings larger, not reaching suture; protibiae with green only at basal one-third; apical notch of tegmen deeper (Fig. 2 A); penis broader at the apex and more distinctly concave at mesal margin (Fig. 2 B) and apical margin of abdominal ventrite V slightly sinuate (Fig. 2 D). This species is also similar to M. semifulva (Ritsema, 1881), but can be distinguished by the different elytral color patterns and male genital structures.</p><p>Description.</p><p>Male. Body (Fig. 1 A, B) flat, surfaces shiny, sides nearly parallel. Eyes black; ventral surface, antennae and legs generally yellowish brown; head, apices of femora and bases of tibiae metallic green with a brownish sheen; tarsi reddish brown; pronotum yellowish brown with a greenish sheen; lateral margin of pronotum and elytra blackish brown; each elytron with one brownish markings from striae 1 to striae 8, reaching apical one-quarter but not reaching the basal margin; remaining areas yellowish green with metallic reflection.</p><p>Head triangular, distinctly constricted behind eyes; anterior margin densely covered by tiny punctures and remaining dorsal surface randomly punctate; ventral surface (Fig. 1 D) densely punctured on gular area, with even pubescence coverage, and genal area with dense elliptical punctures. Mentum (Fig. 1 D) transverse, nearly rectangular, with anterior margin concave. Antenna with 11 antennomeres, including a 3 - segmented club.</p><p>Thorax. Pronotum nearly trapezoidal, widest across posterior one-third; pronotal disc punctate, weakly convex, punctures denser and larger laterally; anterior angles obtuse, moderately protruding anteriad, posterior angles acute; anterior margin convex, slightly concave at middle; lateral margins weakly serrated, posterior margin distinctly bisinuate. Prothoracic hypomera with sparse punctures. Prosternum, meso- and metaventrites smooth, without punctures at middle. Scutellar shield glabrous, nearly semicircular, lateral margins truncate.</p><p>Elytra oblong, widest at basal two-fifths, parallel-sided at basal three-quarters, then gradually narrowing, with apices rounded; each elytron with 10 punctured striae, intervals flat, evenly covered with micro-punctures; lateral margins weakly bordered, apex of each elytron with 1–2 obscure notches (Fig. 1 F); epipleura wide, without punctures.</p><p>Legs. Femora surfaces smooth, weakly dilated. Protibiae (Fig. 1 E) curved, inner side notched and clothed with sparse pubescence on apical one-third. Tarsi simple, tarsomeres I – IV clothed with dense pubescence on ventral surface, tarsomere V longer than the combined length of tarsomeres I – IV.</p><p>Abdomen slightly convergent from abdominal ventrite I towards apex; abdominal ventrite V (Fig. 2 D) with sparse short setae at middle; apical margin slightly sinuate, with dense short setae that are densest at the apex. Abdominal tergite VIII (Fig. 2 C) subquadrate, antero-lateral angles rounded; apical margin truncate, with long setae.</p><p>Male genitalia. Penis (Fig. 2 B) elongate, with rounded lateral margins, narrowing towards the rounded apex, apical orifice elongate, elliptical (from basal fourth to apical fourth), with long subparallel lobes. Tegmen (Fig. 2 A) slender, apical margin with sparse apical setae and a relatively deep notch at middle, basal margin truncate. Internal sac (Fig. 2 E, F) with four sclerites basally, one dorsal sclerite long and slender, strongly curved, hooked at base, one ventral sclerite oblong, two paramedian sclerites short, basally expanded.</p><p>Measurements (in mm). BL 8.02, HL 1.22, HW 1.79, PL 1.86, PW 2.55, EL 5.30, EW 3.02, HL / HW 0.68, PL / PW 0.73, EL / EW 1.74.</p><p>Female (Figs 1 C, G, 2 G, H). Similar to male but with body larger; protibiae straight, without notches; and elytral apices (Fig. 1 G) with 2–3 distinct teeth. Ovipositor short; paraproct and proximal gonocoxite moderately sclerotized; and distal gonocoxite and gonostylus elongated, slender, and well sclerotized (Fig. 2 G, H).</p><p>Measurements (in mm). BL 8.33, HL 1.28, HW 1.84, PL 1.88, PW 2.64, EL 5.45, EW 3.03, HL / HW 0.69, PL / PW 0.71, EL / EW 1.79.</p><p>Etymology.</p><p>The specific epithet is derived from the Latin words alatus and notatus, referring to its wing-like elytral markings.</p><p>Distribution.</p><p>China: Yunnan (Fig. 5).</p></div>	https://treatment.plazi.org/id/13FBBC52F14952379F422F6DA57AC31E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Huang, Yu-Zhou;Gu, Tian-Xuan	Huang, Yu-Zhou, Gu, Tian-Xuan (2025): Taxonomic notes on the little-known genus Metahelotella Kirejtshuk (Coleoptera, Cucujoidea, Helotidae) from southwestern China, with description of a new species and two new records. ZooKeys 1258: 189-198, DOI: 10.3897/zookeys.1258.171034
3EFAE761E3E6571198C012D995360591.text	3EFAE761E3E6571198C012D995360591.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metahelotella immaculata (Ritsema 1891)	<div><p>Metahelotella immaculata (Ritsema, 1891)</p><p>Fig. 3 D, E</p><p>Helota immaculata Ritsema, 1891: 895 (type locality: Myanmar); Wegrzynowicz 2000: 399.</p><p>Metahelotella immaculata: Kirejtshuk 2000: 30; Lee 2009 b: 790.</p><p>Material examined.</p><p>China • 1 ♂; Yunnan, Dehong Prefecture, Yingjiang County, Mangyun Village; 2025. V; Gui-Chang Liu leg.; CYZH . • 2 ♀♀; Yunnan, Dehong Prefecture, Yingjiang County, Nongzhang Town, 890 m; 2025. VII. 19; Gui-Chang Liu leg.; CYZH . India • 1 ♂, 1 ♀; Pedong; 1934; collector unknown; CYZH .</p><p>Diagnosis.</p><p>Body (Fig. 3 D, E) slender and elongate, sides nearly parallel. Head brownish green; pronotum yellowish brown; elytra brownish bronze with greenish sheen; tibiae and apices of femora greenish bronze. Protibia curved in male. Pronotum broad, subrectangular, lateral margins straight, with weak crenulations near anterior angles; elytra oblong, widest at middle.</p><p>Distribution.</p><p>China (new country record): Yunnan (Fig. 5); India; Myanmar; Thailand.</p><p>Remarks.</p><p>The specimens from Yunnan appear morphologically consistent with those from India.</p></div>	https://treatment.plazi.org/id/3EFAE761E3E6571198C012D995360591	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Huang, Yu-Zhou;Gu, Tian-Xuan	Huang, Yu-Zhou, Gu, Tian-Xuan (2025): Taxonomic notes on the little-known genus Metahelotella Kirejtshuk (Coleoptera, Cucujoidea, Helotidae) from southwestern China, with description of a new species and two new records. ZooKeys 1258: 189-198, DOI: 10.3897/zookeys.1258.171034
3D7448BC2E325C8CA6797AB5CB9170CD.text	3D7448BC2E325C8CA6797AB5CB9170CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metahelotella Kirejtshuk 2000	<div><p>Genus Metahelotella Kirejtshuk, 2000</p><p>Metahelotella Kirejtshuk, 2000: 30.</p><p>Type species.</p><p>Helota semifulva Ritsema, 1881: 80, by original designation.</p></div>	https://treatment.plazi.org/id/3D7448BC2E325C8CA6797AB5CB9170CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Huang, Yu-Zhou;Gu, Tian-Xuan	Huang, Yu-Zhou, Gu, Tian-Xuan (2025): Taxonomic notes on the little-known genus Metahelotella Kirejtshuk (Coleoptera, Cucujoidea, Helotidae) from southwestern China, with description of a new species and two new records. ZooKeys 1258: 189-198, DOI: 10.3897/zookeys.1258.171034
9A6F4E482E6957DFA94373F72AF509EE.text	9A6F4E482E6957DFA94373F72AF509EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metahelotella semifulva (Ritsema 1881)	<div><p>Metahelotella semifulva (Ritsema, 1881)</p><p>Figs 3 A – C, 4 A, B</p><p>Helota semifulva Ritsema, 1881: 80 (type locality: Indonesia, Java); Wegrzynowicz 2000: 403.</p><p>Metahelotella semifulva: Kirejtshuk 2000: 30; Lee 2009 b: 786.</p><p>Helota fulvitarsis Ritsema, 1889: 107; Wegrzynowicz 2000: 397; synonymized by Lee 2009 b: 786.</p><p>Metahelotella fulvitarsis: Kirejtshuk 2000: 30.</p><p>Helota difficilis Ritsema, 1891: 896; Wegrzynowicz 2000: 396; synonymized by Lee 2009 b: 786.</p><p>Metahelotella difficilis: Kirejtshuk 2000: 30.</p><p>Material examined.</p><p>China • 1 ♀; Guangxi, Chongzuo City, Longzhou County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.7215&amp;materialsCitation.latitude=22.281" title="Search Plazi for locations around (long 106.7215/lat 22.281)">Binqiao Village</a>, 22.2810 ° N, 106.7215 ° E; 2025. V. 2; local collector leg.; CYZH . • 1 ♂; Yunnan, Dehong Prefecture, Yingjiang County, Mangyun Village; 2025. V; Gui-Chang Liu leg.; CYZH . • 1 ♂; Xizang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.15769&amp;materialsCitation.latitude=29.26578" title="Search Plazi for locations around (long 95.15769/lat 29.26578)">Linzhi City</a>, Medog County; 29.26578 ° N, 95.15769 ° E; alt. 1423.92 m; 2024. VII. 4; Hong-Bin Liang leg.; IZCAS . India • 1 ♂; Pedong; 1934; collector unknown; CYZH .</p><p>Diagnosis.</p><p>Body (Fig. 3 A – C) flat, surfaces shiny, sides nearly parallel. Head metallic green; pronotum and basal three-fifths of elytra yellowish brown, remaining dorsal surface greenish blue; apices of femora and bases of tibiae greenish bronze. Protibiae curved in male, inner side notched. Pronotum trapezoidal, with weak crenulations on lateral margins; elytra oblong, widest at basal one-third.</p><p>Distribution.</p><p>China (new country record): Guangxi, Xizang, Yunnan (Fig. 5); India; Indonesia; Laos; Myanmar; Thailand.</p><p>Remarks.</p><p>This species widely occurs in the Oriental region and is reported from China for the first time. In addition, we received an observation record of M. semifulva (Ritsema, 1881) from Nonggang Natural Reserve, Guangxi, China (Fig. 4 A, B).</p></div>	https://treatment.plazi.org/id/9A6F4E482E6957DFA94373F72AF509EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Huang, Yu-Zhou;Gu, Tian-Xuan	Huang, Yu-Zhou, Gu, Tian-Xuan (2025): Taxonomic notes on the little-known genus Metahelotella Kirejtshuk (Coleoptera, Cucujoidea, Helotidae) from southwestern China, with description of a new species and two new records. ZooKeys 1258: 189-198, DOI: 10.3897/zookeys.1258.171034
