taxonID	type	description	language	source
91109818FFFF120132C578BBFE64F98B.taxon	description	Figure 1	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFF120132C578BBFE64F98B.taxon	discussion	Remarks. Archibald and Cannings (Archibald et al. 2021) created the genus-level collective taxon Dysagrionites to function as a holding bin for fossil odonates referrable to the Dysagrioninae (Cephalozygoptera, Dysagrionidae) that can be clearly separated from species of that subfamily but whose nominal generic affinity is unclear by the preservation and completeness of their currently known specimens. Collective taxa are regulated by the International Commission on Zoological Nomenclature (International Trust for Zoological Nomenclature 1999, for example, articles 42.2.1 and 42.3.1). Archibald et al. (2021) referred three species to the parataxon: Dysagrionites delinei Archibald and Cannings and Dysagrionites sp. A and Dysagrionites sp. B. They erred, however, in that the parataxon definition required that those species are established as members of the Dysagrioninae, and although this strongly appears to be the case, these fragmentary fossils lack enough of the character states that, in combination, firmly place them in the family and, therefore, in this subfamily. Simonsen et al. (2024) subsequently established a diagnosis of Dysagrionidae using wings alone; however, known specimens of Dysagrionites species still lack sufficient morphology for confirmation. We emend the collective taxon definition by adding “ or tentatively so ”: This genus-level collective group name is proposed for any fossil species referable to the Dysagrioninae, or tentatively so, that can be clearly defined at the species level, separating them from other species of the subfamily, but whose orthotaxon generic affinity is unclear by the incomplete preservation of its currently known specimens. Dysagrionites then includes these three species and D. allenbyensis, described here. If our assumption that D. allenbyensis is a dysagrionid is correct, it belongs to the Dysagrioninae, as the origin of its RP 3 – 4 is about two-thirds the distance between the arculus and nodus and IR 2 originates at the subnodus [Petrolestinae: RP 3 – 4 originates more proximally, closer to, or at, the point midway between the arculus and subnodus, and IR 2 originates close to, or on, RP 3 – 4; subfamily diagnoses were summarised by Archibald et al. 2021]. Allenbya holmesae Archibald and Cannings is the only previously known non-anisopteran odonate from the Allenby Formation. It is also tentatively treated as a dysagrionid and cephalozygopteran but is distinct enough to be assigned to a nominal genus (Archibald and Cannings 2022). It is also a fragmentary wing, but in this case, the distal portion is preserved. It is distinct from D. allenbyensis; for example, IR 1 is zigzagged, and the postnodal and postsubnodal crossveins are not aligned.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFD120732C57DCFFD1CFAA5.taxon	description	ZooBank Registration number: urn: lsid: zoobank. org: act: E 1 F 8815 D- 1 BB 4 - 4090 - 99 EE- 08 A 9273 CEE 79 Figure 1	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFD120732C57DCFFD1CFAA5.taxon	materials_examined	Type material. Holotype BBM-P 000020 A and B (part and counterpart). A well-preserved wing with colouration, missing the apical portion. Found by Beverley Burlingame on 2 April 2024 and donated to the Beaty Biodiversity Museum, where it is housed.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFD120732C57DCFFD1CFAA5.taxon	diagnosis	Diagnosis. The wing of Dysagrionites allenbyensis is most like those of Okanopteryx Archibald and Cannings (three species as discussed below) and Stenodiafanus Archibald and Cannings (one species as discussed below). It is distinct from those of Okanopteryx by: CuA space expanded to three cells wide for much of its length from subquadrangle to terminus [O. jeppesenorum Archibald and Cannings maximum four; O. fraseri Archibald and Cannings usually two, sometimes a single row of three]; pigmented fascia wide, extending from nodus to at least 15 and probably 17 cells beyond it [O. macabeensis Archibald and Cannings nine (holotype) to 11]. It is distinct from Stenodiafanus westersidei Archibald and Cannings by dark fascia [S. westersidei: hyaline throughout] and CuA space width [S. westersidei: maximum two cells in equivalent region preserved].	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFD120732C57DCFFD1CFAA5.taxon	description	Description. As above and the following: RP 3 – 4 origin at two-thirds the distance from arculus to subnodus. Postnodal and postsubnodal crossveins almost all aligned. Apex of wing and pterostigma unknown by preservation. No oblique crossvein O. CuA progressively zigzagged in distal quarter; MA zigzagged in distal third (final portion not fully preserved); IR 1 almost linear from its origin three cells from origin of RP- 2; other main veins linear (but distal portions missing). CuA space expanded in the middle, three cells wide for most of its length. Ax 1, Ax 2 preserved, arculus aligned with Ax 2; subnodus distinctly angled, normal obliquity. Membrane hyaline except for dark fascia mid-wing from nodus to the level of at least 15 postnodal crossveins and probably 17 in total. Geologic shear forces may at times somewhat distort fossils at this exposure, and so the apparent narrowness of the wing might be an artefact (Archibald and Cannings 2021 b; Archibald et al. 2023 a).	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFD120732C57DCFFD1CFAA5.taxon	etymology	Etymology. The specific epithet is a toponym referring to the Allenby Formation.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFD120732C57DCFFD1CFAA5.taxon	discussion	Remarks. Dysagrionites allenbyensis is immediately notable for its broad, dark fascia mid-wing. Within the Dysagrionidae, such colouration is found only in species of Okanopteryx (Archibald et al. 2021). The genus is known by three species from Okanagan Highlands localities: O. macabeensis (type species) and O. fraseri from the McAbee locality near Cache Creek, British Columbia, and O. jeppesenorum from Republic, Washington (Archibald et al. 2021). Okanagrion hobani Archibald and Cannings (Dysagrionidae) is also known from both Republic and McAbee but has not been found in the Allenby Formation. The preserved veins of BBM-P 000020 agree with those that, in combination, are partly diagnostic of Okanopteryx: no accessory antenodal crossveins; the subnodus is distinctly angled, with normal obliquity; the origin of IR 2 is rather evenly spaced between RP 1 – 2 and RP 3 – 4; MA is linear to mid-wing, then becomes increasingly zigzagged; MA and MP are close at margin; CuA is increasingly zigzagged from about mid-wing; the postnodal and postsubnodal crossveins are almost all aligned; and there is no intercalary vein in the CuA – A space. However, other character states necessary to assign a wing to the genus in combination with the above cannot be evaluated on this incomplete fossil: the shape and size of the pterostigma; presence of an oblique brace vein; number of cells wide in the costal space distad the pterostigma; and a distinct convergence of RA, RP 1, IR 1, RP 2 at apex. Dysagrionites allenbyensis does not greatly differ from O. macabeensis in particular and could be a regional variant of that species, close in distance (ca. 150 km) and time (age estimates overlap: McAbee: 52.10 ± 0.26 Ma, Vermillion Bluffs: 51.85 ± 0.85 Ma; both Rubino et al. 2021). These preserved character states are also found in Stenodiafanus (see diagnosis in Archibald et al. 2021). The broad, dark fascia of BBM-P 000020 might easily be present convergently in a species of that genus, and the width of the CuA space could vary among its species as it does among those of Okanopteryx. Stenodiafanus westersidei is known only from the Klondike Mountain Formation, about 150 km distant and close in time (estimated ages overlap: Klondike Mountain B 4131 at 51.18 ± 0.09 Ma and Vermillion Bluffs as above; both Rubino et al. 2021). McAbee, the Allenby Formation, and the Klondike Mountain Formation had similar climates and forests (Greenwood et al. 2005; Archibald et al. 2011).	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFA120632DA78ADFBD0FD6A.taxon	description	ZooBank Registration number: rn: lsid: zoobank. org: act: 9540 B 488 - 96 D 6 - 4697 - BCCE- 82 C 7 BAE 16 E 10 Figure 2	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFA120632DA78ADFBD0FD6A.taxon	diagnosis	Diagnosis. Preserved wing portion most similar to that of Dysagrion Scudder by: pterostigma about 6.3 times longer than wide [Dysagrion species: 4.0 – 7.5]; many intercalated linear (or nearly so) supplementary sectors between main veins from RP 1 to probably CuA (see below; cf. figures of Archibald et al. 2021, cited below); IR 1 linear from origin as in Dysagrion fredericii Scudder (see Archibald et al. 2021, fig. 6), slightly zigzagged in Dysagrion pruettae Archibald and Cannings (see Archibald et al. 2021, figs. 17 – 18) and Dysagrion lakesii Scudder (see Archibald et al. 2021, fig. 5), fully zigzagged in Dysagrion packardii Scudder (see Archibald et al. 2021, fig. 4). It differs from wings of Dysagrion by: (1) RA, RP 1, IR 1 subparallel to apex, apex between terminus of RA, RP 1 [Dysagrion: converging at apex, RA at or near apex]; (2) CuA (presumed, see below) linear to termination [Dysagrion: heavily zigzagged]; (3) no brace vein [present in D. pruettae; region poorly or not preserved in other species]; (4) RA – RP 1 space between pterostigma and apex three cells wide in most of distal half, seven at margin [Dysagrion: one]; (5) C – RA space distal to pterostigma two cells wide [Dysagrion: one]; (6) RP 1, IR 1, and RP 2 spaces, each with two linear supplementary sectors, terminate with similar widths [Dysagrion: without supplementary sectors, and RP 1, IR 1 terminate very close, IR 1, RP 2 not]; and (7) subnodus distinctly angled with normal obliquity [Dysagrion: reverse obliquity].	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFA120632DA78ADFBD0FD6A.taxon	description	Description. As for its only species, below.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFA120632DA78ADFBD0FD6A.taxon	etymology	Etymology. The genus name is formed from the Greek - para, “ alongside, ” and Dysagrion, referring to similarities of its wings to those of that genus. Gender, neuter.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFA120632DA78ADFBD0FD6A.taxon	discussion	Remarks. Dysagrion (cf. Archibald et al. 2021, figs. 4 – 6, 16 – 18) is also known from exposure B 4131 of the Klondike Mountain Formation. Its most similar species, D. fredericii, is from the Green River Formation of Wyoming, also in the latter portion of the Ypresian.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFB120632C57FB0FC35FB96.taxon	description	ZooBank Registration number: urn: lsid: zoobank. org: act: AC 2 D 2 DFA- 47 B 2 – 46 C 6 – 8 EE 3 - B 564791 E 4 C 72 Figure 2	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFB120632C57FB0FC35FB96.taxon	materials_examined	Type material. Holotype SR 23 - 005 - 003 A and B (part and counterpart) by monotypy. The well-preserved distal half of a wing found by Emmalin Sosby, 28 May 2023, at exposure B 4131 of the Tom Thumb Tuff Member of the Klondike Mountain Formation in the village of Republic, Washington. Housed in the Stonerose Interpretive Center collections, Republic, Washington.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFB120632C57FB0FC35FB96.taxon	diagnosis	Diagnosis. See genus diagnosis.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFB120632C57FB0FC35FB96.taxon	description	Description. Holotype wing. As in genus diagnosis and: dark, hourglass-shaped fascia in pterostigma region; postnodal, postsubnodal crossveins not aligned. C – RA space apical to pterostigma close to half the width of adjacent RA – RP 1 space.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFB120632C57FB0FC35FB96.taxon	etymology	Etymology. The specific epithet is derived from the surname of Emmalin Sosby, who found and donated the fossil, in recognition of her contribution. Gender, feminine.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFB120432C57987FC44FE4E.taxon	description	Figure 3	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFB120432C57987FC44FE4E.taxon	materials_examined	Material. BBM-P 000021, original PDMA collection number PMF. 2023.001.0031. No counterpart. Distal portions of two wings with poorly preserved fragments of apical-most regions. Found by Beverley Burlingame on 4 August 2023 and donated to the Princeton and District Museum and Archives, then to the Beaty Biodiversity Museum, where it is now housed.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFB120432C57987FC44FE4E.taxon	description	Description. Distal portion of the wing, as in remarks (below), and the following. Preserved portion hyaline. Pterostigma about four times longer than wide; no brace vein; most cells in much of anterior region about twice (or more) as high as wide; many linear (or nearly so) supplementary sectors between main veins obscuring identities of partially preserved main veins posterior to RP 3 – 4. Range and age. Vermilion Bluffs Member of the Allenby Formation about 4 km southwest of the village of Princeton, British Columbia. Ypresian (see above).	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
91109818FFFB120432C57987FC44FE4E.taxon	discussion	Remarks. The wings of BBM-P 000021 are more generalised and difficult to assign to a genus and species than are those of other Allenby Formation non-anisopteran odonates. Some of its morphology, however, is informative: its pterostigma is the general size and shape found in many Okanagan Highlands Dysagrionidae; its postnodal and postsubnodal crossveins are not aligned; its C – RA space distad the pterostigma is more than one cell wide and most likely not more than two; it lacks a brace vein; it has a distinctly angled subnodus of normal obliquity; it has numerous linear supplementary sectors; IR 2, RP 3 – 4, and likely at least some other incompletely preserved main veins are rather strongly curved distally; and there are narrow cells in the posterior region of the wings, many twice as high as wide. Although all preserved distal portions of the BBM-P 000021 wings agree with those of the Dysagrionidae, the character states that might diagnose the species as a member of that family are found in the basal half of their wings, and some Zygoptera also have similar venation, particularly in the distal part of their wings (Archibald et al. 2023 b; Simonsen et al. 2024). Therefore, we tentatively associate BBM-P 000021 with similar odonates that are also known from the Ypresian, especially regionally, pending more complete future specimens. By the above, BBM-P 000021 is similar to the wings of Okanagrion threadgillae Archibald and Cannings (Dysagrionidae) (see Archibald et al. 2021, fig. 20) from Klondike Mountain locality A 0307, about 670 m from B 4131; Primorilestes madseni Rust et al. (Dysagrionidae), from the early Ypresian Fur Formation of Denmark (Rust et al. 2008, fig. 2); and Tynskysagrion brookeae Bechly et al. (Green River Formation, Wyoming, United States of America, placed by Bechly et al. 2020 in Eodysagrioninae, which they treated as a subfamily of Dysagrionidae but was later excluded from that subfamily by Simonsen et al. 2024). We, therefore, treat BBM-P 000021 as Allenby gen. and sp. A, also tentatively in the Dysagrionidae and Cephalozygoptera.	en	Archibald, S. Bruce, Cannings, Robert A. (2024): Three new Odonata species (cf. Cephalozygoptera and cf. Dysagrionidae) from the early Eocene Okanagan Highlands of British Columbia, Canada, and Washington, United States of America. The Canadian Entomologist (e 35) 156: 1-9, DOI: 10.4039/tce.2024.25, URL: https://doi.org/10.4039/tce.2024.25
