taxonID	type	description	language	source
6C5A9CFB9A555034A5E6F71901131082.taxon	diagnosis	Diagnosis. We placed Pristimantis chinguelas sp. nov. in the genus Pristimantis sensu Hedges et al. (2008); Duellman and Lehr (2009) and Padial et al. (2014), subgenus Huicundomantis (sensu Ortega et al. 2022). Pristimantis chinguelas sp. nov. is a medium-sized species distinguished by the following combination of traits: (1) skin on dorsum shagreen without crests or ridges on scapular region, having enlarged dorsolateral tubercles; skin on venter areolate; discoidal fold weak; thoracic fold absent; flanks bearing a row of enlarged tubercles surrounded by numerous large rounded tubercles; (2) tympanic membrane absent, tympanic annulus present, its length about 40 % of the length of eye; supratympanic fold prominent going from the posterior edge of upper eyelid to the level of the insertion of forelimbs; (3) snout short, subacuminate in dorsal view, rounded in profile; canthus rostralis angular; (4) upper eyelid lacking tubercles, about 70 % IOD in males, females unknown; cranial crests absent; (5) dentigerous processes of vomers absent; (6) males with a subgular vocal sac and vocal slits; (7) Finger I slightly shorter than Finger II; discs on fingers broadly expanded, elliptical; (8) fingers lacking lateral fringes; (9) ulnar tubercles absent; (10) heel and tarsus lacking conical tubercles, only low rounded tubercles present; (11) inner metatarsal tubercle elliptical, elevated, about 3 times the size of round outer metatarsal tubercle; supernumerary tubercles low; (12) toes with lateral fringes; basal webbing on feet; Toe V longer than Toe III (disc on Toe III reaches distal edge of the penultimate subarticular tubercle on Toe IV, disc on Toe V reaches the middle to distal edge of the distal subarticular tubercle on Toe IV); (13) in life, dorsum buff yellow to chestnut brown; head having dark supratympanic and canthal stripes; groins, axillae and posterior surface of thighs yellow to dark brown with minute dark flecks; venter cream to yellow with dark flecks; iris gold with fine black reticulations (Fig. 3); (14) average SVL in adult males: 31.0 ± 3.1 mm (25.4 – 34.9 mm; n = 9); females: unknown.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
6C5A9CFB9A555034A5E6F71901131082.taxon	description	Description of the holotype. An adult male (CORBIDI 26772). Measurements (in mm): SVL 34.9; TL 18.5; FL 17.1; HL 12.6; HW 13.6; ED 4.9; TD 2.1; IOD 3.8; EW 2.9; IND 2.8; EN 3.9; FeL 15.9. Head wider than long, narrower than body; snout subacuminate in dorsal view, rounded in profile, lacking papilla or flaps on the tip of the snout (Fig. 4 E); cranial crests absent; nostrils slightly protuberant, directed anterolaterally; canthus rostralis slightly concave in dorsal view, rounded in cross section; loreal region concave; upper eyelid lacking tubercles; tympanic annulus present, visible through the skin, its upper and posterolateral edge concealed by thick, prominent supratympanic fold; tympanic membrane not visible; postrictal tubercles absent. Choanae median, ovoid, non-concealed by palatal shelf of maxilla; dentigerous processes of vomers absent; tongue slightly longer than wide, not notched, posterior half free; vocal slits slightly curved, positioned at posterior half of mouth floor in between tongue and margin of jaw; medium sized vocal sac. Dorsal surfaces of body shagreen; dorsolateral enlarged tubercles present, going from the posterior edge of the upper eyelid to the groin; skin on flanks bearing a row of enlarged tubercles going parallel to the dorsolateral tubercle, below numerous enlarged tubercles cover the rest of the flank; skin on chest and belly areolate, that on throat shagreen, ventral surfaces of limbs smooth, ventral surfaces of thighs coarsely areolate; discoidal fold present, thoracic fold absent. Ulnar tubercles absent; nuptial pads absent; outer palmar tubercle bifid low, as large as ovoid thenar tubercle; subarticular tubercles low, rounded; large supernumerary tubercles at base of fingers, distinct; fingers bearing lateral fringes; Finger I shorter than Finger II; discs on fingers expanded and rounded; pads on fingers surrounded by circumferential grooves on all fingers (Fig. 4 C). Hindlimbs slender; dorsal surfaces of hindlimbs shagreen; posterior surfaces of thighs smooth, ventral surfaces of thighs coarsely areolate; heel lacking tubercle; outer and inner edge of tarsus lacking tubercles; inner metatarsal tubercle elliptical, elevated, twice the size of oval outer metatarsal tubercle; plantar surface with small, low and rounded supernumerary tubercles; subarticular tubercles prominent, rounded; toes bearing lateral fringes; basal webbing between toes III and IV, and IV and V; discs on toes smaller than those on fingers, expanded and rounded; all toes having pads surrounded by circumferential grooves; relative lengths of toes: I <II <III <V <IV; Toe V longer than Toe III (disc on Toe III reaches the distal edge of penultimate subarticular tubercle on Toe IV, disc on Toe V reaches the distal edge of distal subarticular tubercle on Toe IV; Fig. 4 D). Coloration of holotype in life (Fig. 3 A – D). Based on photographs taken in the field. Dorsal surfaces of body and flanks duff yellow covered, skin spaces between tubercles chocolate brown; supratympanic and canthal stripes chocolate brown; groins and anterior surface of thighs creamy yellow with dark flecks; posterior surfaces of thighs, shanks and tarsus orange yellow, some dark flecks present on posterior surface of thighs; venter creamy yellow covered by dark flecks; chest salmon, throat orange, both suffused with brown flecks; ventral surfaces of thighs, shanks, and tarsus yellow; plantar and palmar surfaces creamy white with dense dark flecks; iris gold with thick black reticulations. Coloration of holotype in preservative (Fig. 4 A – E). Dorsal surfaces of body and flanks dark brown fading lighter at the edge of the flanks and venter; gray supratympanic and canthal stripes; groins, anterior, and posterior surfaces of thighs, shanks and tarsus tan; venter, belly and throat creamy yellow suffused with dark flecks; ventral surfaces of limbs creamy yellow; ventral surfaces of fingers and toes cream. Variation. Male CORBIDI 26765 (Fig. 5 C, D) has a single enlarged lateral tubercle instead of a row of tubercles. Skin on dorsum is smoother in males CORBIDI 26766 (Fig. 5 E, F) and 26768 (Fig. 5 G, H). Also, the lateral row of enlarged tubercles is shorter in male CORBIDI 26768, reaching only the middle of the flank. Variation in measurements is given in Table 1 and color variation is shown in Fig. 5. Females unknown. Advertisement call. (Fig. 6) Based on recordings of males CORBIDI 26771 and CORBIDI 26772, the advertisement call consists of one peep-like note (Fig. 6). The average duration of a note is 0.10 s (range 0.06 – 0.15 s). The fundamental and dominant frequencies of the notes are on average 2200.54 Hz (range 2191.00 – 2209.82 Hz), without frequency modulation. Descriptive statistics for bioacoustic variables are shown in Table 2.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
6C5A9CFB9A555034A5E6F71901131082.taxon	distribution	Distribution, natural history and conservation status. Pristimantis chinguelas sp. nov. is only known from the type locality, in Cerro Chinguelas, at 2849 m a. s. l. The type locality is a steep, rocky, open area on the cliff side of the Sapalache – El Carmen road (Fig. 7 A). The vegetation here consists of scattered bushes, montane bamboo (Chusquea spp), some herbs and grass. All individuals were collected at night, calling on leaves, up to 1.8 m. Air temperature was 10 ° C and the humidity was 98 %. During our fieldwork, we heard at least two more groups of calling males, in a steeper area, no more than 100 meters away from the type locality. No other amphibian was recorded in sympatry with the new species. Based on our field data, we estimate the area of occupancy of this species is less than 10 km 2. Nevertheless, the species might occupy adjacent unexplored areas. Information on its population size is unknown. In view of the scarcity of information, and according to the IUCN Red List guidelines (IUCN, 2019) we suggest that P. chinguelas sp. nov. be included in the Data Deficient (DD) category.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
6C5A9CFB9A555034A5E6F71901131082.taxon	etymology	Etymology. The specific epithet is a noun in apposition. It refers to the type locality of the new species, Cerro Chinguelas, a place where a handful of explorers in the late 70 ' s started the scientific research in the Cordillera de Huancabamba.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
ED922C7F56CE56A890BAB3F8CF4EF1BD.taxon	diagnosis	Diagnosis. We placed Pristimantis nunezcortezi sp. nov. in the genus Pristimantis sensu Hedges et al. (2008); Duellman and Lehr (2009) and Padial et al. (2014), subgenus Huicundomantis (sensu Ortega et al. 2022). Pristimantis nunezcortezi sp. nov. is a medium-sized species distinguished by the following combination of traits: (1) skin on dorsum finely shagreened without conspicuous crests, folds or ridges on scapular region; skin on venter coarsely areolate; discoidal fold evident, thoracic fold present; dorsolateral folds absent; flanks areolate; skin on dorsal surfaces of limbs smooth with low rounded tubercles; (2) tympanic membrane present, tympanic annulus present, its length about 42 % of the length of eye; supratympanic fold evident going from the posterior edge of the orbit to the level of the insertion of forelimbs; (3) snout short, subacuminate in dorsal view, rounded in profile; canthus rostralis straight; (4) upper eyelid lacking tubercles, about 50 % IOD in males, females unknown; (5) dentigerous processes of vomers present, low, triangular, moderately separated, posteromedial to choanae; (6) males having vocal slits; (7) Finger I slightly shorter than Finger II (Fig. 9 C); discs on fingers broadly expanded except in finger I, elliptical; (8) fingers lacking lateral fringes; (9) ulnar tubercles rounded, low; (10) heel bearing one conical tubercle, tarsus having low rounded tubercles; (11) inner metatarsal tubercle elliptical, elevated, about 4 times the size of round outer metatarsal tubercle; supernumerary tubercles prominent; (12) toes with lateral fringes; basal webbing on feet; Toe V longer than Toe III (disc on Toe III reaches distal edge of the penultimate subarticular tubercle on Toe IV, disc on Toe V reaches the proximal edge of the distal subarticular tubercle on Toe IV) (Fig. 9 D); (13) in life, dorsum yellowish brown to reddish brown, minute red speckles are visible; light yellow trapezoidal mark on the occipital-scapula region; groins, axillae and posterior surface of thighs having orange-pink bordered black blotches; venter chrome orange; iris gold to orange-yellow on the upper half, copper to scarlet red on the lower half with thin dark reticulations (Figs 8 A, 10 B); (14) SVL of two adult males 31.7 and 38.0 mm; females: unknown.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
ED922C7F56CE56A890BAB3F8CF4EF1BD.taxon	description	Description of the holotype. An adult male (CORBIDI 26786). Measurements (in mm): SVL 38.0; TL 21.8; FL 20.7; HL 14.2; HW 15.7; ED 4.2; TD 2.0; IOD 5.5; EW 2.9; IND 3.1; EN 4.9; FeL 19.2. Head wider than long, narrower than body; snout subacuminate in dorsal view, rounded in profile, lacking papilla or flaps on the tip of the snout (Fig. 9 E); cranial crests absent; nostrils low, directed anterolaterally; canthus rostralis straight in dorsal view, rounded in cross section; loreal region concave; upper eyelid lacking tubercles; tympanic annulus present, visible through the skin, its upper and posterolateral edge concealed by a thick supratympanic fold; tympanic membrane visible; postrictal tubercles absent. Choanae median, ovoid, non-concealed by palatal shelf of maxilla; dentigerous processes of vomers present, triangular, moderately separated, posteromedial to choanae; tongue slightly longer than wide, not notched, posterior half free; vocal slits slightly curved, positioned at posterior half of mouth floor in between tongue and margin of jaw; vocal sac size unknown. Dorsal surfaces of body finely shagreen; skin on flanks areolate; skin on dorsal surface of limbs shagreened bearing low, rounded tubercles; skin on belly coarsely areolate, that on throat weakly areolate, ventral surfaces of limbs areolate, ventral surfaces of thighs coarsely areolate; discoidal fold present, thoracic fold prominent. Ulnar tubercles low, rounded; nuptial pads present; outer palmar tubercle slightly bifid low, as large as ovoid thenar tubercle; subarticular tubercles low, rounded; large, prominent supernumerary tubercles at base of fingers; fingers lacking lateral fringes; Finger I slightly shorter than Finger II; discs on fingers expanded and rounded; pads on fingers surrounded by circumferential grooves on all fingers (Fig. 9 C). Hindlimbs slender; dorsal surfaces of hindlimbs shagreen; posterior surfaces of thighs smooth, ventral surfaces of thighs coarsely areolate; heel having a low conical tubercle; outer edge of tarsus bearing low, rounded tubercles; inner metatarsal tubercle elliptical, elevated, three times the size of oval outer metatarsal tubercle; plantar surface with small, low and rounded supernumerary tubercles; subarticular tubercles prominent, rounded; toes bearing lateral fringes; basal webbing between all toes; discs on toes smaller than those on fingers, expanded and rounded; all toes having pads surrounded by circumferential grooves; relative lengths of toes: I <II <III <V <IV; Toe V longer than Toe III (disc on Toe III reaches the distal edge of penultimate subarticular tubercle on Toe IV, disc on Toe V reaches the proximal edge of distal subarticular tubercle on Toe IV; (Fig. 9 D). Coloration of holotype in life. Based on photographs taken in the field. Dorsal surfaces of body reddish brown, areolate areas on flanks orange, minute red speckles on all dorsum; canthal area yellowish brown; groins, anterior and posterior surface of thighs, and posterior surface of shanks having large orange bordered black blotches; dorsal surfaces of limbs reddish brown; throat, chest, belly and ventral surface of hind limbs chrome orange; plantar and palmar surfaces reddish brown with small black blotches; iris with fine black reticulations, gold on the upper half, copper on the lower half (Fig. 8 A – D). Coloration of holotype in preservative. Dorsal surfaces of body and flanks dark brown fading lighter at the edge of the flanks and venter; groins, anterior, and posterior surfaces of thighs, and shanks black; throat and chest yellow; belly creamy yellow suffused with dark flecks; ventral surfaces of limbs creamy yellow with gray blotches; ventral surfaces of fingers and toes creamy yellow with gray blotches (Fig. 9 A – E). Variation. Measurement’s variation is given in table 1 and color variation in the paratype CORBIDI 26785 is shown in fig. 10.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
ED922C7F56CE56A890BAB3F8CF4EF1BD.taxon	distribution	Distribution, natural history and conservation status. This species is only known from the type locality on the eastern slope of Cerro Chinguelas (Fig. 1), in a small rocky stream (Fig. 7 B) that crosses the Sapalache – El Carmen road, at 2,593 m a. s. l. The streambed can be walked from the road for 200 meters until a rocky wall that blocks access upstream. The habitat is a secondary forest with trees up to 15 m tall, scattered shrubs, tree ferns, bromeliads and herbaceous plants. The soil is wet, covered by leaf litter and some fallen trunks, fungi colonies and lichens. Also, we noticed the presence of farms around this stream. Both individuals CORBIDI 26785 and CORBIDI 26786 were observed at night, during the rainy season, perched on leaves, about 1.8 – 2 m from the ground. No calls were recorded. In addition, we observed P. galdi and P. muscosus in sympatry with the new species. Pristimantis nunezcortezi sp. nov. has presumably a small distribution range, its area of occupancy is far less than 10 km 2 and is only known for a single locality. Since we only recorded two individuals, the species’ distribution extension, population size and conservation status are unknown. Therefore, according to the IUCN Red List guidelines (IUCN, 2019) we suggest that this species should be placed in the Data Deficient (DD) category of the IUCN Red List.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
ED922C7F56CE56A890BAB3F8CF4EF1BD.taxon	etymology	Etymology. the specific epithet, is a patronym for Elio Nuñez-Cortez, a Peruvian ornithologist, in recognition for his contribution to the conservation of the Cordillera de Huancabamba, and for his valuable support and guidance in every one of the expeditions performed by this team. The species epithet is formed from the combination of words of the last name “ Nuñez-Cortez ” as a noun in the genitive case, with the latin suffix “ i ”.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
D620EC3B9A955659B9059E3111E01014.taxon	diagnosis	Diagnosis. Pristimantis yonke sp. nov. is a small species with broad flat head, which may be distinguished by the following combination of traits: (1) skin on dorsum finely tuberculate without crests or ridges on scapular region, incomplete and low dorsolateral fold occasionally present; skin on venter areolate; discoidal fold present; thoracic fold absent; skin on flanks as on dorsum; longitudinal middorsal fold evident; (2) tympanic membrane and tympanic annulus evident, its length about 50 % of the length of eye; supratympanic fold present, going from the distal edge of tympanum to the level of the insertion of forelimbs; (3) snout short, dorsoventrally compressed, acuminate in dorsal view, protruding and inclined posteroventrally in profile, rostral papillae present; canthus rostralis concave; (4) upper eyelid having small, low rounded tubercles, about 70 % IOD in males, 50 % in females; cranial crests absent; (5) dentigerous processes of vomers absent; (6) males with a subgular vocal sac and vocal slits; (7) Finger I slightly shorter than Finger II; discs on fingers broadly expanded, elliptical (Fig. 12 C); (8) fingers having lateral fringes; (9) ulnar tubercles absent; (10) heel and tarsus lacking conical tubercles, only low rounded tubercles present; (11) inner metatarsal tubercle elliptical, elevated, about 3 times the size of round outer metatarsal tubercle; supernumerary tubercles low; (12) toes with lateral fringes; basal webbing on feet; Toe V longer than Toe III (disc on Toe III slightly surpasses distal edge of the penultimate subarticular tubercle on Toe IV, disc on Toe V clearly surpasses the distal edge of the distal subarticular tubercle on Toe IV) (Fig. 12 D); (13) in life, dorsum dark brown to cinnamon brown, with or without dark blotches; flanks same color as dorsum; head having dark supratympanic fold, pale interorbital bar occasionally present; groins, axillae and posterior surface of thighs creamy white to dark brown with or without yellow mottling; throat cream, creamy yellow or yellow with or without dark brown reticulations; belly creamy white or creamy yellow with dark dots, ventral surfaces of thighs yellowish pink or yellow with dark dots; iris gold with thin black reticulations, and a broad horizontal reddish brown streak (Fig. 11 A – D); (14) average SVL in adult males: 19.6 ± 0.3 mm (19.3 – 20.1 mm; n = 4); in females: 30.6 ± 2.4 mm (28.9 – 32.3 mm; n = 2).	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
D620EC3B9A955659B9059E3111E01014.taxon	description	Description of the holotype. An adult male (CORBIDI 26399). Measurements (in mm): SVL 19, 6.3; TL 10.2; FL 8.6; HL 7.3; HW 7.7; ED 2.6; TD 1.2; IOD 2.6; EW 1.9; IND 1.4; EN 2.4; FeL 9.5. Head wider than long, narrower than body; snout acuminate in dorsal view, protruding in profile, having a rostral papilla on the tip of the snout (Fig. 12 E); cranial crests absent; nostrils slightly protuberant, directed anterolaterally; canthus rostralis slightly concave in dorsal view, rounded in cross section; loreal region slightly concave; upper eyelid lacking conical tubercles, only small rounded present; tympanic annulus present, visible through the skin, its upper and posterolateral edge concealed by supratympanic fold; tympanic membrane visible; postrictal tubercles absent. Choanae median, ovoid, non-concealed by palatal shelf of maxilla; dentigerous processes of vomers absent; tongue slightly longer than wide, not notched, posterior half free; vocal slits slightly curved, positioned at posterior half of mouth floor in between tongue and margin of jaw; medium sized vocal sac. Dorsal surfaces of body and flanks finely tuberculate; skin on chest and belly areolate, that on throat shagreen, ventral surfaces of limbs shagreen, ventral surfaces of thighs coarsely areolate; discoidal fold present, thoracic fold absent. Ulnar tubercles absent; nuptial pads absent; outer palmar tubercle ovoid low, as large as ovoid thenar tubercle; subarticular tubercles low, rounded; large supernumerary tubercles at base of fingers, distinct; fingers bearing lateral fringes; Finger I shorter than Finger II; discs on fingers expanded and rounded; pads on fingers surrounded by circumferential grooves on all fingers (Fig. 12 C). Hindlimbs slender; dorsal surfaces of hindlimbs finely tuberculate; posterior surfaces of thighs smooth, ventral surfaces of thighs coarsely areolate; heel bearing low rounded tubercle; outer and inner edge of tarsus lacking tubercles; inner metatarsal tubercle elliptical, elevated, 1.5 × the size of oval outer metatarsal tubercle; plantar surface with small, low and rounded supernumerary tubercles; subarticular tubercles prominent, rounded; toes bearing lateral fringes; basal webbing between toes III and IV, and IV and V; discs on toes slightly smaller than those on fingers, expanded and rounded; all toes having pads surrounded by circumferential grooves; relative lengths of toes: I <II <III <V <IV; Toe V longer than Toe III (disc on Toe III reaches the distal edge of penultimate subarticular tubercle on Toe IV, disc on Toe V barely reaches the distal edge of distal subarticular tubercle on Toe IV); (Fig. 12 D). Coloration of holotype in life. Based on photographs taken in the field. Dorsal surfaces of body and flanks cinnamon brown, two dark brown dorsal stripes that start at dorsal surface of the head, one of them going beyond the level of groins, the other is discontinuous at the level of the groins; supratympanic stripe chocolate brown, canthal area cinnamon brown with irregular pale brown blotches, upper eyelids dark brown; groins and anterior surface of thighs cream; posterior surfaces of thighs, shanks and tarsus cream, some dark brown flecks present on posterior surface of thighs; venter creamy yellow covered by dark spots; chest and throat yellow, both suffused with brown reticulation; ventral surfaces of thighs pinkish yellow with dark spots, shanks and tarsus light yellow; plantar and palmar surfaces light yellow with dense dark brown flecks; iris gold with thick black reticulations and a broad transversal bronze streak (Fig. 11 A ‒ D). Coloration of holotype in preservative. Dorsal surfaces of body and flanks grayish yellow with two dark dorsal stripes that start at the level of the orbits, extending beyond the level of groins; supratympanic stripe gray; groins, anterior, and posterior surfaces of thighs, shanks and tarsus cream; venter, belly and throat creamy yellow suffused with dark flecks; ventral surfaces of limbs creamy yellow; ventral surfaces of fingers and toes gray (Fig. 12 A ‒ E). Variation. Females CORBIDI 26398 and CORBIDI 26929 have a more tubercled dorsal skin, with higher tubercles than rest of the type series. Tympanic annulus is prominent in females CORBIDI 26398, 26929, less visible in males CORBIDI 26399, 26402, 26413 ‒ 14. Supratympanic fold is barely pigmented in female CORBIDI 26929. Middorsal fold is barely visible in male CORBIDI 26402. Variation in measurements is given in table 1 and color variation is shown in Fig. 13.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
D620EC3B9A955659B9059E3111E01014.taxon	distribution	Distribution, natural history and conservation status. This species is known from two localities on the western slopes of the Cordillera de Huancabamba: Laguna Negra trail and Bosque de Ramos, which are 28.8 km (airline) apart, at 2949 and 3000 m a. s. l. respectively (Fig. 1). All individuals were found in primary montane forests where arboreal vegetation consists of Myrcianthes sp. and Podocarpus sp., both trees that reach up to 15 m high. Tree trunks are usually covered with lichens, mosses, and some epiphytic bromeliads. Also, we observed scattered bushes and small clearings inside the forest (Fig. 7 C, D). The soil is wet, muddy in some sections and partially covered by leaf litter. All individuals were found at night, inside bromeliads (1 – 3 m from the ground). Bromeliads where we found CORBIDI 26398 - 99, 26403, 26413 - 14 were close to a rocky stream, whereas CORBIDI 26929 was caught inside a bromeliad in the middle of the forest, without water bodies nearby. Our collections were made during the dry season, and no calling activity was recorded. In addition, we observed P. rhodoplichus in sympatry with P. yonke sp. nov. Based on the extension of our night walks, the sum of the area of occupation in the two localities where we found P. yonke sp. nov. is less than 10 km 2. However, the surroundings remain unexplored, so we do not have accurate information about its geographic distribution, nor about its population size. We suggest that further surveys in the area are needed to collect additional data and assign the species to some threat category. Therefore, following the IUCN guidelines (IUCN, 2019), we suggest this species should be included in the Data Deficient (DD) category of the IUCN Red List.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
D620EC3B9A955659B9059E3111E01014.taxon	etymology	Etymology. The specific epithet is a noun in apposition. It refers to the traditional drink “ yonke ” or “ yonque ”, which is obtained from sugar cane distillation and only drunk by local people from northwestern Andes to keep themselves warm during their journeys through the cold highlands, which may include night camps to get to other villages.	en	Chávez, Germán, Aznaran, Wilmar, Wong, Ivan, Victoriano-Cigüeñas, Karen Y., García-Ayachi, Luis A., Valencia-Málaga, Juan D., Ormeño, Jesús R., Gulman, Michael, Sumiano-Mejía, Ronal, Thompson, Michelle E., Catenazzi, Alessandro (2025): Over the top: Three new species of terrestrial breeding frogs (Anura, Terrarana, Pristimantis) from the highlands of the Cordillera de Huancabamba, northwestern Peru. Evolutionary Systematics 9 (1): 145-166, DOI: 10.3897/evolsyst.9.148522
