taxonID	type	description	language	source
A25F535A3C6BFFFA23B6F9DDFC5FD22D.taxon	description	(Figs. 1, 2, 3 B)	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6BFFFA23B6F9DDFC5FD22D.taxon	materials_examined	Material examined. Holotype: New Caledonia, KANACONO, Stn DW 4705, 22 ° 46 ’ S, 167 ° 19 ’ E, 17 August 2016, 290 – 317 m: F 5.1 mm (MNHN-IU- 2017 - 11657). Paratypes: Papua-New Guinea, KAVIENG, Stn CP 4457, 02 ° 33 ’ S, 150 ° 41 ’ E, 02 September 2014, 133 – 178 m: 1 M 4.2 mm (MNHN-IU- 2014 - 8124). New Caledonia, SPANBIOS, Stn CP 5113, 21 ° 10.1 ’ S, 165 ° 51.1 ’ E, 28 June 2021, 260 – 260 m: 1 F 5.5 mm (MNHN-IU- 2020 - 3563).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6BFFFA23B6F9DDFC5FD22D.taxon	etymology	Etymology. This species is dedicated to the first author’s grandson, Aran, for his enormous love of marine animals.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6BFFFA23B6F9DDFC5FD22D.taxon	description	Description. Carapace: As long as broad; cervical groove slightly distinct, laterally bifurcated; 1 epigastric ridge medially and laterally interrupted; 1 protogastric ridge medially interrupted, not reaching laterally to lateral margin, 1 median short scale between epigastric and protogastric ridges and 1 median scale between protogastric and mesogastric ridges; 1 mesogastric and 1 metagastric ridge both medially and laterally interrupted not continuing to anterior branquial ridges, several minute scales behind metagastric ridge. Posterior branchial region with 5 interrupted transverse ridges, sometimes 1 ridge uninterrupted. Lateral margins convex medially, with 6 – 7 spines: 2 spines in front of and 4 – 5 spines behind anterior cervical groove; first anterolateral, second small, accompanying another spine ventral to between first and second; 2 spines on anterior branchial margin, and 2 – 3 spines on posterior branchial margin, last minute or absent. External orbital limit ending in minute spine; infraorbital margin with 1 - 2 spines. Rostrum broad triangular, 1.4 times as long as broad, length 0.6 postorbital carapace length and breadth 0.4 that of carapace, nearly horizontal in lateral view; distance between distalmost lateral incisions 0.2 distance between proximalmost lateral incisions; dorsal surface with some short setae; lateral margin with 4 deeply incised sharp teeth. Pterygostomian flap rugose, unarmed, anterior margin blunt. Thoracic sternum: As long as broad, lateral extremities gently divergent posteriorly. Sternite III with median shallow notch, anterior margin at each side of notch slightly convex. Sternite IV with anterior part as wide as sternite III, with some minute striae. Sternites IV – VI with a few striae on lateral sides. Sternite III ~ 2.2 × as wide as long; sternite IV nearly 2.7 × as wide as long, and 2.8 × as wide as sternite III. Pleon: Tergites II and III each with 2 transverse ridges on tergite, anterior ridge more distinctly elevated than posterior ridge; tergite of somites IV – V smooth or with short scales; somite VI with 2 medially interrupted ridges, posteromedian margin straight. Males with G 1 and G 2. Eye: Ocular peduncles 1.4 times longer than broad, maximum corneal diameter 0.6 rostrum width. Antennule: Article 1 with 3 well-developed spines, distodorsal larger. Ultimate article with a few short fine setae on distodorsal margin. Antenna: Article 1 with distomesial spine reaching end of article 2. Article 2 with 2 distal spines, distolateral spine subequal or larger than distomesial, reaching or overreaching article 3. Article 3 with distomesial spine. Mxp 3: Ischium with well-developed distal spine on extensor and flexor margins; crista dentata with 12 – 13 denticles. Merus equally long as ischium; flexor margin with 2 subequal strong spines; extensor margin with distal spine. Carpus unarmed. P 1: 2.7 times carapace length, relatively slender, somewhat depressed on palm, more so on fingers. Merus as long as carapace, 1.5 times as long as carpus, with spines arranged roughly in rows, distal spines prominent. Carpus as long as palm, twice longer than broad; dorsal surface with small spines arranged roughly in longitudinal rows; mesial margin with 2 – 3 strong spines. Palm 2.5 times longer than broad, lateral and mesial margins subparallel; spines arranged roughly in rows; dorsolateral row continuing onto lateral margin of fixed finger. Fingers 0.6 times as long as palm, each finger distally with two rows of teeth and spooned; movable finger with row of distal dorsomesial spines. P 2 – 4: Moderately slender. P 2 1.6 times carapace length. Meri successively shorter posteriorly (P 3 merus 0.9 length of P 2 merus, P 4 merus 0.7 length of P 3 merus); P 2 merus 0.8 carapace length, 4 times as long as broad, 1.5 times longer than P 2 propodus. Extensor margin with row of 11 – 12 proximally diminishing spines on P 2 – 3, 1 small distal spine on P 4; ventral margins distally ending in strong spine followed proximally by 1 – 2 spines and several eminences, lateral sides with 2 – 5 small spines on P 4. Carpi with 5 – 6 spines on extensor margin on P 2 - 4; lateral surface with 3 - 4 spines or acute granules sub-paralleling extensor margin; flexor distal margin acute. Propodi 3.5 – 4.0 times as long as broad; extensor margins with 3 – 5 proximal spines; flexor margin with 5 – 6 slender movable spines, terminal spines paired. Dactyli distally ending in well-curved strong spine, length 0.5 – 0.6 that of propodi; flexor margin with 6 – 7 proximally diminishing teeth, terminal tooth prominent. Epipods on P 1 – 2; P 2 epipod reduced. Setae: Three types of setae were observed, (1) short plumose setae, with minute setules along both sides of shaft, densely located on ridges of carapace and pleon (Fig. 3 B); (2) long thick simple setae, sometimes iridescent, sparsely located on carapace, pleon and P 1 – 4; (3) short thick simple setae, sparsely located on carapace, pleon and P 1 – 4. Colour in life. Body colour orange, with median longitudinal whitish broad stripe, from base of rostrum to pleonal somite V; one whitish lateral stripe on each branchial region. P 1 – 4 orange, with diffuse whitish band at distal part of propodi; distal part of P 2 – 4 dactyli reddish. Genetic data. COI and 16 S.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6BFFFA23B6F9DDFC5FD22D.taxon	discussion	Remarks. The new species belongs to the group of species characterized by having all gastric ridges medially and laterally interrupted or scale-like, and the absence of dorsal spines on the branchial regions. The morphologically and genetically closest species is G. lemniscata Macpherson & Robainas-Barcia, 2015, from New Caledonia and Chesterfield Islands. Both species can be distinguished by the following characters: — The scale-like ridges on the meta- and mesogastric areas are arcuate in G. lemniscata, whereas these ridges are transverse and not arcuate in the new species. — The P 2 – 4 meri are 2.5 – 2.7 times as long as broad in G. lemniscata, whereas they are 4 times as long as broad in G. arani. — The short setae on the carapace and pleon are simple, without setules, in G. lemniscata (Fig. 3 A), whereas these setae have short setules along both sides of the shaft in G. arani (Fig. 3 B). Morphologically, the new species is also close to G. poupini Macpherson & Robainas-Barcia, 2015, from the French Polynesia. The main difference between the new species and G. poupini is the presence of epigastric spines in G. poupini, whereas these spines are absent in the new species. Genetically, the new species can be distinguished from G. lemniscata. The new species diverges from G. lemniscata by 2.33 % for 16 S and 10.03 % for COI. No genetic data are available for G. poupini.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6BFFFA23B6F9DDFC5FD22D.taxon	distribution	Distribution. New Caledonia and Papua-New Guinea, 133 – 317 m.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6CFFF423B6FAD5FD5AD7F5.taxon	description	(Figs. 4, 5)	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6CFFF423B6FAD5FD5AD7F5.taxon	materials_examined	Material examined. Holotype: Philippines, Mindoro, Puerto Galera, 13 ° 30.46 ’ N, 120 ° 58.59 ’ E, 28 April 2015, 10 m: ov. F 2.8 mm (UF 43289). Paratypes: same station than holotype: 1 M 2.3 mm (UF 43289 B).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6CFFF423B6FAD5FD5AD7F5.taxon	etymology	Etymology. From the Latin auratus, gold, in reference to the type locality (Mindoro = an alliteration of gold mine in Spanish).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6CFFF423B6FAD5FD5AD7F5.taxon	description	Description. Carapace: Slightly broader than long; cervical groove distinct, laterally bifurcated. Gastric region with some transverse ridges: 1 epigastric ridge unarmed, uninterrupted, medially convex; 2 protogastric ridges, anterior one medially interrupted, without parahepatic spine, posterior ridge short, scale-like, with long thick simple setae; 1 mesogastric ridge medially uninterrupted but not extending laterally to anteriormost of branchial marginal spines; 2 metagastric ridges, anterior one medially interrupted, continuing laterally to anteriorbranchial ridges, posterior ridge short, scale-like. Hepatic region with small spine near first marginal (anterolateral) spine. Anterior branchial region with distinct short ridges. Mid-transverse ridge uninterrupted, preceded by shallow cervical groove, followed by 4 – 5 ridges, 2 of them uninterrupted. Lateral margins well convex medially, with 6 spines: 2 spines in front of and 4 spines behind anterior cervical groove; first anterolateral, well-developed, at same level of lateral limit of orbit, second, small, at midlength between anterolateral spine and anteriormost spine of branchial margin, with small spine ventral to between first and second; 2 spines on anterior branchial region, last small, and 2 spines on posterior branchial margin, last small and obsolescent in some specimens. Lateral limit of orbit unarmed; infraorbital margin with strong spine. Rostrum 1.1 times as long as broad, length 0.5 – 0.6 postorbital carapace length and breadth 0.4 that of carapace; distance between distalmost lateral incisions 0.2 distance between proximalmost lateral incisions; dorsal surface nearly horizontal in lateral view, with numerous small scale-like setose ridges; lateral margin with 4 deeply incised teeth. Pterygostomian flap rugose, unarmed, ridges with short setae, anterior margin acute. Thoracic sternum: 0.8 × as wide as long. Sternite III with median shallow notch. Sternite IV with anterior part wider than sternite III, with some short striae. Sternites IV – VI with a few striae on lateral sides. Sternite III ~ 1.8 × as wide as long; sternite IV nearly 2.7 × as wide as long, and 3.0 × as wide as sternite III. Pleon: Tergites II and III each with 2 transverse uninterrupted ridges on tergite, anterior ridge more distinctly elevated than posterior ridge; tergite of somites IV – VI with posterior minute scale-like ridges; posteromedian margin straight. Males with G 1 and G 2. Eye: Ocular peduncles 1.5 times longer than broad, maximum corneal diameter 0.6 rostrum width. Antennule: Article 1 with 3 well-developed distal spines, distodorsal larger, distomesial spine slightly smaller than others. Ultimate article with tuft of fine setae on distodorsal margin. Antenna: Article 1 with ventral distomesial spine reaching distal margin of article 2. Article 2 with distolateral and distomesial spines subequal, and barely reaching midlength of article 3. Articles 3 and 4 unarmed. Mxp 3: Ischium with flexor and extensor margins ending in blunt angle or with small spine; crista dentata with 20 – 21 denticles. Merus shorter than ischium; flexor margin with 2 subequal spines; extensor margin with small distal spine. Carpus unarmed. P 1: 2.2 times carapace length. Merus as long as carapace length, 1.5 times as long as carpus, with some spines, dorsomesial and distal spines stronger than others. Carpus 0.9 length of palm, 1.4 times as long as broad; dorsal surface with several small spines; mesial margin with row of spines, distal slightly stronger than others. Palm 1.4 times longer than broad, lateral and mesial margins slightly divergent; small spines arranged roughly in dorsal, dorsolateral and dorsomesial rows. Fingers 0.9 times palm length, each finger with two rows of teeth distally spooned; fixed finger with some proximal spines along lateral margin; movable finger unarmed. P 2 – 4: Long and slender. P 2 1.3 times carapace length. Meri successively shorter posteriorly (P 3 merus 0.9 length of P 2 merus, P 4 merus 0.8 length of P 3 merus); P 2 merus 0.5 carapace length, 3.1 times as long as broad, 1.6 times longer than P 2 propodus. Extensor margin with row of 6 – 7 proximally diminishing spines on P 2 – 3, 1 minute distal spine on P 4; ventral margins distally ending in strong spine, lateral sides with 0 – 2 small spines on P 4. Carpi with 3 – 4 spines on extensor margin on P 2 - 3, 0 – 1 distal spine on P 4, distalmost smaller than distal second, sometimes absent; lateral surface with 3 – 4 acute granules sub-paralleling extensor margin; flexor distal margin acute. Propodi 2.8 – 3.0 times as long as broad; extensor margin with 0 – 2 small proximal spines; flexor margin with 5 – 6 slender movable spines, terminal spines paired. Dactyli distally ending in well-curved strong spine, length 0.7 that of propodi; flexor margin with 4 – 5 proximally diminishing teeth, terminal tooth moderately prominent. Epipods present only on P 1. Setae: Three types of setae were observed, (1) short plumose setae, with setules along both sides of shaft, densely located on ridges of carapace and pleon, and scales of P 1 – 4; (2) long thick simple setae, sparsely located on carapace, pleon, and P 1 – 4; (3) short thick simple setae, sparsely located on carapace, pleon and P 1 – 4. Colour in life. Carapace greenish-brownish, distal half of rostrum with numerous white spots. Pleon light brown, with numerous white spots of different size. Corneae whitish with some small red spots. P 1 – 4 reddishbrownish, with whitish distal part of P 1 fingers and P 2 – 4 dactyli. Genetic data. COI.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6CFFF423B6FAD5FD5AD7F5.taxon	discussion	Remarks. Galathea aurata is morphologically and genetically very close to G. melobosis Macpherson & Robainas-Barcia, 2015 from the Mariana Islands, Solomon Islands, Vanuatu, New Caledonia, Chesterfield Islands, Fiji, between 2 and 223 m. Both species can be differentiated by the following aspects: — The rostrum is 1.5 – 1.6 × as long as wide in G. melobosis, instead of as long as wide in the new species. — The distomesial spine of the antennal article 1 exceeds the end of the article 3 in G. melobosis, whereas this spine only reaches the end of the article 2 in G. aurata. — The P 2 – 4 are more slender in G. melobosis than in the new species. The P 2 is 1.9 – 2.0 times the carapace length in G. melobosis, whereas it is about 1.3 times in the new species. The P 2 – 4 propodi are 3.8 – 4.7 times as long as broad in G. melobosis, whereas they are 2.8 – 3.0 times as long as broad in the new species. — Genetically the new species diverges from G. melobosis by 6.70 % for COI.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C6CFFF423B6FAD5FD5AD7F5.taxon	distribution	Distribution. Philippines, Mindoro, at 10 m.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C63FFF023B6FA6EFD86D6F8.taxon	description	(Figs. 3 C, 6)	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C63FFF023B6FA6EFD86D6F8.taxon	materials_examined	Material examined. Holotype: New Caledonia. KOUMAC, Stn KL 37, 20 ° 49.9 ’ S, 164 ° 16.7 ’ E, 81 m, 17 September 2018: M 2.5 mm (MNHN-IU- 2014 - 20126).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C63FFF023B6FA6EFD86D6F8.taxon	etymology	Etymology. From the Latin cornis, thorny, and bi, two, in reference to the pair of protogastric and cardiac spines on the carapace dorsal surface	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C63FFF023B6FA6EFD86D6F8.taxon	description	Description. Carapace: As long as broad; anterior and posterior cervical grooves distinct; dorsal surface with scale-like and interrupted ridges in all regions; mid-transverse ridge laterally interrupted and not scale-like, preceded by distinct cervical groove; transverse groove before cardiac spines; ridges not densely setose, with short simple setae (sometimes with a few short setules) and with some long and median thick plumose setae on protogastric and cardiac ridges. Epigastric region with 4 small spines; 2 submedian protogastric spines, and 2 parahepatic spines on each side; 2 postcervical spines on each side of mid-transverse ridge, 1 branchial spine on each side, behind mid-transverse ridge and 2 median cardiac spines. Lateral margins slightly convex, with 7 spines: 2 spines in front of and 5 strong spines behind anterior cervical groove; first anterolateral, well-developed, distinctly posterior to level of lateral limit of orbit; second small, situated at midlength between anterolateral spine and anterior cervical groove, accompanying another small spine ventral to between first and second; 2 spines on anterior branchial region, and 3 spines on posterior branchial margin. Outer orbital angle acute; infra-orbital margin with 1 or 2 spines. Rostrum 1.8 times as long as broad, length 0.6 that of, breadth 0.3 that of carapace; distance between distalmost lateral incisions 0.3 distance between proximalmost lateral incisions; dorsal surface nearly horizontal in lateral view, with minute setiferous ridges; lateral margin with 4 sharp teeth. Pterygostomian flap rugose with sparse setae, anteriorly rounded; some granules on upper margin near linea anomurica. Thoracic sternum: 1.2 × as long as wide. Sternite III with median shallow notch. Sternite IV as wide as following sternites, with anterior part as wide than sternite III, with some short striae. Sternites IV – VI with a few striae. Sternite III ~ 2.1 × as wide as long; sternite IV nearly 2.1 × as wide as long, and 3.2 × as wide as sternite III. Pleon: Tergites II and III each with 2 transverse uninterrupted ridges on tergite, anterior ridge more distinctly elevated than posterior ridge; tergite of somites IV and V with short median ridge; somite VI with 2 scale-like ridges, posteromedian margin straight. Males with G 1 and G 2. Eye: Ocular peduncles 1.8 times longer than broad, maximum corneal diameter 0.8 rostrum width. Antennule: Article 1 with 3 distal spines, 2 well-developed spines, distodorsal larger than others; distomesial spine small but distinct; additional small spine on lateral margin. Ultimate article with a few short setae not in tuft on distodorsal margin. Antenna: Article 1 with depressed ventral distomesial process clearly not reaching distal margin of article 2. Article 2 with small distomesial and distolateral spines, not reaching midlength of article 3. Article 3 with small distomesial spine. Article 4 unarmed. Mxp 3: Ischium with spine on extensor and flexor distal margins; crista dentata with 20 – 21 denticles. Merus subequal in length to ischium, with 3 subequal spines on flexor margin; extensor margin with small distal spine. Carpus with several spines along extensor margin. P 1: Lost. P 2 and 4: Relatively short, somewhat compressed, sparsely setose with some simple setae and some scattered thick long plumose setae on all articles. Meri successively shorter posteriorly (P 4 merus 0.7 length of P 2 merus), equally broad; P 2 merus 0.8 carapace length, 4.0 times as long as broad, 1.3 times longer than P 2 propodus; P 4 merus 2.7 times as long as broad, as long as P 4 propodus. Extensor margins of P 2 merus with row of 6 proximally diminishing spines, 2 – 3 spines on P 4; lateral surface with 0 – 2 small spines; ventrolateral margins ending in strong terminal spine proximally followed by smaller spine; flexomesial margin with 4 spines on P 2, unarmed on P 4. Carpi each with 3 – 4 spines on extensor margin on P 2 and P 4; lateral surface with row of 2 – 4 small spines or acute granules paralleling extensor row; flexor distal margins with very small distal spine. Propodi 4.5 times as long as broad; extensor margin with 2 – 3 proximal spines on P 2, unarmed on P 4; flexor margin with 5 – 6 movable spines. Dactyli subequal in length, 0.7 length of propodi, ending in incurved, strong, sharp spine; flexor margin with 5 – 6 successively diminishing teeth, terminal tooth prominent. Epipods present on P 1, absent on P 2 – 3. Setae: Two types of setae were observed, (1) moderately short simple setae, not dense, sometimes with a few minute setules along both sides of first half of shaft, located on ridges of carapace and pleon, and scales of P 1 – 4; (2) long thick plumose setae, with setules along one side of shaft, distally forming a tuft (Fig. 3 C), sparsely located on carapace, around protogastric and cardiac spines, pleon, and P 2 – 4. Considering the small size of the unique specimen, with both P 1 lost, the type of setae should be considered as provisional. Genetic data. COI and 16 S.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C63FFF023B6FA6EFD86D6F8.taxon	discussion	Remarks. The new species belongs to the group of species having the carapace with median protogastric and cardiac spines and the antennular article 1 with minute distomesial spine. This group contains G. sentosa Macpherson & Robainas-Barcia, 2015 from Wallis and Futuna area and G. robusta Baba, 1990 from Madagascar and La Reunion from which it can be distinguished by the following characters: — G. robusta has the metagastric region armed with 2 median spines, whereas this region is unarmed in G. bicornis. Furthermore, each anterior branchial region has 1 spine in G. robusta, whereas this spine is absent in the new species. — G. sentosa has the cardiac region with a transverse row of 4 – 6 spines, whereas there are only 2 median spines in the new species. Furthermore, the new species has 1 dorso-branchial spine on each side, behind the mid-transverse ridge, whereas this spine is absent in G. sentosa. Genetically, G. bicornis shows divergences in COI of 11.12 % from G. robusta, and 10.54 % from G. sentosa. The new species is also relatively close to G. pascualae Macpherson & Robainas-Barcia, 2015, from New Caledonia and Vanuatu (10.91 %). However, both species are easily differentiated by the absence of median spines on the metagastric and cardiac regions in G. pascualae. There is no 16 S data available for G. sentosa, but G. bicornis sp. nov. shows genetic divergence of 3.93 % from G. pascualae and 14.02 % from G. robusta.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C63FFF023B6FA6EFD86D6F8.taxon	distribution	Distribution. New Caledonia, 81 m.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C66FFF223B6FDADFD68D155.taxon	description	(Fig. 7)	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C66FFF223B6FDADFD68D155.taxon	materials_examined	Material examined. Holotype: New Caledonia. KOUMAC, Stn KB 641, 20 ° 47.9 ’ S, 164 ° 16.4 ’ E, 14 / 11 / 19, 36 m: ov. F 2.8 mm (MNHN-IU- 2014 - 20098). Paratypes: KOUMAC 2019, Stn KL 09, 20 ° 37.3 ’ S, 164 ° 08.3 ’ E, 15 / 09 / 2018, 67 m: 1 ov. F 3.6 mm (MNHNIU- 2021 - 4277), 19 M 2.0 – 3.5 mm, 8 ov. F 2.5 – 3.7 mm (MNHN-IU- 2014 - 20100), 4 M 2.2 – 3.4 mm, 9 ov. F 2.6 – 3.5 mm, 5 F 2.0 – 2.3 mm (MNHN-IU- 2021 - 5813 to 5816, 5819), 1 ov. F 2.2 mm (MNHN-IU- 2014 - 20099). — Stn KL 22, 20 ° 26.8 ’ S, 163 ° 58.3 ’ E, 16 / 09 / 2018, 47 m: 1 M 4.0 mm (MNHN-IU- 2014 - 20155), 4 M 3.0 – 3.7 mm (MNHN-IU- 2021 - 4291), 1 ov. F 3.0 mm (MNHN-IU- 2021 - 5817).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C66FFF223B6FDADFD68D155.taxon	etymology	Etymology. The name dianthus, pink, refers to the colour of the species. The name is considered as a substantive in apposition.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C66FFF223B6FDADFD68D155.taxon	description	Description. Carapace: As long as broad; cervical groove distinct, laterally bifurcated. Gastric region with 6 transverse ridges: 1 epigastric ridge medially interrupted, unarmed; 2 protogastric ridges, anterior ridge uninterrupted, extending laterally to second lateral spine, convex medially, without parahepatic spine, posterior ridge short with some long thick plumose setae; 1 mesogastric ridge uninterrupted, not extending laterally to anteriormost of branchial marginal spines; 2 metagastric ridges, anterior ridge uninterrupted, sometimes extending laterally to third lateral branchial spine, posterior ridge short. Anterior branchial region with distinct ridges. Mid-transverse ridge uninterrupted, preceded by shallow cervical groove, followed by 5 transverse ridges, 2 – 3 of them uninterrupted. Lateral margins convex medially, with 8 spines: 2 spines in front of and 6 spines behind anterior cervical groove; first anterolateral, well-developed, at level of lateral limit of orbit; second, small, at midlength between anterolateral spine and anteriormost spine of branchial margin, with accompanying strong spine ventral to between first and second; 3 spines on anterior branchial region, and 3 spines on posterior branchial margin, last small. Small spine at lateral limit of orbit; infraorbital margin with strong spine. Rostrum 1.2 – 1.3 as long as broad, length 0.5 postorbital carapace length and breadth 0.4 that of carapace; distance between distalmost lateral incisions 0.25 distance between proximalmost lateral incisions; dorsal surface nearly horizontal in lateral view, with numerous setae; lateral margin with 4 deeply incised sharp teeth. Pterygostomian flap rugose, with spine on anterior ridge, ridges with short setae, anterior margin ending in acute angle. Thoracic sternum: 0.8 times as long as broad, lateral extremities gently divergent posteriorly. Sternite III with median shallow notch and some small spines on anterior margin. Sternite IV with anterior part as wide as sternite III, with some minute striae. Sternites IV – VI with a few striae on lateral sides. Sternite III ~ 1.9 × as wide as long; sternite IV nearly 3.0 × as wide as long, and 2.8 × as wide as sternite III. Pleon: Tergites II and III each with 3 transverse ridges on tergite, anterior ridge more distinctly elevated than posterior ridge; tergite of somite V smooth; somite VI with 2 medially interrupted ridges, posteromedian margin straight. Males with G 1 and G 2. Eye: Ocular peduncles 1.5 times longer than broad, maximum corneal diameter 0.6 rostrum width. Antennule: Article 1 with 2 spines; well-developed distodorsal and distolateral spines, distodorsal larger; distomesial spine minute or obsolescent. Ultimate article with a few short fine setae, not in tuft on distodorsal margin. Antenna: Article 1 with ventral distomesial spine reaching distal margin of article 2. Article 2 with 2 well-developed distal spines, distolateral spine shorter than distomesial and reaching midlength of article 3. Articles 3 and 4 unarmed. Mxp 3: Ischium with well-developed spine on flexor distal margin; extensor margin ending in acute angle; crista dentata with 20 – 21 denticles. Merus as long as ischium; flexor margin with 2 subequal spines; extensor margin with minute spines. Carpus unarmed. P 1: 2.3 – 2.6 times carapace length. Merus 0.8 times length of carapace, 1.4 – 1.6 times as long as carpus, with spines arranged roughly in rows, dorsomesial spines stronger; distal spines prominent. Carpus 0.7 – 0.9 length of palm, 1.3 – 1.5 times as long as broad; dorsal surface with some spines; mesial margin with row of well-developed spines. Palm 1.5 – 1.6 times longer than broad, lateral and mesial margins subparallel; a few small spines arranged roughly in dorsolateral and dorsomesial rows, a few small spines scattered on dorsal side; dorsolateral spines continuing along fixed finger. Fingers 0.7 – 0.9 length of palm, each finger with two rows of teeth distally spooned; movable finger with 2 – 3 spines along proximal half of mesial margin. P 2 – 4: Moderately slender. P 2 1.5 times carapace length. Meri successively shorter posteriorly (P 3 merus 0.8 length of P 2 merus, P 4 merus 0.9 length of P 3 merus); P 2 merus 0.5 carapace length, 3.2 times as long as broad, 1.4 times longer than P 2 propodus; P 3 merus 2.1 times longer than broad, 1.1 times longer than P 3 propodus; P 4 merus 2.4 times as long as broad, 0.9 length of P 4 propodus. Extensor margin of P 2 – 3 meri with row of 5 – 7 proximally diminishing spines, and 1 – 2 spines on P 4; ventral margins distally ending in strong spine followed proximally by several tubercles or eminences; lateral sides unarmed on P 2 – 3, with 2 – 3 small spines on P 4. Carpi with 1 – 3 spines on extensor margin on P 2 – 4; lateral surface with several acute granules sub-paralleling extensor margin; flexor distal margin ending in acute or blunt angle. P 2 – 4 propodi 3.5 – 4.5 times as long as broad; extensor margin with 0 – 2 proximal spines; flexor margin with 6 – 7 slender movable spines on P 2 – 4, terminal spines paired. Dactyli distally ending in well-curved strong spine, length 0.6 – 0.7 that of propodi; flexor margin with 4 – 5 proximally diminishing teeth, terminal tooth prominent. Epipods on P 1. Setae: Four types of setae were observed, (1) short plumose setae, with setules along both sides of shaft, densely located on ridges of carapace and pleon, and scales of P 1 – 4; (2) long thick simple setae, sparsely located on pleon and P 1 – 4; (3) short thick simple setae, sparsely located on pleon and P 1 – 4; (4) long thick plumose setae, with long setules along one side of shaft, sparsely located on P 1 – 4. Colour in life. Ground colour of carapace and pleon orange or light reddish, with minute red spots, more numerous on the anterior part of carapace and pereiopods. Posterior part of carapace and pleon with some dark spots. Genetic data. COI and 16 S.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C66FFF223B6FDADFD68D155.taxon	discussion	Remarks. The morphologically and genetically closest relative of the new species is G. cymothoe Macpherson & Robainas-Barcia, 2015, from Vanuatu, New Caledonia, and Chesterfield Islands, from which G. dianthus can be distinguished by the following characters: — The epigastric and parahepatic spines on the carapace are present in G. cymothoe, whereas these spines are absent in the new species. — The rostrum is narrower in G. cymothoe (nearly 1.8 times longer than broad) than in G. dianthus (1.2 times longer than broad). — The genetic distances between the two species are 11.64 % and 4.73 % for COI and 16 S, respectively.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C66FFF223B6FDADFD68D155.taxon	distribution	Distribution. New Caledonia, 36 – 67 m.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C64FFED23B6F93DFCDED3C9.taxon	description	(Fig. 8)	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C64FFED23B6F93DFCDED3C9.taxon	materials_examined	Material examined. Holotype: New Caledonia. SPANBIOS, Stn DW 5230, 18 ° 58.7 ’ S, 163 ° 19.4 ’ E, 21 July 2021, 364 – 368 m: ov. F 5.0 mm (MNHN-IU- 2021 - 9124). Paratypes: New Caledonia. SPANBIOS, Stn CP 5113, 21 ° 10.1 ’ S, 165 ° 51.1 ’ E, 28 June 2021, 260 – 260 m: 1 F 6.0 mm (MNHN-IU- 2021 - 5974). — Stn CP 5131, 20 ° 34 ’ S, 164 ° 58.7 ’ E, 01 July 2021, 354 – 368 m: 1 M 5.8 mm (MNHN-IU- 2021 - 5996). — Stn CP 5132, 20 ° 33.8 ’ S, 164 ° 58.4 ’ E, 01 July 2021, 398 – 431 m: 4 ov. F 4.7 - 6.8 mm (MNHN-IU- 2016 - 9606). — Stn CP 5200, 20 ° 48.7 ’ S, 165 ° 26.1 ’ E, 15 July 2021, 306 – 452 m: 1 M 4.8 mm (MNHNIU- 2021 - 5972), 1 M 3.4 mm (MNHN-IU- 2021 - 1916). — Stn CP 5266, 20 ° 27.9 ’ S, 164 ° 50.1 ’ E, 26 July 2021, 414 – 411 m: 1 M 5.2 mm (MNHN-IU- 2021 - 4444). — Stn DR 5283, 21 ° 24.5 ’ S, 166 ° 10.2 ’ E, 29 July 2021, 365 – 252 m: 1 M 4.3 mm (MNHN-IU- 2021 - 5970). Vanuatu. MUSORSTOM 8, Stn CP 1087, 15 ° 10.18 ’ S, 167 ° 14.07 ’ E, 6 Octuber 1994, 394 – 421 m: 1 ov. F 5.0 mm (MNHN-IU- 2013 - 15935).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C64FFED23B6F93DFCDED3C9.taxon	etymology	Etymology. From the Latin gibbus, humped, in reference to the hump on the protogastric area.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C64FFED23B6F93DFCDED3C9.taxon	description	Description. Carapace: As long as broad; cervical groove distinct, laterally bifurcated; most ridges on gastric region interrupted or scattered scale-like ridges; epigastric region with 10 – 15 spines; 3 – 6 small hepatic spines on each side; 1 – 4 small parahepatic spines on each side lateral to anterior protogastric ridge; anterior protogastric ridge medially convex, with distinct elevation with 2 small median spines and 2 – 4 long thick iridescent setae; anterior branchial region unarmed; one small postcervical spine on each side, rarely obsolescent. Mid-transverse ridge uninterrupted, preceded by shallow cervical groove, followed by 5 – 6 ridges. Lateral margins slightly convex medially, with 8 spines: 2 spines in front of and 6 spines behind anterior cervical groove; first anterolateral, well-developed, at level of orbit, second small but distinct, located at midlength between first spine and anterior cervical groove, with 2 spines ventral to between first and second; 3 spines on anterior branchial margin, and 3 spines on posterior branchial margin. Small outer orbital spine; infraorbital margin with strong spine. Rostrum lanceolate, 1.5 – 1.6 times as long as broad, length 0.5 that of, breadth 0.3 that of carapace; distance between distalmost lateral incisions 0.25 distance between proximalmost lateral incisions; dorsal surface nearly horizontal in lateral view, with some unirramous setae; lateral margin with 4 deeply incised sharp teeth. Pterygostomian flap rugose, with sparse short setae, anterior margin bluntly angular. Thoracic sternum: 0.8 × as wide as long. Sternite III with median shallow notch. Sternite IV with anterior part as wide as sternite III, with some short striae. Sternites IV – VI with a few striae. Sternite III ~ 2.3 × as wide as long; sternite IV nearly 2.8 × as wide as long, and 3.0 × as wide as sternite III. Pleon: Tergites II and III each with 4 transverse uninterrupted ridges, anterior ridge more distinctly elevated than posterior ridge; tergite of somites IV and V with 3 ridges; somite VI with 2 scale-like ridges, posteromedian margin straight. Males with G 1 and G 2. Eye: Ocular peduncles 1.5 times longer than broad, maximum corneal diameter 0.5 rostrum width. Antennule: Article 1 with 2 well-developed distodorsal and distolateral spines, distodorsal larger; distomesial spine minute; 1 – 2 small spines on lateral margin. Ultimate article with a few long fine setae not in tuft on distodorsal margin. Antenna: Article 1 with distomesial spine slightly exceeding distal margin of article 2. Article 2 with 2 well-developed subequal distal spines or distolateral spine slightly longer than distomesial and not reaching end of article 3, sometimes 1 additional small mesial spine. Articles 3 and 4 unarmed. Mxp 3: Ischium with well-developed spine on flexor distal margin; crista dentata with 20 – 21 denticles. Merus shorter than ischium; flexor margin with 2 spines, proximal stronger than distal; extensor margin with 1 well-developed distal spine. Carpus unarmed. P 1: 3.4 (females) to 4.0 times (males) carapace length. Ischium with some ventral spines. Merus 1.3 times length of carapace, 1.5 – 2.0 times as long as carpus, with spines arranged roughly in rows, dorsomesial spines stronger; distal spines prominent. Carpus 0.7 – 0.8 length of palm, 2.7 – 3.5 times as long as broad; dorsal surface with small spines arranged roughly in longitudinal rows; mesial row of well-developed spines. Palm 2.8 – 3.3 times longer than broad, lateral and mesial margins with small spines arranged roughly in dorsolateral and dorsomesial rows, some small spines scattered on dorsal side. Fingers 0.7 – 0.8 length of palm, each finger distally with two rows of teeth, spooned; mesial margin of movable finger and lateral margin of fixed finger unarmed. P 2 – 4: Moderately long and slender. P 2 2.0 times carapace length. Meri successively shorter posteriorly (P 3 merus 0.9 length of P 2 merus, P 4 merus 0.8 length of P 3 merus); P 2 merus 0.7 – 0.9 carapace length, 4.0 times as long as broad, 1.2 – 1.4 times longer than P 2 propodus; P 3 merus 4.2 times longer than broad, 1.1 – 1.2 times longer than P 3 propodus; P 4 merus 3.1 times as long as broad, 1.1 length of P 4 propodus. Extensor margins of meri with row of 8 – 10 proximally diminishing spines on P 2 – 3, 3 – 5 spines on P 4; flexor margins distally ending in strong spine followed proximally by 1 – 4 small spines and several tubercles or eminences; lateral sides with some minute spines on P 2 – 4. Carpi with 5 – 8 spines on extensor margin, distalmost longer than distal second; lateral surface with small spines and acute granules sub-paralleling extensor margin on P 2 – 4; flexor distal margin with small spine. Propodi 5.2 – 6.0 times as long as broad; extensor margin with 5 – 8 small proximal spines on P 2 – 4, terminal spines paired; flexor margin with 5 – 7 slender movable spines on P 2 – 4. Dactyli distally ending in well-curved strong spine, length 0.5 – 0.6 that of propodi; flexor margin with 5 – 6 proximally diminishing teeth, terminal tooth prominent. Epipods on P 1. Setae: Three types of setae were observed, (1) short simple iridescent setae, sometimes with minute setules along both sides of first half of shaft, densely located on ridges of carapace and pleon, and scales of P 1 – 4; (2) long thick iridescent plumose setae, with setules along both sides of shaft and reaching nearly to end of shaft, sparsely located on carapace, pleon, and P 1 – 4; (3) long thick iridescent simple setae, sparsely located on carapace, pleon and P 1 – 4. Genetic data. COI and 16 S.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C64FFED23B6F93DFCDED3C9.taxon	discussion	Remarks. Galathea gibbosa sp. nov. belongs to the group of species characterized by the presence of more than 4 epigastric spines and often with spinules on hepatic and branchial regions, the lateral margin of the carapace has one small but distinct spine between the anterolateral spine and the anteriormost branchial marginal spine, and the antennular article 1 has 2 well-developed terminal spines, being the distomesial spine very small or obsolescent, e. g., G. pascualae Macpherson & Robainas-Barcia, 2015, G. crinita Macpherson & Robainas-Barcia, 2015, G. pubescens Stimpson, 1858, G. scolopia Macpherson & Robainas-Barcia, 2015, G. tagaloa Macpherson & Robainas-Barcia, 2015 from the central and western Pacific, and G. villosa Macpherson & Robainas-Barcia, 2015 from Madagascar and Mozambique. The morphologically and genetically closest species is G. villosa and both species can be distinguished by the anterior protogastric ridge, with a median prominent convexity with 2 small median spines having some long and thick setae in the new species, whereas this ridge is not prominent, unarmed and without long thick setae in G. villosa. This morphological difference is also distinguishing the new species from G. pascualae and G. tagaloa. The new species is easily distinguished from G. pubescens and G. crinita by the absence of numerous anterior branchial and postcervical spines, which are present in G. pubescens and G. crinita. Galathea gibbosa is also different from G. scolopia by the shape of the rostrum, extremely narrow in G. scolopia and relatively broad in the new species. The genetic divergences between G. gibbosa and other related species are, for COI: 3.91 %, 7.42 %, 7.73 %, and 12.56 % for G. villosa, G. crinita, G. pubescens, and G. scolopia, respectively. For 16 S, we only have reliable data respect to G. crinita (1.02 %) and G. scolopia (1.73 %).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C64FFED23B6F93DFCDED3C9.taxon	distribution	Distribution. New Caledonia and Vanuatu, 252 – 452 m.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7BFFEF23B6FAA9FD79D7D9.taxon	description	(Fig. 9)	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7BFFEF23B6FAA9FD79D7D9.taxon	materials_examined	Material examined. Holotype: Papua New Guinea. BIOPAPUA, Stn CP 3659, 04 ° 14 ’ S, 152 ° 17 ’ E, 22 September 2010, 508 m: F 4.0 mm (MNHN-IU- 2011 - 1940).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7BFFEF23B6FAA9FD79D7D9.taxon	etymology	Etymology. From the Latin, penna, in reference to the presence of some tuft of plumose setae on the carapace.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7BFFEF23B6FAA9FD79D7D9.taxon	description	Description. Carapace: As long as broad; cervical groove distinct, laterally bifurcated. Gastric region with some transverse ridges: 2 epigastric ridges, anterior ridge medially interrupted, with 2 median spines, posterior ridge short, scale-like; 2 protogastric ridges, anterior ridge medially interrupted, without parahepatic spine, posterior ridge long, with some median long and thick plumose setae, short median ridge behind this ridge; 2 mesogastric ridges, anterior ridge medially uninterrupted, not extending laterally to anteriormost of branchial marginal spines, posterior ridge short; 2 metagastric ridges, anterior one uninterrupted, not extending laterally to anterior branchial ridges and with some median long thick plumose setae, posterior ridge scale-like. Hepatic region unarmed. Anterior branchial region with distinct ridges. Mid-transverse ridge uninterrupted, preceded by shallow cervical groove. Posterior branchial region with 6 transverse ridges, 2 ridges uninterrupted. Lateral margins convex medially, with 6 spines: 2 spines in front of and 4 spines behind anterior cervical groove; first anterolateral, well-developed, at level of lateral limit of orbit; second, small, at midlength between anterolateral spine and anteriormost spine of branchial margin, with small accompanying spine ventral to between first and second; 2 spines on anterior branchial region, last small, and 2 spines on posterior branchial margin, last small. Lateral limit of orbit unarmed; infraorbital margin with strong spine. Rostrum 1.8 times as long as broad, length 0.6 postorbital carapace length and breadth 0.3 that of carapace; distance between distalmost lateral incisions 0.25 distance between proximalmost lateral incisions; dorsal surface nearly horizontal in lateral view, with numerous setae; lateral margin with 4 deeply incised sharp teeth. Pterygostomian flap rugose, unarmed, ridges with short setae, anterior margin bluntly angular. Thoracic sternum: As long as broad, lateral extremities gently divergent posteriorly. Sternite III with median shallow notch. Sternite IV with anterior part as wide as sternite III, with some minute striae. Sternites IV – VI with a few striae on lateral sides. Sternite III ~ 2.1 × as wide as long; sternite IV nearly 2.0 × as wide as long, and 3.5 × as wide as sternite III. Pleon: Tergites II and III each with 2 transverse uninterrupted ridges on tergite, anterior ridge more distinctly elevated than posterior ridge; tergite of somite IV with posterior ridge medially interrupted; tergite of somites V and VI with 2 ridges. Eye: Ocular peduncles 1.5 times longer than broad, maximum corneal diameter 0.7 rostrum width. Antennule: Article 1 with 2 spines; well-developed distodorsal and distolateral spines, distodorsal larger; distomesial spine distinct, clearly not reaching midlength of distolateral spine. Ultimate article with tuft of long setae on distodorsal margin. Antenna: Article 1 with ventral distomesial spine reaching distal margin of article 2. Article 2 with 2 well-developed distal spines, lateral spine longer than mesial and reaching midlength of article 3. Articles 3 and 4 unarmed. Mxp 3: Ischium with well-developed spine on flexor distal margin; extensor margin ending in acute angle; crista dentata with 21 denticles. Merus shorter than ischium; flexor margin with 3 spines, decreasing in size distally; extensor margin with distal spine. Carpus unarmed. P 1: 3.5 times carapace length. Merus 1.2 times length of carapace, 1.6 times as long as carpus, with spines arranged roughly in rows, dorsomesial spines stronger; distal spines prominent. Carpus 0.9 length of palm, twice longer than broad; dorsal surface with some small spines; mesial margin with well-developed spines. Palm 2.1 times longer than broad, lateral and mesial margins subparallel; some spines arranged roughly in dorsolateral and dorsomesial rows, some small spines scattered on dorsal side. Fingers 0.8 times palm length, each finger distally with two rows of teeth, spooned; fixed finger with some small proximal spines. P 2 – 4: Moderately slender. P 2 2.1 times carapace length. Meri successively shorter posteriorly (P 3 merus 0.8 length of P 2 merus, P 4 merus 0.8 length of P 3 merus); P 2 merus 0.8 carapace length, 4.7 times as long as broad, 1.2 times longer than P 2 propodus. Extensor margin with row of 6 – 8 proximally diminishing spines on P 2 – 3, 1 distal spine on P 4; ventral margins distally ending in strong spine, lateral sides unarmed. Carpi with 3 – 5 spines on extensor margin on P 2 – 4, distalmost smaller than distal second, sometimes absent; lateral surface with several acute granules sub-paralleling extensor margin; flexor distal margin acute. Propodi 4.3 – 5.0 times as long as broad; extensor margin with 0 – 2 small proximal spines; flexor margin with 5 – 7 slender movable spines, terminal spines paired. Dactyli distally ending in well-curved strong spine, length 0.6 – 0.8 that of propodi; flexor margin with 6 – 7 proximally diminishing teeth, terminal tooth moderately prominent. Epipods absent on pereiopods. Setae: Two types of setae were observed, (1) short plumose setae, with setules along both sides of shaft, densely located on ridges of carapace and pleon, and scales of P 1 – 4; (2) long thick plumose setae, with setules along both sides of shaft, sparsely located on carapace, pleon, and P 1 – 4. Genetic data. COI.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7BFFEF23B6FAA9FD79D7D9.taxon	discussion	Remarks. The new species is referred to the group of species characterized by the presence of one small but distinct spine between the anterolateral spine and the anteriormost branchial marginal spine, two well-developed distal spines on the antennular basal article, the possession of two epigastric spines and the absence of hepatic and parahepatic spines, and epipods. It resembles Galathea yamashitai Miyake & Baba, 1967 from East China Sea, although both species can be distinguished by the following characters: — The antennal article 3 has a distomesial spine in G. yamashitai, whereas this spine is absent in the new species. — The second lateral spine of the carapace is present in the new species, instead of absent in G. yamashitai. — Pleonal tergites 2 – 3 with 2 ridges in the new species, whereas there are 4 – 5 ridges in G. yamashitai. Unfortunately, genetic data for G. yamashitai are not available. The closest species are G. lopisma Macpherson, Rodríguez-Flores & Machordom, 2023, from the Mozambique Channel, and G. setigera Macpherson & Robainas-Barcia, 2015, from the Kei Islands. The divergences for COI are 15.06 % with G. setigera and 17.63 % with G. lopisma; there are no data for 16 S.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7BFFEF23B6FAA9FD79D7D9.taxon	distribution	Distribution. Papua New Guinea, 508 m.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7EFFEA23B6FF11FD4CD51D.taxon	description	(Fig. 10)	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7EFFEA23B6FF11FD4CD51D.taxon	materials_examined	Material examined. Holotype: Papua-New Guinea, PAPUA NIUGINI, Stn PB 19, 05 ° 05.1 ’ S, 145 ° 48.6 ’ E, 30 December 2012, 10 m: ov. F 2.4 mm (MNHN-IU- 2014 - 13935). Paratype: same data than holotype: 1 M 2.2 mm (MNHN-IU- 2014 - 20839).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7EFFEA23B6FF11FD4CD51D.taxon	etymology	Etymology. From the Greek, rhanis, drop, spot, in reference to the small size of the species.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7EFFEA23B6FF11FD4CD51D.taxon	description	Description. Carapace: As long as broad; anterior cervical groove indistinct. Five ridges on gastric region: 1 epigastric ridge, medially convex and interrupted, with 2 epigastric spines; 1 protogastric ridge uninterrupted, without parahepatic spines; 1 mesogastric ridge extending laterally to anteriormost of branchial marginal spines, uninterrupted or with minute interruption medially; 2 metagastric ridges, anterior ridge medially interrupted and extending laterally to branchial margin, posterior ridge short. Mid-transverse ridge uninterrupted, preceded by shallow cervical groove, followed by 5 ridges, 1 – 2 uninterrupted ridges. Lateral margins medially convex, with 6 spines: 1 spines in front of and 5 spines behind indistinct anterior cervical groove; first anterolateral, well-developed, at same level of lateral limit of orbit; 2 spines on anterior branchial region, last small, and 3 spines on posterior branchial margin, last small. External orbital limit ending in small spine; infra-orbital margin with strong spine. Rostrum broad triangular, 1.1 – 1.2 times as long as broad, length 0.5 postorbital carapace length and breadth 0.4 that of carapace, nearly horizontal in lateral view; distance between distalmost lateral incisions 0.3 distance between proximalmost lateral incisions; dorsal surface with some short setae; lateral margin with 4 deeply incised sharp teeth. Pterygostomian flap rugose, with facial small spine on anterior part, anterior margin ending in small spine. Thoracic sternum: As long as broad, lateral extremities gently divergent posteriorly. Sternite III with median shallow notch, anterior margin at each side of notch slightly convex. Sternite IV with anterior part as wide as sternite III, with some minute striae. Sternites IV – VI with a few striae on lateral sides. Sternite III ~ 1.8 × as wide as long; sternite IV nearly 3 × as wide as long, and 2.5 × as wide as sternite III. Pleon: Tergite II with 2 transverse ridges on tergite, anterior ridge more distinctly elevated than posterior ridge; tergites III and IV smooth or with some scales, with anterior ridge only; tergite of somites V and VI smooth. Males with G 1 and G 2. Eye: Ocular peduncles 1.2 times longer than broad, maximum corneal diameter 0.6 rostrum width. Antennule. Article 1 with 3 well-developed distal spines, distodorsal larger; distomesial spine reaching midlength of distolateral spine. Ultimate article with a few short setae not in tuft on distodorsal margin. Antenna. Article 1 with distomesial process reaching distal margin of article 2. Article 2 with distomesial spine barely shorter than distolateral spine, exceeding midlength of article 3. Articles 3 and 4 unarmed. Mxp 3. Ischium with well-developed distal spine on flexor margin; extensor margin unarmed; crista dentata with 26 – 27 denticles. Merus subequal in length to ischium, with 2 – 3 small spines on flexor margin, proximal one located at midlength, distal one at terminal end, median spine slightly smaller than others or obsolescent; extensor margin with distal spine. P 1. Lost in both specimens. P 2 – 4. Relatively slender, somewhat compressed, moderately setose. P 2 1.5 times carapace length. Meri successively shorter posteriorly (P 3 merus 0.9 length of P 2 merus, P 4 merus 0.8 length of P 3 merus); P 2 merus 0.6 carapace length, 3 times as long as broad, 1.3 times longer than P 2 propodus; P 3 merus 3.0 times as long as broad, 1.3 times length of P 3 propodus; P 4 merus 2.5 times as long as broad, 1.1 length of P 4 propodus. Extensor margins with row of 6 proximally diminishing spines on P 2 – 3, 4 spines on P 4; lateral surface unarmed on P 2 – 3, 0 – 1 spines on P 4; flexolateral margin with strong terminal spine. Carpi with 3 – 4 spines on extensor margin; lateral surface with row of 2 – 4 small spines or acute granules paralleling extensor row; flexor distal margins with spine. Propodi 4 times as long as broad; extensor margin with 0 – 2 proximal spines on P 2 – 4; flexor margin with 3 – 4 slender movable spines. Dactyli subequal in length, 0.7 length of propodi, ending in incurved, strong, sharp spine; flexor margin with prominent triangular terminal tooth preceded by obsolescent 4 – 5 teeth. Epipods present on P 1 – 3. Setae: Three types of setae were observed, (1) short plumose setae, with setules along both sides of shaft, densely located on ridges of carapace and pleon; (2) long thick iridescent simple setae, sparsely located on pleon and P 2 – 4; (3) short thick iridescent simple setae, sparsely located on pleon and P 2 – 4. Genetic data. COI.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7EFFEA23B6FF11FD4CD51D.taxon	discussion	Remarks. Galathea rhanis sp. nov. belongs to the group of species having an uninterrupted mesogastric ridge between the anteriormost branchial marginal spines, one pair of epigastric spines, the pterygostomian flap with 1 facial spine on the anterior part, epipods on P 1 – 3 and the antennal article 3 with distal spines. The new species is closely related to G. paulayi Macpherson & Robainas-Barcia, 2015 from the French Polynesia and Kiribati and G. stilla sp. nov. (see below). The new species is easily distinguished from both species by the following characters: — The parahepatic spines are absent in G. rhanis, whereas these spines are present in the other two species. — The carapace lateral margin without a spine between the anterolateral spine and the anteriormost spine of the branchial margin in G. rhanis, whereas this spine is always present in G. paulayi and G. stilla. — The genetic divergences with G. paulayi and G. stilla are 17.63 % and 18.57 % for COI, respectively.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7EFFEA23B6FF11FD4CD51D.taxon	distribution	Distribution. Papua New Guinea, at 10 m.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7CFFE523B6FD05FD07D4CD.taxon	description	(Fig. 11, 12)	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7CFFE523B6FD05FD07D4CD.taxon	materials_examined	Material examined. Holotype: Kiribati, Phoenix Islands, 4 ° 31 ’ S, 172 ° 13 ’ W, 10.5 m, 18 September 2015: M 2.8 mm (UF 51343). Paratype: same data than holotype: ov. F 3.0 mm (UF 51346).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7CFFE523B6FD05FD07D4CD.taxon	etymology	Etymology. From the Latin, stilla, drop, in reference to the small size of the species.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7CFFE523B6FD05FD07D4CD.taxon	description	Description. Carapace: As long as broad; anterior cervical groove indistinct. Seven ridges on gastric region: 1 epigastric ridge, with 2 epigastric spines, medially convex and medially uninterrupted; 2 protogastric ridges, anterior one uninterrupted, with 1 parahepatic spine at each side, posterior ridge uninterrupted or scale-like, with 2 median long thick simple setae; 2 mesogastric ridges, anterior ridge uninterruptedly extending laterally to anteriormost of branchial marginal spines, posterior one short; 2 metagastric ridges, anterior ridge uninterrupted and fused with anterior branchial ridge, posterior ridge short. Mid-transverse ridge uninterrupted, preceded by shallow cervical groove, followed by 5 ridges, 2 – 3 of them uninterrupted. Lateral margins medially convex, with 7 spines: 2 spines in front of and 5 spines behind indistinct anterior cervical groove; first anterolateral, well-developed, at same level of lateral limit of orbit, accompanying another spine ventral to between first and second, second spine small or minute; 2 spines on anterior branchial region, last small, and 3 spines on posterior branchial margin, last small. External orbital limit ending in small spine; infra-orbital margin with strong spine. Rostrum broad triangular, 1.2 times as long as broad, length 0.5 postorbital carapace length and breadth 0.4 that of carapace, nearly horizontal in lateral view; distance between distalmost lateral incisions 0.2 distance between proximalmost lateral incisions; dorsal surface with some short setae; lateral margin with 4 deeply incised sharp teeth. Pterygostomian flap rugose, with facial spine on anterior part, anterior margin ending in small spine. Thoracic sternum: As long as broad, lateral extremities gently divergent posteriorly. Sternite III with median shallow notch, anterior margin at each side of notch slightly convex. Sternite IV with anterior part as wide as sternite III, with some minute striae. Sternites IV – VI with a few striae on lateral sides. Sternite III ~ 2.2 × as wide as long; sternite IV nearly 1.6 × as wide as long, and 1.4 × as wide as sternite III. Pleon: Tergites II and III each with 2 transverse ridges on tergite, anterior ridge more distinctly elevated than posterior ridge; tergite of somites IV and V with posterior ridge medially interrupted; somite VI with 2 medially interrupted ridges, posteromedian margin straight. Males with G 1 and G 2. Eye: Ocular peduncles 1.2 times longer than broad, maximum corneal diameter 0.7 rostrum width. Antennule. Article 1 with 3 well-developed distal spines, distodorsal larger. Ultimate article with a few short setae not in tuft on distodorsal margin. Antenna. Article 1 hardly visible from dorsal view, with depressed ventral distomesial process exceeding distal margin of article 3. Article 2 with distomesial spine smaller than distolateral, exceeding midlength of article 3. Articles 3 and 4 unarmed. Mxp 3. Ischium with well-developed distal spine on flexor margin; extensor margin unarmed; crista dentata with 26 – 28 denticles. Merus subequal in length to ischium, with 3 spines on flexor margin, proximal one located at midlength, distal one at terminal end, median spine smaller than others; extensor margin with distal spine. Carpus with distal spine on flexor margin. P 1. 2.2 times carapace length. Merus 0.8 times carapace length, 1.5 times as long as carpus, with rows of spines, mesial and distal spines strong. Carpus 0.9 length of palm, 1.5, times longer than broad, lateral and mesial margins subparallel, dorsal surface with some spines in longitudinal rows; mesial surface with row of well – developed spines; and row of small spines along lateral margin. Palm 1.8 times longer than broad, lateral and mesial margins subparallel; small spines roughly in rows on dorsal; lateral row continued on to lateral margin of fixed finger; mesial row continuing on the mesial margin of movable finger. Fingers 0.7 as long as palm, each finger distally with two rows of teeth, spooned. P 2 – 4. Relatively slender, somewhat compressed. P 2 1.7 times carapace length. Meri successively shorter posteriorly (P 3 merus 0.9 length of P 2 merus, P 4 merus 0.8 length of P 3 merus); P 2 merus 0.7 carapace length, 3.0 times as long as broad, 1.5 times longer than P 2 propodus; P 3 merus 2.5 times as long as broad, 1.3 times length of P 3 propodus; P 4 merus 2.5 times as long as broad, 1.3 length of P 4 propodus. Extensor margins with row of 7 – 9 proximally diminishing spines on P 2 – 3, 6 spines on P 4; lateral surface unarmed on P 2 – 3, 1 – 2 spines on P 4; flexolateral margins with strong terminal spine and 1 – 2 additional spines on terminal half; flexomesial margin with terminal spine on P 2 – 3. Carpi with 4 – 5 spines on extensor margin; lateral surface with row of 2 - 4 small spines or acute granules paralleling extensor row; flexor distal margins with spine. P 2 – 4 propodi each 3.5 – 4.5 times as long as broad; extensor margin with 4 proximal spines on P 2 – 4; flexor margin with 4 – 5 slender movable spines, terminal spines paired; 2 – 3 proximal spines on lateral side. Dactyli subequal in length, 0.7 length of propodi, ending in incurved, strong, sharp spine; flexor margin with prominent triangular terminal tooth preceded by obsolescent 5 teeth. Epipods present on P 1 - 3. Setae: Three types of setae were observed, (1) short plumose setae, with setules along both sides of shaft, densely located on ridges of carapace and pleon and sparsely on P 1 – 4; (2) long thick iridescent simple setae, sparsely located on carapace, pleon and P 1 – 4; (3) long thick plumose setae, with long setules along one side of shaft, sparsely located on P 1 – 4. Genetic data. COI.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7CFFE523B6FD05FD07D4CD.taxon	discussion	Remarks. Galathea stilla belongs to the group of species having an uninterrupted ridge between the anteriormost branchial marginal spines, the carapace lateral margin having one small spine between the anterolateral spine and the anteriormost branchial marginal spine, one pair of epigastric spines, the pterygostomian flap with 1 facial spine, and epipods on P 1 – 3. The new species is closely related to G. paulayi Macpherson & Robainas-Barcia, 2015 from the French Polynesia and Line Islands. The new species is easily distinguished from G. paulayi by the shape of the P 2 - 4 meri, slenderer in G. paulayi than in the new species. For instance, the P 2 merus is 3.5 – 3.9 times as long as broad in G. paulayi, whereas it is nearly 3.0 times in the new species. The genetic divergence between these two species is 12.07 % (COI). The new species is also genetically close to G. amamiensis Miyake & Baba, 1966, and G. boucheti Macpherson & Robainas-Barcia, 2015, both from the Indian and Western Pacific areas. However, the new species has epipods on P 1 – 3, whereas G. amamiensis and G. boucheti only have epipods on P 1. Genetically, they are also very different from the new species (11.96 % and 11.55 %, respectively for COI).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C7CFFE523B6FD05FD07D4CD.taxon	distribution	Distribution. Kiribati, Phoenix Islands, 10.5 m.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C73FFE723B6FBFDFCF8D3E5.taxon	description	(Fig. 13)	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C73FFE723B6FBFDFCF8D3E5.taxon	materials_examined	Type material. Holotype: Philippines, MUSORSTOM 3, Stn DR 117, 12 ° 31 ’ N, 120 ° 39 ’ E, 3 June 1985, 92 – 97 m: M 1.8 mm (MNHN-IU- 2016 - 502). Paratypes: same data than holotype: 1 M 1.7 mm, 2 F 1.6 – 1.7 mm (MNHN-IU- 2014 - 13802).	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C73FFE723B6FBFDFCF8D3E5.taxon	etymology	Etymology. From the Greek, psekas, small drop, in reference to the small size of the species. De s cription. Carapace: Length 0.9 × width; transverse ridges granulated. Gastric region with numerous short or scale-like ridges; epigastric region with 2 - 3 pairs of minute submedian spines or acute granules; anterior protogastric ridge medially interrupted, not extending laterally to carapace margin; anterior mesogastric ridge scale-like; metagastric ridge laterally interrupted by cervical groove, laterally scale-like. Mid-transverse ridge laterally interrupted, preceded by shallow cervical groove, followed by 2 laterally interrupted ridges, and some short or scale-like ridges on branchial and intestinal areas. Lateral margins medially convex, with 7 – 8 spines: 2 spines (anterolateral, hepatic) in front and 5 – 6 spines (3 anterior branchial spines, 2 – 3 posterior branchial spines) behind distinct anterior cervical groove. Anterolateral spine well-developed, not reaching anteriorly to level of lateral orbital spine. Hepatic spine small, set slightly dorsomesial from lateral margin. First anterior branchial spine slightly longer than second and third spines. Posterior branchial spines progressively smaller posteriorly, first spine subequal to anterior branchial spines. Rostrum widely triangular, downwards directed, 0.5 × pcl; length-width ratio 0.9; lateral margins unarmed, slightly convex, with distinct pair of sub-apical spines; dorsally concave. Pterygostomian flap with sharp anterior spine, upper margin with minute spines or granules; entire surface with series of scale-like ridges. Thoracic sternum: Sternal plastron slightly wider than long (length-width ratio 0.9), lateral margins divergent posteriorly. Sternite 3 about twice as wide as long; anterior margin slightly convex, with median notch. Pleon: Tergites 2 and 3 with anterior and posterior transverse ridges; tergite 4 with anterior ridge only; tergites 5 and 6 smooth, without elevated ridges. Males with G 1 absent. Eye: Eye stalk a little wider than long, peduncle mesially with minute spines and plumose setae; cornea narrower than peduncle. Antennule: Article 1 with 3 distal spines, distomesial spine distinct; 2 distolateral spines, and 2 lateral spines, proximal as long as distomesial spine. Antenna: Article 1 with prominent mesial process, distally reaching end of peduncle. Article 2 with distinct mesial and lateral spines. Article 3 usually unarmed or with minute distomesial spine. Article 4 unarmed. Mxp 3: Ischium with pronounced distal spines on flexor and extensor margins; crista dentata with regular row of 18 – 19 spines along entire margin. Merus slightly shorter than ischium at midlength, with strong distal spines on extensor margins, smaller than spine at midlength on flexor margin, otherwise unarmed. Cheliped (P 1): missing. Walking legs (only P 3; P 2 and P 4 missing in all specimens): Merus length 0.5 × pcl; extensor margin with row of 5 spines, proximally diminishing, with prominent distal spine; flexor margin with distal spine. Carpus with 2 spines on extensor margin, distal spine prominent; flexor margin unarmed. Propodus slightly shorter than menus, length 6 × height; extensor margin unarmed; flexor margin with 3 slender movable spines in addition to distal pair. Dactyli 0.8 × propodus length, ending in incurved, strong, sharp spine; flexor margin with 6 movable spines, otherwise unarmed. Setae: Unfortunately, most setae are lost in all specimens. Two types of setae were observed, (1) short plumose setae, with setules along both sides of shaft, densely located on ridges of carapace and pleon; (2) long thick plumose setae, with long setules along one side of shaft, sparsely located on carapace and pleon. Genetic data. No data	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C73FFE723B6FBFDFCF8D3E5.taxon	discussion	Remarks. The genus Nanogalathea was described by Tirmizi & Javed (1980) to include a small species of squat lobster (N. raymondi) collected during the International Indian Ocean Expedition in the Andaman Sea, at 77 m. Recently, a second species (N. longispina) was described by Tiwari et al. (2024) from Kerala, Arabian Sea. The genus is characterized by its small size, a leaf-like rostrum, without supraocular spines, and carapace with weak ridges. The new species is clearly different from N. raymondi by the following aspects: — The lateral margins of the rostrum are straight in N. raymondi, whereas these margins are convex in the new species. Furthermore, the distal spines of the rostrum seem more distinct in the new species than in N. raymondi and N. longispina. — The epigastric region is armed with 2 – 3 pairs of spines in the new species, whereas this region is unarmed in N. raymondi and N. longispina. — The hepatic margin of the carapace is unarmed in N. longispina, whereas there is a marginal spine in the new species.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
A25F535A3C73FFE723B6FBFDFCF8D3E5.taxon	distribution	Distribution. The Philippines, between 92 and 97 m.	en	Macpherson, Enrique, Rodríguez-Flores, Paula C., Machordom, Annie (2025): New species of Galathea Fabricius, 1793 and Nanogalathea Tirmizi & Javed, 1980 (Crustacea: Decapoda, Galatheidae) from the Western Pacific. Zootaxa 5570 (3): 447-483, DOI: 10.11646/zootaxa.5570.3.2, URL: https://doi.org/10.11646/zootaxa.5570.3.2
