taxonID	type	description	language	source
9BCF807BD2855EBF815AD189C85910E3.taxon	description	Figs 1, 2, 3, 4, 14 A, 15 A, 16 Vernacular name: 淡黄博狼蛛	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
9BCF807BD2855EBF815AD189C85910E3.taxon	diagnosis	Diagnosis. Bogdocosa flavida is similar to B. multivaga and B. yunshani sp. nov. in having a pair of epigynal lobes (Figs 1 B, 2 D, 4 C; 8 D, 10 C; 11 D, 13 C), but can be distinguished by ventral arm of median apophysis rod-shaped ventrally in B. flavida (Figs 2 B, 3 B, D, G, 14 A) (vs. digitiform in B. multivaga, Figs 8 B, 9 B, D, G, 14 C; sectorial in B. yunshani sp. nov., Figs 11 B, 12 B, D, G, 14 D). Females can be distinguished by two epigynal lobes closely approximated with individual lobes 2 times as wide as long in B. flavida (Figs 1 B, 2 D, 4 C) (vs. interlobar space narrower than lobe width with individual lobes 1.5 times as wide as long in B. multivaga, Figs 8 D, 10 C; interlobar space equal to lobe width and individual lobes as wide as long in B. yunshani sp. nov., Figs 11 D, 13 C).	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
9BCF807BD2855EBF815AD189C85910E3.taxon	description	Description. Male (Fig. 4 A) total length 5.65. Carapace 2.98 long, 2.03 wide; abdomen 2.50 long, 1.75 wide. Eye sizes and interdistances: AME 0.14, ALE 0.16, PME 0.30, PLE 0.24; AME – AME 0.07, AME – ALE 0.04, PME – PME 0.19, PME – PLE 0.24. Clypeus 0.10 high. Leg measurements: I 7.91 (2.14, 2.76, 1.73, 1.28); II 7.06 (2.01, 2.23, 1.69, 1.13); III 7.37 (1.95, 2.39, 1.98, 1.05); IV 10.91 (2.70, 3.40, 3.36, 1.45). Palp (Figs 2 A – C, 3 A – G, 14 A). Terminal apophysis broad, with small tip. Prolateral arm of median apophysis sail-shaped ventrally. Retrolateral arm of median apophysis membranous. Ventral arm of median apophysis rod-shaped and narrower than retrolateral arm of median apophysis in ventral view. Lobe of embolic base extending ventrally. Embolus roundly bent. Conductor trifurcates. Female (Fig. 4 B) total length 7.27. Carapace 3.56 long, 2.54 wide; abdomen 3.80 long, 2.48 wide. Eye sizes and interdistances: AME 0.13, ALE 0.12, PME 0.37, PLE 0.29; AME – AME 0.11, AME – ALE 0.07, PME – PME 0.24, PME – PLE 0.31. Clypeus height 0.19. Leg measurements: I 9.47 (2.75, 3.23, 2.01, 1.48); II 8.82 (2.54, 2.94, 1.94, 1.40); III 8.86 (2.56, 2.61, 2.47, 1.22); IV 13.71 (3.62, 4.04, 4.28, 1.77). Epigyne (Figs 1 B, C, 2 D, E, 4 C, D). Epigynal lobes wider than long and closely approximated. Median plate square. Copulatory openings situated bilaterally on the median plate, with their orifices posteriorly oriented. Spermathecal heads oval.	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
9BCF807BD2855EBF815AD189C85910E3.taxon	distribution	Distribution. The species is known from Gansu, Qinghai and Xinjiang in China (sub. Pardosa xinjiangensis (Li and Lin 2016) most likely occurs in Afghanistan and Jammu and Kashmir. Syntype series includes specimens from “ Yarkand to Bursi ” that may refer to India) (Fig. 16).	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
1E6CFC4A6A885B0D950C876F6A4CCAD8.taxon	description	Figs 5, 6, 7, 14 B, 15 B, 16 Vernacular name: 克氏博狼蛛	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
1E6CFC4A6A885B0D950C876F6A4CCAD8.taxon	diagnosis	Diagnosis. Bogdocosa kronebergi is distinguished from all other congeners by the rectangular ventral arm of median apophysis ventrally and oval median plate.	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
1E6CFC4A6A885B0D950C876F6A4CCAD8.taxon	description	Description. See Ponomarev et al. (2008). Habitus as shown in Figs 7 A, B, male palp as in Figs 5 A – C, 6, 14 B, epigyne as in Figs 5 D, E, 7 C, D.	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
1E6CFC4A6A885B0D950C876F6A4CCAD8.taxon	distribution	Distribution. This species is distributed from Transcaucasia east to Xinjiang, north to 49 ^ N in northwesternmost Kazakhstan and south to northeastern Iran. It is reported from Armenia (first record), Azerbaijan, Iran, Kazakhstan, Turkmenistan, Uzbekistan, Tajikistan and China (Fig. 16).	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
799942750AE652E89C15FE0159AB3733.taxon	description	Figs 8, 9, 10, 14 C, 15 C, 16 Vernacular name: 多瓣博狼蛛	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
799942750AE652E89C15FE0159AB3733.taxon	diagnosis	Diagnosis. Bogdocosa multivaga is similar to B. flavida and B. yunshani sp. nov. in having a pair of epigynal lobes (Figs 8 D, 10 C; 1 B, 2 D, 4 C; 11 D, 13 C), but can be distinguished by ventral arm of median apophysis digitiform ventrally in B. multivaga (Figs 8 B, 9 B, D, G, 14 C) (vs. rod-shaped in B. flavida, Figs 2 B, 3 B, D, G, 14 A; sectorial in B. yunshani sp. nov., Figs 11 B, 12 B, D, G, 14 D). Females can be distinguished by interlobar space narrower than lobe width with individual lobes 1.5 times as wide as long in B. multivaga (Figs 8 D, 10 C) (vs. two epigynal lobes closely approximated with individual lobes 2 times as wide as long in B. flavida, Figs 1 B, 2 D, 4 C; interlobar space equal to lobe width and individual lobes as wide as long in B. yunshani sp. nov., Figs 11 D, 13 C).	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
799942750AE652E89C15FE0159AB3733.taxon	description	Description. Male (Fig. 10 A) total length 6.13. Prosoma 3.19 long, 2.28 wide; opisthosoma 2.99 long, 1.99 wide. Eye sizes and interdistances: AME 0.12, ALE 0.10, PME 0.28, PLE 0.24; AME – AME 0.08, AME – ALE 0.03, PME – PME 0.17, PME – PLE 0.21. Clypeus height 0.18. Leg measurements: I 7.97 (2.16, 2.67, 1.89, 1.25); II 7.30 (2.02, 2.39, 1.75, 1.14); III 7.19 (1.95, 2.28, 1.88, 1.08); IV 11.25 (2.88, 3.44, 3.46, 1.47). Palp (Figs 8 A – C, 9 A – G, 14 C). Terminal apophysis hook-shaped, with short and thick end. Prolateral arm of median apophysis sclerotized. Ventral arm of median apophysis digitiform and narrower than retrolateral arm of median apophysis ventrally. Embolus bent into a human-ear-shape in ventral view. Lobe of embolic base with arc-shaped end. Conductor with two bifurcations. Female (Fig. 10 B) total length 7.80. Prosoma 3.33 long, 2.45 wide; opisthosoma 4.66 long, 2.77 wide. Eye sizes and interdistances: AME 0.12, ALE 0.10, PME 0.29, PLE 0.24; AME – AME 0.09, AME – ALE 0.06, PME – PME 0.20, PME – PLE 0.28. Clypeus height 0.15. Leg measurements: I 8.07 (2.33, 2.72, 1.85, 1.17); II 7.14 (2.11, 2.32, 1.63, 1.08); III 7.76 (2.08, 2.42, 2.09, 1.17); IV 12.26 (3.20, 3.57, 3.92, 1.57). Epigyne (Figs 8 D, E, 10 C, D). Epigynal lobes equal in length and width. Interlobar space narrower than lobe width. Median plate trapezoid, as long as wide. Copulatory openings located on lateral margins of median plate. Spermathecal heads circular.	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
799942750AE652E89C15FE0159AB3733.taxon	distribution	Distribution. China (Beijing, Inner Mongolia, Ningxia, Gansu, Shaanxi, Shanxi, Hebei, Shandong) (Fig. 16).	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
AC3BDD5A86645E0BB013B8B5DDAFA9A3.taxon	type_taxon	Type species. Bogdocosa baskuntchakensis Ponomarev & Belosludtsev, 2008.	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
AC3BDD5A86645E0BB013B8B5DDAFA9A3.taxon	diagnosis	Diagnosis. Males of Bogdocosa can be distinguished from those of all other lycosid genera by the strong erected setae on the male sternum (vs. absent) and median apophysis with 3 branches (vs. 2). Females of Bogdocosa differ from those of most lycosid genera by having the median plate of the epigyne covered with setae together with the lack of septum (vs. setae absent or present on septum only (Arctosa C. L. Koch, 1847, Evippinae) and having strongly sclerotized anterior margins of the epigyne (epigynal lobes), extending the median plate (little in B. kronebergi, and strongly in other species; absent in Aulonia C. L. Koch, 1847 and Halocosa Azarkina & Trilikauskas, 2019).	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
AC3BDD5A86645E0BB013B8B5DDAFA9A3.taxon	description	Description. Medium-sized (4.0 – 8.5). Carapace pear-shaped, yellowish-brown, with wide light median band starting anteriorly from PLE and reaching posterior edge, two longitudinal dark bands, and light marginal stripes. Cervical groove and radial furrows indistinct. Eye region black except space between PLE. Sternum yellowish-brown to grey-brown, males with spine-like black setae. Legs yellowish-brown. Leg formula: 4123 or 4132. Abdomen oval, yellowish-brown to grey-brown, with wide median band, lateral sides covered with black or brown setae. Heart mark distinct, lanceolate. Spinnerets pale yellow. Palp (Figs 2 A – C, 3 A – G, 5 A – C, 6 A – G, 8 A – C, 9 A – G, 11 A – C, 12 A – G, 14 A – D) with tibial bulge. Cymbium droplet-shaped, 1.6 ‒ 1.8 times longer than wide, tip short, about 1 / 4 th of cymbial length and lacking claw-like setae, but with numerous apical strong spines. Bulb oval, ca 1.25 ‒ 1.30 times longer than wide. Subtegulum large, about 0.27 ‒ 0.41 of bulbal length, located postero-prolaterally. Tegulum long with total length of tegulum and conductor matching the bulb in length, with distinct median notch and retrolateral subtriangular lobe. Sperm duct makes deep loop around median notch. Conductor membranous, bifid or trifid. Median apophysis located medially, with 3 arms: prolateral, retrolateral and ventral. Terminal part with palea, terminal apophysis small, with hook-shaped tip, embolus filamentous, roundly bent, with or without lobe directed anteriorly on the embolic base. Epigyne (Figs 1 B, C, 2 D, E, 4 C, D, 5 D, E, 7 C, D, 8 D, E, 10 C, D, 11 D, E, 13 C, D) with pair of epigynal lobes located anteriorly or laterally and almost fusing anteriorly. Median plate oval or pentagonal-rectangular, covered with setae. Copulatory openings crack-like. Spermathecae subdivided into two parts, well distinct in all species, anterior part with distinct stalks and posterior part well distinct in type species only. Fertilization ducts crescent-shaped.	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
AC3BDD5A86645E0BB013B8B5DDAFA9A3.taxon	distribution	Distribution. China (Beijing, Inner Mongolia, Xinjiang, Ningxia, Qinghai, Gansu, Shaanxi, Shanxi, Hebei, Shandong), Russia (North Caucasus), Central Asia: Kazakhstan, Uzbekistan, Tajikistan, Azerbaijan and northern Iran (Fig. 16).	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
564440EE705558D7B87B46A2384D201F.taxon	description	Figs 11, 12, 13, 14 D, 15 D, 16 Vernacular name: 蕴山博狼蛛	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
564440EE705558D7B87B46A2384D201F.taxon	etymology	Etymology. The specific name is a patronym honouring Mr. Yunshan Yang who collected this new species.	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
564440EE705558D7B87B46A2384D201F.taxon	diagnosis	Diagnosis. Bogdocosa yunshani sp. nov. is similar to B. flavida and B. multivaga in having a pair of epigynal lobes (Figs 11 D, 13 C; 1 B, 2 D, 4 C; 8 D, 10 C), but differ by ventral arm of median apophysis approximately sectorial ventrally in B. yunshani sp. nov. (Figs 11 B, 12 B, D, G, 14 D) (vs. rod-shaped in B. flavida, Figs 2 B, 3 B, D, G, 14 A; digitiform in B. multivaga, Figs 8 B, 9 B, D, G, 14 C). Females can be distinguished by interlobar space equal to lobe width and individual lobes as wide as long in B. yunshani sp. nov. (Figs 11 D, 13 C) (vs. two lobes closely approximated with individual lobes 2 times as wide as long in B. flavida, Figs 1 B, 2 D, 4 C; interlobar space equal to lobe width and individual lobes 1.5 times as wide as long in B. multivaga, Figs 8 D, 10 C).	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
564440EE705558D7B87B46A2384D201F.taxon	description	Description. Male (holotype, SWUC - T-LY- 27 - 01) (Fig. 13 A) total length 5.09. Carapace 2.76 long, 1.90 wide; abdomen 2.29 long, 1.53 wide. Eye sizes and interdistances: AME 0.15, ALE 0.09, PME 0.25, PLE 0.19; AME – AME 0.07, AME – ALE 0.02, PME – PME 0.17, PME – PLE 0.20. Clypeus 0.11 high. Legs light yellowish-brown, with black markings. Leg measurements: I 7.15 (1.97, 2.39, 1.69, 1.10); II 6.77 (1.80, 2.24, 1.58, 1.15); III 6.57 (1.85, 1.93, 1.80, 0.99); IV 10.29 (2.67, 2.97, 3.33, 1.32). Palp (Figs 11 A – C, 12 A – G, 14 D). Terminal apophysis hook-shaped, with long and thin end. Prolateral arm of median apophysis membranous and ventral arm of median apophysis sectorial ventrally. Ventral arm of median apophysis wider than retrolateral arm of median apophysis. Embolus bent into a human-ear-shape. Conductor bifurcated. Female (paratype, SWUC - T-LY- 27 - 02) (Fig. 13 B) total length 8.05. Carapace 3.26 long, 2.39 wide; abdomen 4.51 long, 2.96 wide. Eye sizes and interdistances: AME 0.13, ALE 0.12, PME 0.27, PLE 0.23; AME – AME 0.09, AME – ALE 0.03, PME – PME 0.21, PME – PLE 0.28. Clypeus 0.11 high. Leg measurements: I 7.96 (2.27, 2.84, 1.56, 1.29); II 7.53 (2.14, 2.50, 1.65, 1.24); III 7.61 (2.19, 2.26, 2.00, 1.16); IV 12.06 (3.04, 3.67, 3.79, 1.56). Epigyne (Figs 11 D, E, 13 C, D). Epigynal lobes as long as wide. Interlobar space equal to lobe width. Median plate trapezoid, longer than wide. Copulatory openings situated on lateral margins of median plate. Spermathecal heads circular.	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
564440EE705558D7B87B46A2384D201F.taxon	distribution	Distribution. China (Xinjiang) (Fig. 16).	en	Zhang, Xiang-Yun, Marusik, Yuri M., Nadolny, Anton A., Zhang, Zhi-Sheng, Wang, Lu-Yu (2025): A review of the wolf spider genus Bogdocosa (Araneae, Lycosidae). Zoosystematics and Evolution 101 (4): 1445-1458, DOI: 10.3897/zse.101.161058
