identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
00DE5EF153B15290AEDC4BD1012614FD.text	00DE5EF153B15290AEDC4BD1012614FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax amasanga Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax amasanga sp. nov.</p><p>Figs 5 G, H, 7 E – H, 8 I</p><p>Type material.</p><p>Holotype • ♂ (QCAZ -I-278673): “ Ecuador. Orellana, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.40004&amp;materialsCitation.latitude=-0.68315" title="Search Plazi for locations around (long -76.40004/lat -0.68315)">Estación Científica Yasuní</a> PUCE, 496 m, -0.68315, -76.40004, 12 - 2016. Winkler, Brian Four ” / “ Caterino DNA voucher, Ext. MSC-13224, Morphosp. 278673 ” / “ QCAZ -278673 ” . Paratype • 1 ♂: “ Ecuador F. co Orellana, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.876114&amp;materialsCitation.latitude=-0.6138889" title="Search Plazi for locations around (long -75.876114/lat -0.6138889)">Chiruisla Km 0</a>, 2, 218 m, 00°36'50"S, 75°52'34"W, 08-13DEC2005, J. Vieira ” / “ Ex: Winkler trap, Primary forest ” (QCAZ -I-280398) .</p><p>Diagnosis.</p><p>BL = 2.09 mm (n = 1). Body densely setose and brown, finely punctate above; head rounded at base, vertexal foveae impressed, without vertexal horns or tubercles at base; frons narrowing and rising to common antennal base; eyes reduced, with two or three ommatidia; gular teeth well-developed; antennae with scape very long, slightly sinuate at base, antennomere II (pedicel) slightly longer than III – VI combined, globose, narrow at base, widening towards apex, convex along outer margin, slightly concave on inner, protruding slightly at apical internal corner, III – VIII all bead-like, V and VII slightly larger and bearing elongate setae, IX – XI forming loose club, IX and X bearing elongate setae, XI densely setose; pronotum slightly longer than wide, widest near front, sides rounded; pronotal disk with lateral and median subbasal foveae; disk with strongly marked lateral longitudinal impressions; elytra rather short (wingless), humeri evenly sloped, sides rounded, each elytron with sutural and lateral dorsobasal foveae, lateral fovea with strong impression running posterad ~ 2 / 3 elytron length; posterior margin of elytron with outer submarginal tooth; subhumeral fovea absent; dorsal surface of abdomen distinctly punctate; protibia swollen; junction of mesepimeron and metaventrite deeply foveate; metatrochanter with inner margin slightly expanded, with blunt tooth at inner corner; penultimate ventrite distinctly depressed at middle, swollen at sides, last abdominal ventrite bearing small median apical marginal flange. Aedeagus (Fig. 5 G, H) with rather reduced basal bulb, slightly rounded dorsally, with oval dorsobasal diaphragm; basal apodemes forming wide elongate plate; tegmen not obviously articulated with basal bulb, dorsally subdivided into two asymmetric processes, one shorter, thinner strongly arched, the other longer, wider, curved ventrally to acute apex; additionally, below tegmen, two thin, curved, spine-like processes, project apically and basally from midpoint of tegmen, distal process flattened at base.</p><p>Distribution.</p><p>This species is known only from the Yasuní Biological Station in Amazonian Ecuador.</p><p>Remarks.</p><p>This species and the next are extremely similar in external morphology, and, coming from the same locality, were not initially considered to be distinct species. Both are also similar to the preceding species from the Rio Anzu area, ca 200 km west (and 1000 m higher in elevation). While the male genitalia separate these species clearly, distinguishing the two Yasuní species from each other based on external morphology is challenging (particularly given that both are known only from single males). In both, males have antennomere II asymmetrically swollen and the male metatrochanter apically dentate. In this species the penultimate male abdominal ventrite appears slightly lengthened and more distinctly depressed than in the following. The shape of the swollen second antennomere also differs slightly, being more distinctly swollen along the outer margin in this species, and less flanged at the inner apex. Also, the male metatrochanters are distinct, with that of B. amasanga basally toothed, that of B. yasuni markedly prolonged at the apex.</p><p>We have two female Bibrax specimens from Yasuní that appear to be distinct from each other. One of them was collected in the same sample as this species: the two individuals differ considerably in size, with a smaller one comparable in size to both B. amasanga and B. yasuni (described below). The second female specimen is significantly larger and differs in depth and length of the elytral impressions, as well as in minor details of antennomere lengths. At present it is impossible to associate either of these with either of the described species from this site: it is possible that the larger specimen represents yet a third species from the site. More material and possibly DNA sequences will be necessary to make definitive associations.</p><p>Etymology.</p><p>This species’ name refers to a mythical ‘ spirit of the jungle’, Amasanga, who came to the people in dreams and taught them to hunt</p></div>	https://treatment.plazi.org/id/00DE5EF153B15290AEDC4BD1012614FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
1C80667BA7EB59C3B427704DD50338B4.text	1C80667BA7EB59C3B427704DD50338B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax arachnoides Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax arachnoides sp. nov.</p><p>Figs 5 E, F, 7 A – D</p><p>Type material.</p><p>Holotype • ♂ (MECN -EN 40778): “ Ecuador: Pastaza, -1.4267, -78.0439, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.0439&amp;materialsCitation.latitude=-1.4267" title="Search Plazi for locations around (long -78.0439/lat -1.4267)">Rio Anzu Reserve</a>, 1342 m, 11.XI.2024, M. Caterino, sifted litter ” / “ Caterino DNA voucher, Ext. MSC-13131, Morphosp. RAn.014 ” / “ MECN -EN 40778 ” . Paratypes (6 ♂, 1 ♀) • Ecuador: Pastaza, Sumak Kawsay in situ Reserve, 26-JAN-2024, Díaz-Guevara (MECN -EN 23781 -23787)</p><p>Diagnosis.</p><p>BL = 1.68 mm (4). Body densely setose, densely and finely punctate beneath; head rounded at base, vertexal foveae shallowly impressed, short vertexal tubercles present at base; frons narrowing and rising to common antennal base, with shallow median sulcus on surface; eyes (of male) strongly reduced, comprising short longitudinal series of two or three indistinct facets; gular teeth well-developed; antennae with scape very long, slightly sinuate, antennomere II (pedicel) slightly longer than II – V combined, in male slightly convex on outer margin, produced at inner apical corner, III – VIII all bead-like, but V and VII larger and bearing elongate setae, IX – XI forming loose club, X and XI slightly more closely associated than IX and X; pronotum elongate, ~ 1.25 × longer than wide, widest near front, sides rounded; pronotal disk with lateral and median subbasal foveae; disk with only weak lateral longitudinal impressions; elytra rather short (wingless), humeri evenly sloped, sides rounded, each elytron with sutural and lateral basal foveae, lateral fovea with strong impression running posterad ~ 1 / 3 elytron length; subhumeral fovea absent; protibia swollen; male metatrochanter with inner margin expanded, subquadrate, distal corner projecting perpendicularly as small, truncate tooth; last male abdominal ventrite weakly depressed, bearing small basal marginal tooth and larger median apical marginal flange; penultimate male ventrite not obviously modified. Aedeagus (Fig. 5 E, F) with subspherical basal bulb, bearing large, round dorsal diaphragm; basal apodemes paired, thin, slightly curved; tegmen short, widened from base to middle, sclerotized portion bent strongly ventrad into a hook with deeply divided, densely bristled lobes extending laterodistad; between them an unsclerotized, setose median lobe extends from tip.</p><p>Distribution.</p><p>This species is known from two localities in the foothills of the Andes, in western Pastaza Province, Ecuador.</p><p>Remarks.</p><p>This species and the next two are very similar and share the probable synapomorphy of a swollen antennomere II. They may share this synapomorphy with B. chullachaqui, which is otherwise quite distinct in its large eyes, short body setae, and various other characters. Bibrax arachnoides has slightly different modifications of the last male abdominal ventrite (with a small basal tooth), metatrochanter (more generally subquadrate along apical margin with a perpendicular distal tooth), and antennomere II (more or less symmetrically swollen). The male genitalia of all these species are unambiguously distinct.</p><p>Etymology.</p><p>We name this species for the somewhat spider-like appearance, common to members of Bibrax, as a result of their geniculate antennae appearing to be an extra pair of legs.</p></div>	https://treatment.plazi.org/id/1C80667BA7EB59C3B427704DD50338B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
F6DB8F00AF395D00BA52115553932086.text	F6DB8F00AF395D00BA52115553932086.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax aratrifer Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax aratrifer sp. nov.</p><p>Figs 2 G, H, 3 E – H</p><p>Type material.</p><p>Holotype • ♂ (MECN -EN 38735): “ Ecuador: Napo, -0.5952, -77.8994, Est. Biol. Yanayacu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.8994&amp;materialsCitation.latitude=-0.5952" title="Search Plazi for locations around (long -77.8994/lat -0.5952)">Piha Tr.</a>, 2386 m, 9.IX.2024, sifted litter, M. Caterino ” / “ MECN -EN 38735 ” . Paratypes (4 ♂, 1 ♀, same general locality as type) • 1: same data as type • 1: -0.5983, -77.8954, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.8954&amp;materialsCitation.latitude=-0.5983" title="Search Plazi for locations around (long -77.8954/lat -0.5983)">Stream Tr.</a>, 2192 m • 1: -0.5986, -77.8949, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.8949&amp;materialsCitation.latitude=-0.5986" title="Search Plazi for locations around (long -77.8949/lat -0.5986)">Stream Tr.</a>, 2185 m • 1: -0.5958, -77.8927, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.8927&amp;materialsCitation.latitude=-0.5958" title="Search Plazi for locations around (long -77.8927/lat -0.5958)">Aburrian Tr.</a>, 2283 m • 1: -0.5947, -77.8945, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.8945&amp;materialsCitation.latitude=-0.5947" title="Search Plazi for locations around (long -77.8945/lat -0.5947)">Piha Tr.</a>, 2248 m (MECN -EN 38736 -38739, 41045) .</p><p>Diagnosis.</p><p>BL = 1.32 mm (n = 3). Eyes of both sexes with single facet, head subquadrate posteriorly, prolonged in front of eyes, deeply depressed at vertexal foveae, slightly elevated posteriorly; antennal base prolonged and narrowed anterad; antennal scape sinuate, narrowed to base and before apex, flagellomeres II – VI progressively shorter, VII slightly larger, VIII small, IX – XI forming weak, loose club; pronotum with lateral lobes rounded, median and lateral longitudinal pronotal sulci well developed; elytra very short (flightless in both sexes), together emarginate along posterior margin, each with single distinct, median basal fovea, with short longitudinal elytral sulci short extending posterad; male with visible abdominal ventrite 2 elevated at middle, elevation posteriorly concave, densely setose; male ventrites 3 and 4 simply depressed medially; other secondary sexual characters not evident. Aedeagus (Fig. 2 G, H) with large basal bulb with round dorsal diaphragm, bulb ventrally bearing strong basal shelf (which articulates internally with sternite 6); tegmen constricted at base, appearing articulated with basal bulb, bent strongly laterad, thence narrowing evenly distad, apex with very small spatulate tip.</p><p>Distribution.</p><p>This species is known only from the cloud forests of the Yanayacu Biological Station, Napo province, Ecuador.</p><p>Remarks.</p><p>Among the smaller, flattened, and flightless species mentioned previously, the unique, elevated, curved transverse ridge on male ventrite 2 easily distinguish this species.</p><p>Etymology.</p><p>The name of this species refers to this abdominal modification, meaning in Latin ‘ plow-bearing’.</p></div>	https://treatment.plazi.org/id/F6DB8F00AF395D00BA52115553932086	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
86E21074F99F59998C2EC0991D6198FF.text	86E21074F99F59998C2EC0991D6198FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax canelazo Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax canelazo sp. nov.</p><p>Figs 5 A, B, 6 A – D</p><p>Type material.</p><p>Holotype • ♂ (MECN -EN 40836): “ Ecuador: Tungurahua, -1.4361, -78.3105, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.3105&amp;materialsCitation.latitude=-1.4361" title="Search Plazi for locations around (long -78.3105/lat -1.4361)">NaturetrekCandelariaRes</a>, 2241 m, 13.XI.2024, M. Caterino, sifted litter ” / “ Caterino DNA voucher, Ext. MSC-13189, Morphosp. Cnd.005 ” / “ MECN -EN 40836 ” . Paratypes (1 ♂ 1 ♀, same general locality as type) • 1: -1.4343, -78.3114, 2145 m • 1: -1.4361, -78.3105, 2241 m (MECN -EN 40332, 40379) .</p><p>Additional material.</p><p>• 1 ♂ (MECN -EN 40794): “ Ecuador: Tungurahua, -1.3812, -78.2895, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.2895&amp;materialsCitation.latitude=-1.3812" title="Search Plazi for locations around (long -78.2895/lat -1.3812)">Rio Machay Reserve</a>, 2416 m, 12.XI.2024, M. Caterino, sifted litter ” / “ Caterino DNA voucher, Ext. MSC-13147, Morphosp. Mch.010 ; (1 ♂ 8 ♀, all Rio Machay Reserve) • 3: same data as preceding • 4: -1.3895, -78.2929, 2239 m • 1: -1.3801, -78.2889, 2434 m • 1: -1.3829, -78.2909, 2382 m (MECN -EN 40190, 40206, 40215, 40220, 40229, 40283, 40285, 40292, 40296) .</p><p>Diagnosis.</p><p>BL = 1.10 mm (n = 2). Body identical in most respects to B. onorei, with the following exceptions: antennomeres II-IV slightly more slender, elongate, antennomere III, in particular, nearly as long antennomere II; median longitudinal pronotal sulcus more deeply impressed; male with only last two abdominal ventrites modified, penultimate with weak depression in distal third, weakly margined by transverse carina, last ventrite short, concave, with small slightly transverse apical marginal tooth. Aedeagus (Fig. 5 A, B) with basal bulb short, distal surface flat, bearing large oblique diaphragm on left dorsolateral surface; with pair of basal apodemes well separated, obliquely truncate with outer edge longer; tegmen trunk-like, basally cylindrical, extending distad from middle of apical surface of basal bulb, bent ventrad, flattening toward broad paddle-shaped apex; tegmen with short ventral tooth and shorter, weakly sclerotized lobe extending distad below apex of tegmen.</p><p>Distribution.</p><p>This species is known from cloud forests of two localities at similar elevations on either side of the Rio Pastaza in Tungurahua province, Ecuador, both of them reserves managed by the EcoMinga Foundation.</p><p>Remarks.</p><p>There is little to separate this species externally from other small, flattened species described here. It is relatively small, and the male abdominal modifications are comparatively subtle, with only a small distal tooth at the apex of a shallowly depressed last ventrite. There is slight variation in aedeagal shape between the two localities, the ‘ apical paddle’ of the type population more elongate, narrower, and apically rounded, while that at Rio Machay is shorter, more quadrate, and pointed at middle. But for the present we prefer to recognize this as intraspecific variation. Discovery of additional populations of either form would allow further assessment of their respective status.</p><p>Etymology.</p><p>The name of this species refers to the popular Ecuadorian beverage, canelazo, made and enjoyed during Fiesta de Quito, an annual festival happening around the time of the discovery of this species (late November). The name also echoes the type locality Candelaria.</p></div>	https://treatment.plazi.org/id/86E21074F99F59998C2EC0991D6198FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
309305BED1D556498EC334F52D49FBB4.text	309305BED1D556498EC334F52D49FBB4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax cerroblanco Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax cerroblanco sp. nov.</p><p>Figs 2 K, L, 4 E – H</p><p>Type material.</p><p>Holotype • ♂ (MECN -EN 38548): “ Ecuador: Guayas, -2.159579, -80.02109, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.02109&amp;materialsCitation.latitude=-2.159579" title="Search Plazi for locations around (long -80.02109/lat -2.159579)">Bosque Protector Cerro Blanco</a>, Q. Cusumbo, 25.IV.2022, Winkler, C. Cujigualpa ” / “ Caterino DNA voucher, Ext. MSC-13079, Morphosp. CrBl.A.009 ” / “ MECN -EN 38548 ” . Paratypes (2 ♂, 5 ♀) • same data as type (MECN -EN 38545 -38547, 38549-38552) .</p><p>Diagnosis.</p><p>BL = 1.89 mm (n = 3). Body with dense, matte-like pubescence; head with single large ommatidium on each side (in both sexes); vertexal foveae shallowly impressed; head rounded posteriorly, prolonged anteriorly into narrowly elongate antennal base; gular teeth well-developed; scape very elongate, weakly curved at base, male antennomere II (pedicel), slightly swollen, ~ 2 × as long as III, evenly widened to apex, antennomeres III – V ~ 1.5 × longer than wide, VI – VIII shorter, VII slightly wider than VI or VIII; IX – XI slightly larger, forming loose, weak club; pronotum elongate, ~ 1.5 × longer than wide, widest just anterior of midpoint, evenly narrowed to base and apex; lateral pronotal foveae deeply impressed, setose, joined by weak transverse sulcus that passes through smaller, nude median fovea, continuing laterad to weakly interrupt lateral margin; shallow longitudinal impressions extend anterad and posterad from lateral and median foveae forming weak sulci; elytra narrow at base (wingless), widening to apex, each with sutural and one lateral dorsobasal foveae, shallow sulci extending posterad from foveae; subhumeral fovea absent; wingless in both sexes; protibiae strongly swollen; male ventrite 5 with broad, shallowly concave, smooth median area, male ventrite 6 with minute median, apical tooth. Aedeagus (Fig. 2 K, L) with basal bulb large, irregularly subspherical, with prominent basal apodeme; diaphragm located dorsobasally; tegmen dorsoventrally flattened, widened from base as it curves dorsad and distad, apex split ~ 1 / 4 from tip, one side with serrate margin narrowing to acute point, other side slightly longer and ending in slightly expanded, blunt apical plate.</p><p>Distribution.</p><p>This species is known only from Ecuador’s coastal mountains just northwest of Guayaquil.</p><p>Remarks.</p><p>This species is among the more convex, less strongly flattened species, having the pronotum gradually narrowed posteriorly, not abruptly constricted behind prominent lateral lobes. Unlike several of these, it does not have male antennomere II expanded, nor does it exhibit modifications of the male metatrochanter. The combination of these, along with a male penultimate metaventrite that is broadly and shallow concave, will distinguish it from any other species currently known. The aedeagus, having a flattened, shallowly bifid tegmen in which the right lobe is marginally denticulate, is highly distinctive.</p><p>Etymology.</p><p>We name this species for the protected area in which it occurs, El Bosque Protector Cerro Blanco. Semi-surrounded by the greater Guayaquil metropolitan area, this oasis of biodiversity has exceptional conservation value.</p></div>	https://treatment.plazi.org/id/309305BED1D556498EC334F52D49FBB4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
2E8A8F1D989E58C89F7E907D80309570.text	2E8A8F1D989E58C89F7E907D80309570.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax chocoensis Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax chocoensis sp. nov.</p><p>Figs 1 D – F, 2 C, D</p><p>Type material.</p><p>Holotype • ♂ (MECN -EN 40872): “ Ecuador: Esmeraldas, 0.56759, -79.06351, Quinindé. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.06351&amp;materialsCitation.latitude=0.56759" title="Search Plazi for locations around (long -79.06351/lat 0.56759)">Reserva Biológica Canandé</a>. 400 m. 14-oct-2022. Winkler., A Pazmiño | D Díaz. ” / “ Caterino DNA voucher, Ext. MSC-13015, Morphosp. Can.A.001 ” / “ MECN -EN 40872 ”.</p><p>Diagnosis.</p><p>BL = 1.79 mm (n = 1). Body densely setose, densely and finely punctate beneath; head rounded at base, vertexal foveae shallowly impressed; frons elevated, elongate to common antennal base, with shallow median sulcus on surface; eyes (of male) well developed, convex, with&gt; 20 fine facets; gular teeth well-developed; antennae with scape very long, slightly sinuate, antennomere II (pedicel) as long as III-VI combined, III – VIII all bead-like, but V and VII larger, IX – XI forming loose club; pronotum elongate, widest near front, with lateral and median subbasal foveae, as well as row of small secondary foveae along basal margin; disk with lateral and median longitudinal impressions; elytra moderately elongate (winged) with sides rounded, each elytron with sutural and lateral foveae, lateral fovea with strong impression running posterad ~ 1 / 2 elytron length; subhumeral fovea absent; protibia swollen, with modified inner marginal spurs; last abdominal ventrite slightly flattened, densely setose on each side. Aedeagus (Fig. 2 C, D) with large basal bulb bearing prominent basal flange where it articulates with sternite VI; tegmen articulated with basal bulb, tubular at base, strongly arched dorsad, then curved distad and flattening and narrowing to subacute apex.</p><p>Distribution.</p><p>This species is known only from the coastal forests of Esmeraldas, Ecuador.</p><p>Remarks.</p><p>This species more closely resembles the described species, B. bradleyi in particular, in having large male eyes, apparently well-developed flight wings (at least in the male), and pronotum without lobed sides. Its pronotum is more elongate than either B. bradleyi or B. popeye, and it exhibits distinct secondary sexual characters of the protibial spurs, while lacking metatibial modifications.</p><p>Etymology.</p><p>The name of this species refers to the Chocó biodiversity hotspot, a biome largely limited to northwestern Ecuador and neighboring Colombia, where this species is found.</p></div>	https://treatment.plazi.org/id/2E8A8F1D989E58C89F7E907D80309570	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
2E7D8CFDB4D15B2CAB57E0731658DCC3.text	2E7D8CFDB4D15B2CAB57E0731658DCC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax chullachaqui Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax chullachaqui sp. nov.</p><p>Figs 1 A – C, 2 A, B</p><p>Type material.</p><p>Holotype • ♂ (MECN -EN 23779): “ Ecuador. Napo, Archidona. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.59197&amp;materialsCitation.latitude=-0.658219" title="Search Plazi for locations around (long -77.59197/lat -0.658219)">Pacto Sumaco</a>, -0.658219, -77.59197. Malaise, 22-ene-2024. A Pazmiño | M Barreno ” / “ Caterino DNA voucher, Ext. MSC-13221, Morphosp. PS.A.003 ” / “ MECN -EN 23779 ”.</p><p>Diagnosis.</p><p>BL = 1.89 mm (n = 1); body densely setose and brownish-orange; eyes prominent, borne on small prominence, comprising ~ 26 ommatidia; gular teeth well-developed, curved slightly dorsad; head rounded posteriorly, prolonged in front of eyes to prominent antennal base, vertexal foveae slightly impressed; frons finely punctate above; antennal scape thick and sinuate, slightly narrowed before apex (in lateral view), antennomere II (pedicel) swollen, approximately as long as III – V together, III – V similar in size, short, VI and VIII shorter, VII wider than long, IX – XI forming a club, IX and X transverse, X slightly wider than IX, X, and XI closely associated; pronotum rounded, widest anterad, with lateral lobes slightly produced; lateral longitudinal pronotal impressions very shallowly impressed; elytra longer than wide, slightly widened beyond midpoint, each with sutural and one lateral basal foveae, lateral with shallow longitudinal impression extending posteriorly ~ 2 / 3 elytron length; subhumeral fovea absent; flight wings present protibia swollen; metatrochanter not modified, metaventrite between second and third pair of legs with oblique carinae posterad a shallow median depression; last abdominal ventrite bearing small median apical marginal flange; third ventrite slightly flattened, posterior margin straight at middle. Aedeagus (Fig. 2 A, B) with basal bulb large, elongate oval, with blunt, apically knobbed pair of basal apodemes; diaphragm rather small, round, located just basad tegmen articulation; tegmen widened from base as it curves dorsad and distad, apex divided, with an elongate, sinuate subacute process extending beneath shorter, broad dorsal brush.</p><p>Distribution.</p><p>This species is known only from the lower (~ 1500 m), southern slopes of the Volcán Sumaco, in the western Amazon, Napo Province, Ecuador.</p><p>Remarks.</p><p>This species is most similar to B. bradleyi and B. popeye in having well-developed eyes and flight wings (at least in the male sex), while having the pronotum rather simply expanded toward the front, with only weak longitudinal sulci. It is distinctive in its dense covering of short, stiff setae, while the short swollen second antennomere and compact antennal club are also distinctive. While lacking any secondary sexual characters on the legs, the depressed male metaventrite is unusual.</p><p>Etymology.</p><p>The species is named after Chullachaki, a mythical ‘ one-footed’ spirit of the Amazonian jungle. He is purportedly able to change into any type of rainforest animal, guarding the forest against unwise uses.</p></div>	https://treatment.plazi.org/id/2E7D8CFDB4D15B2CAB57E0731658DCC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
B4E0DBE183275AF69739B85FDCAC0924.text	B4E0DBE183275AF69739B85FDCAC0924.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax Fletcher 1927	<div><p>Bibrax Fletcher, 1927</p><p>Type species.</p><p>Bibrax bradleyi Fletcher, 1927, by original designation. Type locality. Barro Colorado Island, Panama.</p><p>Diagnosis.</p><p>Among Metopiasini, Bibrax has been diagnosed (e. g., by Asenjo et al. 2024; Mario Chaul and Lopes-Andrade 2024; Chandler, unpublished) by having a scape as long as the funicular antennomeres combined (unlike Rhinoscepsis), the abdomen laterally margined, with distinct paratergites (unlike Metopioxys and Metopiosoma), the second antennomere or pedicel (at least up until now) at least 2 × as long as the third, the lack of acute pronotal and vertexal tubercles (seen in Metopiellus), having distinct lateral and / or median longitudinal pronotal sulci, having abdominal sternite III (first visible) longer than the metacoxae, and having the anterolateral corners of the gula acutely projecting (the size of which are, in light of new species, quite varied).</p><p>Secondary sexual characters, when present, frequently involve modifications of the male abdominal sternites, the male tibial apices, male antennal pedicel, and male trochanters. In some species the sexes are dimorphic in eye size, although in several species described below reduced eyes are present in both sexes. The male genitalia, while highly varied in particulars, are consistent in having a large basal bulb that is articulated with an elongate distal tegmen. Parameres are absent. Asenjo et al (2024) demonstrated the basal articulation of the basal bulb with an “ apodemal plate ” on the inner surface of sternite VIII, and all of the species have a variously developed process (sometimes paired) on the basal bulb for this purpose. The tegmen can be simple or quite elaborate, sometimes subdivided, and is highly diagnostic of each species.</p></div>	https://treatment.plazi.org/id/B4E0DBE183275AF69739B85FDCAC0924	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
C4ACA30D22C05DC4B1D6FC5EE544C4D4.text	C4ACA30D22C05DC4B1D6FC5EE544C4D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax grandis Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax grandis sp. nov.</p><p>Fig. 8 F – H</p><p>Type material.</p><p>Holotype • ♀ (MECN -EN 40146): “ Ecuador: Cotopaxi, -0.4168, -79.0044, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.0044&amp;materialsCitation.latitude=-0.4168" title="Search Plazi for locations around (long -79.0044/lat -0.4168)">Bosque Integral Otonga</a>, 14.VIII.2024, 2093 m, M. Caterino | A. Pazmiño, Sifted leaf litter ” / “ Caterino DNA voucher, Ext. MSC-12889, Morphosp. Ot.A.054 ” / “ MECN -EN 40146 ”.</p><p>Diagnosis.</p><p>BL = 1.73 mm (n = 1). Body large, very densely setose; eyes (of female) with multiple facets, head rounded at sides, short anterior to eyes, with pair of lyriform carinae; gular teeth well-developed; antennal tubercle short, wider apically; antennal scape cylindrical, coarsely punctate, antennomeres II – V more slightly enlarged and more densely setose than VI – X; pronotum with anterior lobes weakly angulate laterally, median and lateral longitudinal sulci deeply impressed; elytra moderately long, not together emarginate along posterior margin; dorsal elytral sulci (from mid-basal foveae) distinct, reaching just beyond elytral midpoint; abdominal tergites densely setose; posterior corners of abdominal laterotergites rounded.</p><p>Male. Unknown.</p><p>Distribution.</p><p>This species is known only from the higher elevations of the Bosque Integral Otonga, in northern Cotapaxi province, Ecuador.</p><p>Remarks.</p><p>This species is highly distinctive among known Bibrax species. Although known only from a single female, it is unusual enough to warrant recognition. It is most distinctive in its much shorter antennal base, lyriform frontal carinae, and produced subangulate pronotal sides. While the eyes are reduced, they exhibit a few more ommatidia than most of the reduced-eye species described here. The male might be expected to differ in eye size, and its discovery would also permit assessment of some other phylogenetically useful characters, such as presence of secondary sexual characters of the abdominal ventrites, mesotibiae, and metatrochanters.</p><p>Etymology.</p><p>While appearing related to many of the smaller, flattened species described in this study, this species is considerably larger than most of them, leading to our naming it B. grandis .</p></div>	https://treatment.plazi.org/id/C4ACA30D22C05DC4B1D6FC5EE544C4D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
F90D17D8E11352CBB86039883BA851B7.text	F90D17D8E11352CBB86039883BA851B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax longiventer Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax longiventer sp. nov.</p><p>Figs 2 I, J, 4 A – D</p><p>Type material.</p><p>Holotype • ♂ (MECN -EN 40723): “ Ecuador: Napo, -0.61523, -77.59071, Archidona, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.59071&amp;materialsCitation.latitude=-0.61523" title="Search Plazi for locations around (long -77.59071/lat -0.61523)">Pacto Sumaco</a>, 1805 m, 27.IX.2024, A. Pazmiño &amp; D. Díaz, Winkler ” / “ Caterino DNA voucher, Ext. MSC-13068, Morphosp. VS.007 ” / “ MECN -EN 40723 ” . Paratypes (2 ♂ 2 ♀) • same data as type (MECN -EN 40639, 40608, 40651, 40655) .</p><p>Diagnosis.</p><p>BL = 1.88 mm (n = 3). Body rather large, densely setose, reddish-orange; eyes of both sexes strongly reduced, borne on small prominence, comprising two or three indistinct ommatidia; head subquadrate posteriorly, prolonged in front of eyes to prominent antennal base, deeply depressed at vertexal foveae, slightly elevated posteriorly; antennal scape sinuate, narrowed to base and before apex, flagellomeres II – VI progressively shorter, VII slightly larger, VIII small, IX – XI forming weak, loose club; gular teeth not prominent; pronotum with lateral lobes strongly produced, rounded, pronotum strongly constricted posteriorly; median and lateral longitudinal pronotal impressions well developed; elytra rather short, with narrowed humeri (flightless), each with sutural and one lateral basal foveae, lateral fovea with short longitudinal impression extending posterad; abdominal paratergites wide; protibia swollen; male metaventrite with small flat glabrous area between and anterad metacoxae; male abdominal segment 5 markedly elongate, sternite especially, prolonged anterad into emarginate ventrite 4, deeply concave, with small, tubular denticle at basal margin; legs without obvious secondary sexual characters. Aedeagus (Fig. 2 I, J) with basal bulb large, basally truncate, dorsally angulate, with long basal apodeme; tegmen comprising two separate processes, a shorter, strongly arched and apically acute rod on left side; right-side process longer, terminating in dorsoventrally flattened, subrectangular plate.</p><p>Distribution.</p><p>This species is known only from the isolated Volcan Sumaco, in the western Amazonian basin, Ecuador.</p><p>Remarks.</p><p>The remarkably prolonged male sternite 5 in this species is unique among known Bibrax, while the strongly laterally lobed (and posteriorly constricted) pronotum may also help distinguish females from those of other species. While among the ‘ smaller’, flattened species described herein, this species is also larger than most others.</p><p>Etymology.</p><p>The name of this species refers to the uniquely elongate last abdominal ventrite.</p></div>	https://treatment.plazi.org/id/F90D17D8E11352CBB86039883BA851B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
512A30269CAC5FE98BAB9DE0F3D2D47A.text	512A30269CAC5FE98BAB9DE0F3D2D47A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax onorei Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax onorei sp. nov.</p><p>Figs 2 E, F, 3 A – D</p><p>Type material.</p><p>Holotype • ♂ (MECN -EN 40873): “ Ecuador: Cotopaxi, -0.4176, -79.0040, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.004&amp;materialsCitation.latitude=-0.4176" title="Search Plazi for locations around (long -79.004/lat -0.4176)">Bosque Integral Otonga</a>, 13.VIII.2024, 2063 m, M. Caterino &amp; A. Pazmiño, Sifted leaf litter ” / “ Caterino DNA voucher, Ext. MSC-12844, Morphosp. Ot.A.009 ” / “ MECN -EN 40873 ” . Paratypes (3 ♂, 5 ♀, same general locality as type) • 2: same data as type • 3: -0.4167, -79.0043, 2097 m • 1: -0.4192, -79.0031, 1966 m • 1: -0.4172, -79.0040, 2073 m • 1: -0.4169, -79.0042, 2086 m (MECN -EN 38587 -38593, 40897) .</p><p>Diagnosis.</p><p>BL = 1.10 mm (n = 3). Eyes of both sexes with single facet, head subquadrate posteriorly, with transverse basal ridge on vertex, prolonged in front of eyes; antennal base prolonged and narrowed anterad, antennal insertions swollen; antennal scape sinuate, narrowed to base and before apex, antennomere II (pedicel) slightly swollen, ~ 1.5 × as long as III (Fig. 3 A – C), III conical, IV – VIII bead-like, alternating slightly in size (IV, VI, VIII smaller), IX – XI forming club, X and XI more closely associated than IX with X; gular processes evident but not large; pronotum with anterior lobes rounded (Fig. 3 A), strongly constricted posteriorly; lateral longitudinal sulci conspicuous; median pronotal sulcus weak; elytra very short, together emarginate along posterior margin; dorsal elytral sulci short and weakly developed; abdominal paratergites with posterior corners angulate; male with visible abdominal ventrites 2–4 depressed medially (Fig. 3 D); male lacking modification of mesotibial apex. Aedeagus (Fig. 2 E, F) with short, broad basal bulb, with oval dorsobasal diaphragm, diaphragmatic sclerite indistinct; short digitiform process extending from left, middle edge of basal bulb; slightly longer, narrow apically quadrate process extending from medioventral margin; longest process extending out of middle of basal bulb (articulated), a well sclerotized, basally bent, apically narrowed and slightly sinuate blade, terminating in blunt and weakly tufted apex; dorsally an articulated, weakly sclerotized process extends over basal two-thirds of median blade.</p><p>Distribution.</p><p>This species is known only from the higher elevations of the Bosque Integral Otonga, in northern Cotopaxi province, Ecuador.</p><p>Remarks.</p><p>This species resembles several other new species described here, all of which differ considerably from any previously described ones. The species in this group ( B. bradleyi, B. aratrifer, B. longiventer, B. cerroblanco, B. canelazo, B. pectinifer, B. arachnoides, B. amasanga, and B. yasuni) share strongly reduced eyes and lack of wings in both sexes, a small, somewhat flattened body, strongly impressed lateral longitudinal pronotal sulci that set off distinct lateral pronotal lobes, and exhibit modified abdominal ventrites in the males (Fig. 3 D). It is principally in the latter characters (as well as marked differences in male genitalia (Fig. 2 E, F) that distinguish the species.</p><p>Etymology.</p><p>This species name honors Dr. Giovanni Onore, a pioneer of entomology in Ecuador, who facilitated the fieldwork and worked to protect the site that led to the discovery of this species.</p></div>	https://treatment.plazi.org/id/512A30269CAC5FE98BAB9DE0F3D2D47A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
B095B9DDDBB45AD58693B73FC6D8B2E1.text	B095B9DDDBB45AD58693B73FC6D8B2E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax pectinifer Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax pectinifer sp. nov.</p><p>Figs 5 C, D, 6 E – H</p><p>Type material.</p><p>Holotype • ♂ (MECN -EN 23776): “ Ecuador. Napo, Archidona. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.59197&amp;materialsCitation.latitude=-0.658219" title="Search Plazi for locations around (long -77.59197/lat -0.658219)">Pacto Sumaco</a>, -0.658219, -77.59197. Malaise, 22-ene-2024. A Pazmiño | M Barreno ” / “ Caterino DNA voucher, Ext. MSC-13222, Morphosp. PS.A.004 ” / “ MECN -EN 23776 ” . Paratypes (3 ♂ 1 ♀) • same data as type (MECN -EN 23774, 23777, 23775, 23778) .</p><p>Diagnosis.</p><p>BL = 1.55 mm (n = 3). Body densely setose, brownish-orange; eyes of both sexes strongly reduced, borne on small prominences, comprising three or four indistinct ommatidia; head rounded posteriorly, prolonged in front of eyes to prominent antennal base, two deep depressions in lateral view: one in front of eyes and another one at vertexal fovea; antennal scape slightly sinuate, narrowed anterior to apex, antennomere II (pedicel) longer than antennomeres III – IV combined, III – VI progressively shorter, VII slightly larger, VIII small, IX and X similar sized, IX – XI forming club, antennomere XI rounded and slightly longer than antennomeres IX and X; gular teeth not prominent; pronotum with lateral lobes slightly produced, rounded, pronotum weakly constricted and gradually narrowed to base; median and lateral longitudinal pronotal impressions well developed; elytra rather short, with narrowed humeri, each with sutural and one lateral basal foveae, lateral fovea with broad longitudinal impression extending posterad; abdominal paratergites wide, projecting at posterior corners; penultimate male abdominal ventrite weakly depressed at middle with comb-like series of erect setae along apicolateral margins; last ventrite with similar setal comb less prominent. Aedeagus (Fig. 5 C, D) with basal bulb large and spherical, with pair of short, blunt basal apodemes; tegmen deeply divided into two separate processes of similar lengths, one sinuate, with concave apical margin, the other with spine-like end and curved dorsally.</p><p>Distribution.</p><p>This species is known only from the lower (~ 1500 m), southern slopes of the Volcán Sumaco, in the western Amazon, Napo Province, Ecuador.</p><p>Remarks.</p><p>This species is among the smaller, flattened species, and accordingly differs mainly in the male abdominal characters. The paired combs of setae on the last two male ventrites are unique and readily distinguish the species, although these combs may be somewhat obscured by adhering particles of dirt.</p><p>Etymology.</p><p>We name this species for its males’ ‘ comb-bearing’ abdominal sternites.</p></div>	https://treatment.plazi.org/id/B095B9DDDBB45AD58693B73FC6D8B2E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
6C0A52DE851F54FCA7E1171010F196E4.text	6C0A52DE851F54FCA7E1171010F196E4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bibrax yasuni Tapuy-Avilés & Díaz-Guevara & Caterino 2025	<div><p>Bibrax yasuni sp. nov.</p><p>Figs 5 I, J, 8 A – E</p><p>Type material.</p><p>Holotype • ♂ (QCAZ -I-278674): “ Ecuador. Orellana, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.40004&amp;materialsCitation.latitude=-0.68315" title="Search Plazi for locations around (long -76.40004/lat -0.68315)">Estación Científica Yasuní</a> PUCE, 253 m, -0.68315, -76.40004, 09 - 2017. Winkler., A. Argoti &amp; R. E. Cárdenas ” / “ Caterino DNA voucher, Ext. MSC-13225, Morphosp. 278674 ” / “ QCAZ -278674 ” . Paratype • 1 ♂: “ Ecuador: Yasuni National Park, Yasuni Biological Station, 0°40'32"S, 76°23'50"W ” / “ 29 June 1999, Berlese, 15 km West of station, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.39722&amp;materialsCitation.latitude=-0.6755556" title="Search Plazi for locations around (long -76.39722/lat -0.6755556)">Monkey Plot</a>, CEC # 034, CECarlton ” (QCAZ -I-280395) .</p><p>Diagnosis.</p><p>BL = 1.76 mm (n = 1). Body setose and reddish, finely punctate above; head rounded posteriorly, vertexal foveae slightly impressed; frons narrowing and rising to common antennal base; eyes reduced, with two or three ommatidia; gular teeth well-developed; scape elongate, weakly sinuate, antennomere II (pedicel) slightly longer than III – VI combined, narrow at base and widening towards distal end, with outer margin straight, inner apical angle protruding forming flattened, slightly translucent, rounded spatulate process, III-VIII bead-like, V and VII slightly longer and bearing elongate setae, antennomeres IX – XI forming loose club, IX and X slightly transverse and bearing elongate setae, XI densely setose; pronotum elongate, widest near front, sides rounded; pronotal disk with lateral and median subbasal foveae; disk with strongly marked lateral longitudinal impressions; elytra rather short (wingless), sides rounded, each elytron with sutural and lateral basal foveae, lateral fovea with strong impression running posterad; subhumeral fovea absent; outer posterior corner of each elytron with tooth; dorsal surface of abdomen largely impunctate; protibia swollen; metatrochanter with distal corner extended as strong spine; metafemora swollen, distinctly clavate; last abdominal ventrite bearing small median apical marginal flange; penultimate ventrite shorter, barely depressed at middle. Aedeagus (Fig. 5 I, J) with round basal diaphragm, short and irregularly subspherical basal bulb; basal apodemes conjoined, broad, flat, with prolonged apical corners; tegmen narrow and tubular at base, expanded distally, dorsally hooded, with strong, blunt ventral keel, two distal processes that are similar in length extend from ventral apex, one thinner, slightly sinuate and apically subacute, the other wider, with basal brush of setae along outer margin, apically bifid with sclerotized, acute tip and secondary, setose, unsclerotized apical lobe; expansive membranous internal sac surrounding much of tegmen apex.</p><p>Distribution.</p><p>This species is known only from the Yasuní Biological Station in Amazonian Ecuador.</p><p>Remarks.</p><p>As discussed above, this species is very similar to the preceding. Aside from the distinctive male genitalia, B. yasuni is most distinct in the shape of antennomere II of the male, having a straight outer margin and flanged inner apical corner, the male penultimate abdominal ventrite, being shorter and barely depressed at the middle, and in the apically prolonged male metatrochanter.</p><p>Etymology.</p><p>This species is named for the Yasuní Biological Station, its type locality.</p></div>	https://treatment.plazi.org/id/6C0A52DE851F54FCA7E1171010F196E4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Tapuy-Avilés, Yarina;Díaz-Guevara, David R.;Caterino, Michael S.	Tapuy-Avilés, Yarina, Díaz-Guevara, David R., Caterino, Michael S. (2025): First record of Bibrax Fletcher, 1927 (Coleoptera, Staphylinidae, Pselaphinae) from Ecuador, with descriptions of twelve new species. ZooKeys 1250: 105-133, DOI: 10.3897/zookeys.1250.156763
