identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
CD5DE5502516433395CDC17239B7DAA3.text	CD5DE5502516433395CDC17239B7DAA3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phyllanthus catingicola D. Santos, Loiola & Marques-Torres 2025	<div><p>Phyllanthus catingicola D. Santos, Loiola &amp; Marques-Torres, sp. nov. (Figures 1, 2)</p><p>Type:— BRAZIL. Ceará, Chaval, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.254723&amp;materialsCitation.latitude=-3.0494444" title="Search Plazi for locations around (long -41.254723/lat -3.0494444)">Pedra da Santa</a>, 03º02’58”S, 41º15’17”W, 09 April 2024, fl., fr., F.D. Santos 871 (holotype: EAC!; isotypes: EAC!, HUEFS!, K!, PEUFR!, RB!, UFG!) .</p><p>Diagnosis: — Phyllanthus catingicola differs from P.hyssopifolioides by having a cylindrical main stem (vs. flattened in P. hyssopifolioides), 5-merous staminate flowers with elliptical sepals (vs. 6-merous and orbicular), pistillate flowers with patelliform disk and capitate stigma (vs. 6-lobed and punctiform).</p><p>Description: —Erect herbs, 7–12 cm tall, monoecious; non-phyllantoid branching. Main stem cylindrical, glabrous, non-papillose, greenish; secondary branches flat, winged, glabrous, non-papillose, greenish. Cataphylls absent. Stipules ca. 1 mm long, lanceolate, apex acute, base truncated, margin entire, membranaceous, surface smooth, glabrous, persistent. Leaves alternate distichous, present on the main stem and secondary branches; petiole 0.5 mm long, cylindrical, glabrous, non-papillose; leaf blade 0.5–0.8 × 3.0–5.0 mm, oval or ovate-elliptical, base rounded, apex obtuse, sometimes rounded, margin entire, glabrous on both sides, membranaceous, concolor; brochidodromous venation with 4 or 5 pairs of secondary veins, printed on both sides. Cymules unisexual, axillary distributed along the main stem and secondary branches. Bracts ca. 1 mm long, lanceolate, apex acute, base truncate, scarious. Staminate cymules with 1 to 3 flowers; Pistillate cymules with 1 or 2 flowers. Staminate flowers: pedicel 0.3–0.6 mm long, cylindrical, non-papillose, glabrous; sepals 5, 0.5–0.6 mm long, elliptical or elliptical-obovate, apex obtuse or rounded, disk with 5-segmentes obtriangular, surface smooth, stamens 3, free, anthers horizontally dehiscent. Pistillate flowers: pedicel 0.3–0.4 mm long, cylindrical, non-papillose, glabrous; sepals 6, sometimes 5, 0.4–0.5 mm long, oblong, apex obtuse or rounded, membranaceous, margin entire; patelliform disk; ovary ca. 0.25 × 0.25, globose-depressed, non-papillose, styles 3, bifid, free, patent, stigma capitate. Capsule 0.7 × 0.7 mm, globose, greenish, glabrous, pedicellate, calyx and styles persistent. Seeds 0.20–0.22 mm long, trigonous, dark brown, verruculose.</p><p>Paratypes: — Brazil. Ceará: Chaval, Pedra da Carnaúba, 03º02’58”S, 41º15’17”W, 23 March 2024, fl., fr., M.I.B. Loiola 3046 (EAC).</p><p>Distribution and habitat: — Phyllanthus catingicola appears to be endemic to the northeastern region of Brazil, where it has so far been collected in the municipality of Chaval in the state of Ceará (Fig. 3). It has large populations (&gt; 50 individuals) in several rocky outcrops that are difficult to access and grows in moist, sandy and clayey soil on rocky outcrops in the Caatinga domain at around 17 meters of altitude. This species forms large populations associated with species of Cactaceae, Lythraceae and Poaceae .</p><p>Etymology: —The specific epithet “ catingicola ” refers to the Caatinga domain, in which the species is endemic. Furthermore, the new species is the only one from the section Loxopodium endemic to this domain, which is also rarely found in the Phyllanthus genus.</p><p>Phenology: —It was collected with flowers and fruits in March and April.</p><p>Preliminary conservation status: —Species classified as Data Deficient (DD), as it has an estimated area of occupancy of 10 km 2, and is only known from one location, not located in Conservation Units. In addition, the type of vegetation where it was found is common in the state of Ceará, indicating that with greater collection efforts its occurrence could be observed in other areas of the state.</p></div>	https://treatment.plazi.org/id/CD5DE5502516433395CDC17239B7DAA3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Santos, Diego;Sousa, Luana Mateus De;Torres, Alícia Marques;Silva, Marcos José Da;Loiola, Maria Iracema Bezerra	Santos, Diego, Sousa, Luana Mateus De, Torres, Alícia Marques, Silva, Marcos José Da, Loiola, Maria Iracema Bezerra (2025): A New Species of Phyllanthus (Phyllanthaceae) from Northeastern Brazil. Phytotaxa 694 (1): 93-100, DOI: 10.11646/phytotaxa.694.1.8, URL: https://doi.org/10.11646/phytotaxa.694.1.8
