identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
4A3B59B8A2C75E299C5876E1ED834C73.text	4A3B59B8A2C75E299C5876E1ED834C73.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mongolodiaptomus nakhonphanomensis Watiroyram & Monongdern & Koompoot 2025	<div><p>Mongolodiaptomus nakhonphanomensis sp. nov.</p><p>Figs 2, 3, 4, 5, 6, 7</p><p>Type locality.</p><p>A pond in the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.13426&amp;materialsCitation.latitude=17.500408" title="Search Plazi for locations around (long 104.13426/lat 17.500408)">Nong Hua Ngua Village</a> (Fig. 1 D), Na Khun Yai Sub-district, Na Wa District, Nakhon Phanom Province, north-eastern Thailand; coordinates: 17°30'01.47"N, 104°08'03.35"E, altitude: 153 m above sea level.</p><p>Material examined.</p><p>Holotype: one adult male dissected and mounted on one slide (THNHM -IV-21117); allotype: one adult female dissected and mounted on one slide (THNHM -IV-21118); paratypes: three adult males and three adult females dissected and mounted on one slide each (NPU 2025–01-04) . All specimens collected from the type locality on 30 July 2023 by Piyathida Monongdern.</p><p>Etymology.</p><p>The specific name nakhonphanomensis refers to Nakhon Phanom Province in north-eastern Thailand, where the new species was discovered. The Latin suffix “-ensis” denotes origin or place.</p><p>Description.</p><p>Adult male. Body length excluding caudal setae 2.1 mm (holotype) (other males: mean = 2.0, n = 3) (Fig. 2 A). Rostrum (Fig. 1 B) with two spiniform processes. Pedigers 4 and 5 incompletely fused laterally (Fig. 1 A). Lateral wings (Fig. 2 A) asymmetrical: left wing prolonged downwards, the right wing expanded distally in lateral direction; both tipped with thin spine. Urosome (Fig. 2 E) orientated obliquely posteriad towards right side. Genital somite short, asymmetrical, with small seta on outer distal margin on right side. Urosomites 2–3 each with long hairs on mid-ventral surface; former somite longer than latter. Urosomite 4 with right posterolateral margin dilated; dorsoposterior margin expanded, overlapping part of following somite. Anal somite symmetrical; anal operculum small, with posterior margin slightly concave. Caudal rami (Fig. 2 A, E) symmetrical; each ramus expanded distally, with setules along both inner and outer margins; with six setae (setae II – VII): setae II – VI plumose, seta VII bare. Right ramus without chitinous processes or ventral prominences.</p><p>Antennules asymmetrical, with the right antennule transformed and geniculated, exhibiting sexual dimorphism. Right side antennula (Fig. 2 C, D) 22 - segmented. Setal formula: 1 + a (I), 3 + a (II), 1 + a (III), 1 + a (IV), 1 + a (V), 1 (VI), 1 + a (VII), 1 + s (VIII), 2 + a (VIX), 1 + s (X), 1 + s (XI), 1 + a + s (XII), 1 + a + s (XIII), 2 + a + s (XIV), 2 + a + s (XV), 2 + a + s (XVI), 2 + s (XVII), 1 + s (XVIII), 2 + s (XIX), 3 + s (XX), 2 (XXI), 4 + a (XXII). Segments 13 to 20 enlarged and elongated; segments 18 and 19 geniculated; segment 20 (antepenultimate) with comb-like process (four teeth). Left side antennula (Fig. 4 A) 25 - segmented, reaching beyond the caudal setae. Setal formula as follows: 1 + a (I), 3 + a (II), 1 + a (III), 1 (IV), 1 + a (V), 1 (VI), 1 + a (VII), 1 + s (VIII), 2 + a (IX), 1 (X), 1 (XI), 1 + a + s (XII), 1 (XIII), 1 + a (XIV), 1 (XV), 1 + a (XVI), 1 (XVII), 1 (XVIII), 1 + a (XIX), 1 (XX), 1 (XXI), 2 (XXII), 2 (XXIII), 2 (XXIV), 4 + a (XXV).</p><p>Antenna (Fig. 4 B): coxa with one seta on inner side; basis with two setae on inner distal margin. Exp seven-segmented, with 1, 3, 1, 1, 1, 1, 1 setae laterally, three setae apically. Enp two-segmented; Enp - 1 with two setae at 1 / 2 length, Enp - 2 with nine setae along inner margin and seven setae apically, accompanied by short row of spinules on distal outer margin.</p><p>Mandible (Fig. 4 C): coxa with five bi-cuspidate teeth and one blunt tooth on gnathobase; with small dorsal seta on distolateral corner. Basis with four setae along inner margin. Enp two-segmented; Enp - 1 with four setae on distal inner corner. Enp - 2 with nine setae apically, accompanied by row of spinules along outer margin. Exp 4 - segmented, with 1, 1, 1, and 3 setae, respectively.</p><p>Maxillule (Fig. 4 D): praecoxal arthrite with ten robust setae apically and four fine setae on dorsal side. Coxal endite with three fine setae; coxal epipodite with eight robust setae. Basis with two endites, each armed with four setae; basal exite with one small seta. Enp reduced, represented by eight setae apically. Exp with seven apical setae.</p><p>Maxilla (Fig. 4 E): proximal and distal praecoxal endites with four and three setae, respectively. Two coxal endites with three setae each. Allobasis with three setae on basal endite. Enp two-segmented, each with three setae.</p><p>Maxilliped (Fig. 4 F): coxa with four endites, bearing 1, 2, 3 and 4 setae, respectively. Basis with three setae distally plus row of strong spinules along inner margin. Enp six-segmented, with 2, 3, 2, 2, 2 and 4 setae, respectively.</p><p>P 1 – P 4 (Fig. 5 A – D). P 1 with two-segmented Enp and three-segmented Exp; P 2 – P 4 with three-segmented Enp and Exp; Exp longer than Enp. Coxa on P 1 – P 4 with pinnate seta at distal inner corner. P 4 basis with small seta at distal outer margin. Exp - 1 P 1 – P 4 with one robust spine on outer margin and pinnate seta on inner margin. Exp - 2 as Exp - 1, but P 1 without spine on outer margin. Exp - 3 with three pinnate setae on inner margin and three pinnate setae apically on P 2 – P 4, but two on P 1 inner margin. Enp - 1 P 1 – P 4 with one pinnate seta on inner margin. P 1 Enp - 2 with three setae on inner margin, two setae apically, one pinnate seta on outer margin. P 2 – P 4 Enp - 2 with two pinnate setae on inner margin. Enp - 3 P 2 – P 4 with three setae on inner margin, two setae apically and two pinnate setae on outer margin.</p><p>P 5 (Figs 3, 5E, F) asymmetrical, with right leg markedly enlarged. Intercoxal plate with rounded distal margin. Right side of P 5: coxa with spiniform seta on posterior lobe, not reaching mid-length of basis. Basis approximately twice as long as wide, with narrow hyaline lamella along proximal inner margin and hyaline prominence at mid-distal region; with small seta at 3 / 4 length of outer margin. Exp three-segmented: Exp - 1 shorter than wide, with unproduced distal outer corner and semicircular hyaline prominence distally. Exp - 2 elliptical, with distal portion broader than proximal one, about 2.5 times as long as wide; with strong blunt spine at 1 / 2 length of outer margin and blunt process distally. Spine curved backwards, slightly shorter than half segment length. Exp - 3 as sickle-shaped claw, about 1.5 times as long as Exp - 2; with inner margin serrated. Enp 1 - segmented, conical, gradually tapering distally, reaching the mid-length of Exp - 2; with spinulated tip. Left side of P 5: coxa with thin seta at distal inner margin, reaching beyond mid-half of basis. Basis with narrow hyaline lamella along distal inner margin; with short, thin posterolateral seta on outer margin. Exp three-segmented (Fig. 5 E, F): Exp - 1 longer than wide, gradually tapering in posterior end; with a hair field at distal half of inner margin. Exp - 2 oval, smaller than Exp - 1; with inner robust seta, accompanied by spinular field along inner margin. Exp - 3 reduced to a short, bare segment with rounded tip. Enp one-segmented, conical, gradually tapering distally; reaching end of Exp - 1, with spinulate tip.</p><p>Adult female. Body length excluding caudal setae 2.5 mm (allotype) (other females: mean = 2.3, n = 3) (Fig. 6 A). Antennules symmetrical, with setal formula identical to that of the left antennule of male. Antenna and mouth appendages as in male. Cephalosome and pediger 1, as well as pedigers 4 and 5, completely fused. Pediger 5 (Figs 6 A, 7 B) with asymmetrical posterolateral wings: right wing short, rounded, with strong large spine dorsally, tiny spine on posterior margin; left wing longer, subtriangular, with tiny spine dorsally, larger spine posteriorly. Urosome (Fig. 7 A) three-segmented; genital double-somite longer than two subsequent urosomites and caudal rami combined. Genital double-somite (Fig. 7 A) asymmetrical; left side broadly expanded laterally at the proximal part, with strong spine; right side with proximal part slightly expanded laterally. Both spines articulated, inserted on somite prominence. Urosomite 2 symmetrical, shorter than wide, partly covered by posterior end of genital double-somite. Anal somite and caudal rami as in male (Fig. 7 A).</p><p>P 5 (Figs 6 B, C, 7 C, D) asymmetrical. Coxa with a stout spine on a prominence at the distolateral corner in dorsal view; coxal spine reaching proximal margin of Exp - 1. Basis with thin seta at 1 / 2 length of outer margin. Exp three-segmented, Enp two-segmented. Exp - 1 rectangular, about twice as long as wide. Exp - 2 triangular: right side shorter and stouter than left side (Fig. 6 C 1 – 2); inner margin with row of strong spinules; two longitudinal ridges ventrally; distal lateral seta strongly modified into short, stout spine (Fig. 6 D). Exp - 3 reduced to short, stout spine and long, thin seta. Enp cylindrical, extending beyond half-length of Exp - 1. Enp - 1 shorter than Enp - 2; Enp - 2 with circular row of spinules at narrowed apical end.</p><p>Variations.</p><p>In addition to body size differences, segment 20 of the right antennule in males have a comb-like process with two to five teeth (Fig. 2 D). The female P 5 Enp varies in length, extending from the middle to the distal end of Exp - 1. The size of the hyaline lamella on the right male P 5 basis also varies, appearing in long and narrow on the inner margin and rounded (Fig. 5 E) in the proximal-distal region.</p><p>Ecology and distribution.</p><p>At present, the new species has been recorded in seven samples, all collected from the Nawa District, Nakhon Phanom Province. It occurs in both temporary and permanent waterbodies. Physical and chemical parameters during sampling were: pH 6.2–7.01, temperature 27.2–31.2 ° C, conductivity 14.73–108.6 µS / cm, total dissolved solids (TDS) 7.34–52.5 mg / l. The co-occurring copepod species include Dentodiaptomus javanus (Grochmalicki, 1915), Eodiaptomus draconisignivomi Brehm, 1952, E. phuphanensis Sanoamuang, 2001, Heliodiaptomus elegans Kiefer, 1935, Mongolodiaptomus dumonti Sanoamuang, 2001, M. malaindosinensis (Lai &amp; Fernando, 1978) and Neodiaptomus yangtsekiangensis Mashiko, 1951 .</p></div>	https://treatment.plazi.org/id/4A3B59B8A2C75E299C5876E1ED834C73	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Watiroyram, Santi;Monongdern, Piyathida;Koompoot, Kamonwan	Watiroyram, Santi, Monongdern, Piyathida, Koompoot, Kamonwan (2025): Mongolodiaptomus nakhonphanomensis sp. nov. (Copepoda, Calanoida, Diaptomidae) from north-eastern Thailand, with a note on M. pectinidactylus (Shen & Tai, 1964). Zoosystematics and Evolution 101 (4): 1965-1983, DOI: 10.3897/zse.101.161679
308DB2FB497D5BE1B2D72262FD50B083.text	308DB2FB497D5BE1B2D72262FD50B083.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mongolodiaptomus pectinidactylus (Shen & Tai 1964)	<div><p>Mongolodiaptomus pectinidactylus (Shen &amp; Tai, 1964)</p><p>Figs 8, 9</p><p>Sampling sites.</p><p>Samples were collected from four ponds in the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.084236&amp;materialsCitation.latitude=17.585197" title="Search Plazi for locations around (long 104.084236/lat 17.585197)">Lao Pattana Village</a>, Lao Pattana Sub-district, Na Wa District, Nakhon Phanom Province, north-eastern Thailand. The coordinates and elevations are as follows: 1) 17°35'4.59"N, 104°05'9.56"E, 151 m above sea level ; 2) 17°35'0.15"N, 104°06'0.25"E, 152 m above sea level; 3) 17°34'6.05"N, 104°06'4.47"E, 147 m above sea level; 4) 17°35'6.71"N, 104°05'3.24"E, 155 m above sea level .</p><p>Material examined.</p><p>Four adult males, each dissected and mounted on separate slide (NPU 2025–05–08); three adult females prepared for scanning electron microscopy (SEM) imaging. All specimens were collected on 4 August 2023 by Piyathida Monongdern.</p><p>Adult male. Urosomites 2–3 with hairs on ventral side. Caudal rami symmetrical, without chitinous prominence on right ramus ventrally. P 5 (Figs 8 A, 9 A, B) asymmetrical, intercoxal plate with rounded distal margin. Right side of P 5: coxa with spiniform seta inserted on distal margin dorsally, not reaching mid-length of basis. Basis about twice as long as wide, with narrow hyaline lamella along proximal inner margin and hyaline prominence at middle of segment dorsally; row of tiny spinules on ventral surface (Fig. 8 B); small seta at 2 / 3 of segment length on ventral side. Exp three-segmented: Exp - 1 shorter than wide, with unproduced distal outer corner and semicircular hyaline prominence distally. Exp - 2 (Fig. 8 C, D) elongated, with broader middle part, about 2.5 times as long as wide; with blunt-tipped spine at 1 / 2 length of outer margin and blunt process distally; spine straight, shorter than half of segment length. Exp - 3 modified as sickle-shaped end claw with serrated inner about 1.5 times as long as Exp - 2. Enp one-segmented, conical, not reaching the mid-length of Exp - 2; with spinulated tip. Left side of P 5: coxa with thin seta at distal inner margin, reaching to mid-half of basis. Basis with narrow hyaline lamella along distal inner margin; with short, thin posterolateral seta on outer margin. Exp three-segmented: Exp - 1 longer than wide, gradually tapering in posterior end; with hair field at distal half of inner margin; Exp - 2 oval, smaller than Exp - 1; with inner robust seta, accompanied by spinular field along inner margin. Exp - 3 reduced to short, bare segment with rounded tip. Enp one-segmented, conical, gradually tapering distally; not reaching end of Exp - 1, with spinulate tip.</p><p>Adult female. Pediger 5 with almost symmetrical posterolateral wings; dorsal spine smaller than posterior spine on both wings. Genital double-somite asymmetrical; left side broadly expanded laterally at proximal part and with larger spine. Both spines on genital double-segment articulated, inserted on small somite prominence and directed down-laterally. Right margin concave, left margin convex. Anal somite and caudal rami as in male. P 5 (Fig. 8 E) symmetrical. Coxa with stout spine on prominence at distolateral corner in dorsal view; coxal spine reaching proximal margin of Exp - 1. Basis with thin seta on distolateral margin in dorsal view, reaching beyond middle of Exp - 1. Exp three-segmented and Enp two-segmented. Exp - 1 enlarged, rectangular, more than 2.0 times as long as wide. Exp - 2 (Fig. 8 F, G) subtriangular, symmetrical, both margins with row of strong spinules; two longitudinal ridges visible in ventral view; distal lateral seta modified into short, stout spine. Exp - 3 reduced to small spine, fused to base of Exp - 2 accompanied by one short and one long, thin seta. Enp subconical, extending beyond half-length of Exp - 1. Enp - 1 shorter than Enp - 2; Enp - 2 with circular row of spinules at narrowed apical end.</p><p>Variations.</p><p>The male right P 5 basis has one or two rows of tiny spinules amongst individuals in the population (Figs 8 B, 9 B).</p><p>Distribution.</p><p>Mongolodiaptomus pectinidactylus has been sporadically recorded across China, Vietnam and Thailand (Sanoamuang and Dabseepai 2021). In Thailand, it was previously reported from only two localities in the Roi Et and Ubon Ratchathani Provinces (Sanoamuang 2002). More recently, additional records have been reported from the Sakon Nakhon and Nakhon Phanom Provinces (Koompoot and Watiroyram 2025). The species has been found in both temporary and permanent waterbodies (Sanoamuang and Dabseepai 2021), similar to the observations made in the present study.</p><p>Differential diagnosis.</p><p>Compared to the generic diagnoses by Kiefer (1938) and Ranga Reddy et al. (2000), the new species is assigned to the genus Mongolodiaptomus, based on the following characters: (1) urosomites 2–3 of the male with hairs on the ventral surface; (2) the presence of a spiniform seta on the male P 5 coxa; (3) the lateral spine on the right male P 5 Exp - 2 inserted at the 1 / 2 length of outer margin, accompanied by two accessory spines. The male P 5 intercoxal sclerite is rounded and unproduced, differing from the condition described by Kiefer (1938), but similar to that observed in M. calcarus, M. dumonti, M. gladiolus, M. mephistopheles and M. pectinidactylus . Additionally, the male lacks a chitinous prominence on the ventral side of the right caudal ramus, which also contrasts with Kiefer’s (1938) description.</p><p>The new species is distinguished from its congeners by the distinctive morphology of the right male P 5 Exp - 2 and the right caudal ramus, as well as by the absence of a chitinous prominence on the ventral side of the right caudal ramus. This combination of characters makes the new species most similar to M. pectinidactylus, a member of the M. mephistopheles (Brehm, 1933) species group sensu Sanoamuang and Watiroyram (2018) and Sanoamuang and Koompoot (2024). Based on traditional taxonomy, particularly the morphology of pediger 5 in females, the urosome and the P 5 in both sexes, the species can be distinguished from M. pectinidactylus as follows: the female of the new species has an asymmetrical lateral wing on pediger 5, with the right dorsal spine and the left posterior spine being larger than those of M. pectinidactylus . Additionally, the female P 5 is asymmetrical, featuring a longer left Exp - 2 and a stouter right Exp - 2, whereas the P 5 is symmetrical in M. pectinidactylus . The seta on the female P 5 Exp - 2 is also distinctly shorter and stouter compared to that of M. pectinidactylus . The male of the new species lacks a row of spinules on the frontal surface of the P 5 basis, whereas this feature is present in M. pectinidactylus . In addition, the male P 5 Exp - 2 of the new species is slightly more distally expanded than that of M. pectinidactylus and the lateral spine is bent outwards, in contrast to the straight spine in M. pectinidactylus (Table 2).</p></div>	https://treatment.plazi.org/id/308DB2FB497D5BE1B2D72262FD50B083	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Watiroyram, Santi;Monongdern, Piyathida;Koompoot, Kamonwan	Watiroyram, Santi, Monongdern, Piyathida, Koompoot, Kamonwan (2025): Mongolodiaptomus nakhonphanomensis sp. nov. (Copepoda, Calanoida, Diaptomidae) from north-eastern Thailand, with a note on M. pectinidactylus (Shen & Tai, 1964). Zoosystematics and Evolution 101 (4): 1965-1983, DOI: 10.3897/zse.101.161679
