Thysanoplisna signata (Walker, 1854)

(Figs 5–12, 42, 43, 57, 58)

Lithosia signata Walker, 1854, List of the specimens of lepidopterous insects in the collection of the British Museum, 2: 495 (Type locality: “ Shanghai, North China ”).

Type material examined. Holotype (by monotypy) (Fig. 5): ♀, “N. China | (Fortune) | 55–87” / “3. Lithosia signata .” / green ring “ Type ” label / QR-code label with unique ID “ NHMUK 010401739 ” (NHMUK).

Additional material examined. CHINA: 1 ♀, Shanghai ( Fortune), 54–8, NHMUK010606147 , gen. prep. No.: NHMUK010313548 (NHMUK); 18 ♂♂, 13 ♀♀, Jiangxi /Fujian border, Wuy Shan, 50 km SE of Yingtan, 27°56'N 117°25'E, 1600m, iv.2002, V . Sinyaev & local collectors leg., gen. prep. No.: MWM 35.990, 35.992 (♀♀) (MWM/ ZSM); 2 ♂♂, same data as previous but v.2002, gen. prep. No.: ZSM Arct. 2021-447 (MWM/ ZSM) ; 1 ♂, 2 ♀♀, Jiangxi Prov., Xipaihe Village, 1500m, 27°54'N 117°20'E, vi.2003, V . Sinyaev & his team leg., gen. prep. Nos.: MWM 35.985 (♂), MWM 35.986 (♀) (MWM/ ZSM); 1 ♂, 1 ♀, same data as previous but vii.2003 (MWM/ ZSM); 5 ♂♂, 4 ♀♀, Hainan Dao, He Ling Mts and Wuzhi Shan, 1000–1800m, early ii–early v.2000, light trap, J.L. Li leg., coll. Maier, gen. prep. Nos.: MWM 35.987, 35.988 (♂♂), 36.033 (♀) (MWM/ ZSM) ; 17 ♂♂, 7 ♀♀, [Fujian Prov.] Kuatun (2300m), 27°40'N 117°40'E, J. Klapperich [leg.]: 2.v.1938 (♂, gen. prep. No.: MWM 35.979), 3.v.1938 (♀), 16.v.1938 (♂, 2 ♀♀, gen. prep. No.: MWM 35.980 (♀)), 17.v.1938 (8 ♂♂, 2 ♀♀), 19.v.1938 (2 ♂♂), 21.v.1938 (2 ♂♂, 1 ♀), 22.v.1938 (♂, ♀), 27.v.1938 (♀), 16.vii.1938 (♀), 1.viii.1938 (♂), 13.viii.1938 (♂), and 18.viii.1938 (♀, gen. prep. No.: MWM 35.983) (MWM/ ZSM) ; 1 ♀, same data as previous, 20.v.1946, unique ID: NHMUK010606153, gen. prep. No.: NHMUK010313552 (NHMUK); 1 ♂, 1 ♀, [Fujian Prov.] Kuatun, Prov. Fukien, 2.ix.1938, [H.] Höne [leg.], gen. prep. Nos.: MWM 35.981 (♀), MWM 35.982 (♂) (MWM/ ZSM) .

Notes. The species was described from a single female, which was lettered by Walker (1854) in the original description as ‘a’, and this specimen should be treated as holotype by monotypy. Hampson’s (1900) subsequent citation of the material examined as “N. China (Fortune), 2♀, type” should be regarded as the author studied two specimens from North China collected by Fortune, including the type one (A. Zilli, pers. comm.).

Diagnosis. The forewing length is 16.5–18.0 mm in males and 17.0– 19.5 mm in females. Thysanoplisna signata is most similar to Th. brevimacula and the detailed comparison is provided above in the diagnosis of the latter.

Distribution. East and South China (Fukien, Jiangxi, Guangdong, and Hainan Provinces, and Shanghai) (Walker 1854; Daniel 1954 (partim., as ‘ Lithosia signata ssp. directa ’); Kishida 2011).