Bonaspeiini Zahniser & Dietrich, 2013: 50 .
Diagnosis (modified from Zahniser & Dietrich (2013). Medium sized leafhoppers; forewings often submacropterous (Fig. 7I) to brachypterous (Fig. 11M) and in the former, hind wings strap-like (Figs 7J, 17M) and in the latter hind wings reduced in size (Fig. 11N) or absent. Connective wider than long ( Colistra Fig. 7H, Proekoides Fig. 16K–O, Proekes Fig. 23H) with arms strongly divergent with stem often very short (Fig. 7H) or absent; equidistant in Xhoreus gen. n. Fig. 20K; rarely longer than wide (e.g., Hadroca Theron, 1974). Male pygofer usually deeply incised (dorsal bridge very short e.g., Colistra, Fig. 7A–B, Bonaspeia Linnavuori, 1961, Caffrolix Linnavuori, 1961, Curvostylus Davies, 1987) with segment 10 large (except Xhoreus gen. n.). Ovipositor sometimes distinctly protruding far beyond pygofer apex, e.g., Proekes (Figs 22F–J, 24J, 27C, D, G,).
Remarks. The tribe Bonaspeiini, like many other Deltocephalinae tribes, is poorly defined morphologically, there being no unique characters found in all genera. The characters noted in the above diagnosis are a reduced set compared to that given by Zahniser & Dietrich (2013). Of the characters used by these authors to define the tribe the color range of ivory, greenish, or brownish is considered too wide ranging to be diagnostic. In addition, the absence of transverse carinae on the fore margin of the head (a situation found in most Deltocephalinae) was only mentioned by Zahniser & Dietrich (2013) because it is a diagnostic character for Selenocephalini, which was paired with some Bonaspeiini in molecular studies (pers. com. J. Zahniser). A group of genera in Bonaspeiini, (including those studied here) have reduced wings but this could be due to convergent evolution and habitat but the strap-like hind wings found in all submacropterous forms is unusual. Based on the above findings, the validity of the tribe awaits further studies on the many other similar new genera and species which have been found in the Savanna and Fynbos Biomes.