Cardiodactylus pelagus Otte, 2007a

(Figs. 4, 6F, 10E, 12H, 13P, 15F, 49)

Cardiodactylus pelagus Otte, 2007a: 348; 2007: 30 (confirmation of depository).

Type material. Male holotype: Malaysia. Sarawak, Nanga Pelagus near Kapit, 180–585 m, 7–14.VIII.1958, Maa (BPBM) (not examined) .

Distribution. Malaysia, western part of Borneo.

Material examined. Malaysia. Sarawak State, foot of Mt. Dulit, junction of rivers Tinjar & Lejok, 10–24.IX.1932, Oxford Univ. Exp., B. M. Hobby & A. W. Moore, B.M.1933-254: 1♂, fish trap 1, identified Cardiodactylus gaimardi by L. Chopard (MNHN-EO-ENSIF3511) ; 1♀, #28, traps 1, identified Cardiodactylus gaimardi by L. Chopard (BMNH) ; 1♂, traps, identified Cardiodactylus gaimardi by L. Chopard (BMNH) . East Malaysia, 4 th Division, Baram District, Gunong [Gunung = Mount] Mulu National Park, 4°03’N 114°56’E, 150 m, 12–18.V.1978, 1♀, malaise in mixed dipterocarp forest, N. M. Collins (BMNH.1978-11) . Sarawak, Mulu National Park not far from borders with Brunei and Indonesia, 100–300 m, primary forest, on leaf of bush, at night, 24–27.III.2012, A. Gorochov, M. Berezin, E. Tkatsheva, I. Kamskov, 1♂, 2♀ (ZIN) . Borneo I., environs of Kuching City, Kubah National park on Matang Mt., 200–500 m, primary forest, on leaf of tree not far from soil, at night, 10–17.III.2012, A. Gorochov, M. Berezin, E. Tkatsheva, I. Kamskov, 1♀ (ZIN) .

Diagnosis. Species of small size, coloration contrasted, dark brown to black, with yellowish patterns. Male genitalia close to that of C. singapura with parallel carinated dorsal ridges and apex of pseudepiphallic sclerite not rounded. Species close to C. variegatus n. sp. but differing by coloration of pronotum (dorsal disk laterally yellow but median area mostly dark brown; lateral lobe yellow ventrally), and by coloration of FIII orange brown, less contrasted.

Redescription. In addition to the characters given by Otte (2007): size small for the species group, coloration dark brown to black with contrasted yellow brown patterns (Fig. 49A–D). Head dorsum yellow brown with 4 dark brown bands including 2 complete lateral ones and 2 thin median punctuated bands; area posterior to eyes yellow brown with 2 short dark brown lines (Fig. 6F). Pronotum: Dorsal disk yellow brown laterally, median area mottled with dark brown and with black spots. Lateral lobes dark brown dorsally, ventral half yellow. Legs yellow brown, femora mottled with dark brown, tibiae with dark brown rings. FIII orange brown, almost homogeneous, knees slightly darker than rest of femora. Tarsomeres 1–2 yellow brown, their apex dark brown. Hind knees black or dark brown. Cerci yellow brown, base faintly mottled with dark brown, apex with dark brown rings.

Male: FW coloration (Fig. 10E) mostly dark brown to black (cells and veins), with yellow to whitish areas including bases of anal veins and CuA, harp veins, oval region posterior to mirror and transverse veins in apical field. Bases of chords orange brown. M/R/Sc area and cells dark orange brown. Projections of Sc and more ventral veins of lateral field yellow. FW venation (Fig. 10E): 1A slightly bisinuated. CuP absent. Harp with 2 w-shaped veins. Mirror area: mirror (d1) oval, separated in two parts near mid-length; d2 as wide as mirror. Apical field with 3 cell alignments posterior to mirror (n = 3). Lateral field with 4–7 projections of Sc (m = 5; n = 3) and 3 more ventral veins (n = 3).

Male genitalia (Fig. 49E–F): Pseudepiphallus short, narrowed twice, at mid-length and preapically. Lateral ridges parallel, thick, wider anteriorly, their dorsal edges carinated. Pseudepiphallic sclerite with short triangular anterior expansions. Posterior pseudepiphallic apex trapezoidal, with a small rounded translucent area. Rami parallel, with very short apical stems. Ectophallic arc complete, v-shaped. Ectophallic fold with strong lateral sclerites; apex trilobate, membranous, median lobe elongate. Endophallic sclerite long, with a triangular posterior expansion and short lateral arms. Endophallic apodeme with a dorsal crest and wide lateral lamellas. Membrane of endophallic cavity finely plicate.

Female: FW coloration (Fig. 12H) mostly dark brown with contrasted yellow cells. Base of FWs yellow brown, including bases of anal veins, CuA and anterior half of M. CuA/M area yellow brown basally, then dark brown, with a yellowish spot at mid-length, extended on M/R area. R/Sc area and veins orange brown. Projections of Sc and more ventral veins yellow, the cells between them dark brown. FW venation (Fig. 12H): 10 (n = 2) strong longitudinal veins on dorsal field; lateral field with 7–8 (n = 2) longitudinal veins including 4–5 projections of Sc and 3–4 more ventral veins. Ovipositor of average length, apex with both dorsal and ventral edges slightly denticulate (Fig. 13P).

Female genitalia (Fig. 15F): Copulatory papilla triangular, with baso-lateral sclerites; apex rounded and slightly sclerotized, slightly folded ventrally.

Juvenile: unknown.

Remark. The female specimen photographed by Otte (2007a) probably does not belong to C. pelageu, but to C. borneoe n. sp., according to the pattern of coloration of the material examined here.