Gonaphodiellus bimaculosus (Schmidt, 1909)

(Fig. 6-12)

Aphodius bimaculosus Schmidt, 1909: 19 .

Aphodius (Gonaphodiellus) bimaculosus; Schmidt 1913: 134; Dellacasa 1988: 98.

Aphodius (Gonaphodiellus) xalapensis Galante et al. 2003: 304 (new synonymy).

Gonaphodiellus bimaculosus; Skelley et al. 2007: 4.

Type locality. Mexico.

Type repository. Naturhistoriska Riksmuseet, Stockholm, Sweden (type examined).

Redescription. Length 5.0-6.0 mm, oblong, moderately convex, shiny, glabrous. Brown-yellowish; head, pronotum disc and elytra brownish, elytra usually with a yellowish round spot on preapical declivity (Fig. 10), sometimes elytra entirely brownish (Fig. 9); legs dark brown; antennal club brownish. Head with epistome feebly convex on disc, subregularly and rather densely punctured; clypeus anteriorly subtruncate, obtusely round at sides, lateral margins finely bordered, edge very shortly sparsely bristled; genae obtusely round, very shortly ciliate, protruding from the eyes; frontal suture finely impressed; front rather sparsely regularly punctured. Pronotum transverse, moderately convex, subregularly punctured; punctation somewhat denser on sides; lateral margins almost parallelsided, finely bordered, edge very shortly sparsely bristled; hind angles widely truncate, inwardly sinuate; base bisinuate. Scutellum triangularly elongate, with few sparse and irregular punctures on basal half. Elytra slightly broadened posteriorly, with a small denticle at shoulder; striae fine, superficially punctured, faintly crenulate; interstriae entirely shiny, slightly convex, finely sparsely punctured. Hind tibiae upper spur as long as half of first tarsal segment; first tarsal segment as long as following four segments combined. Male: pronotum not narrowed anteriorly, relatively somewhat more sparsely punctured on disc; fore tibiae slender and more elongate, their spur stouter, downward bent and outward curved; metasternal plate longitudinally feebly excavate at middle; aedeagus Fig. 11-12. Female: pronotum slightly narrowed anteriorly, relatively somewhat more densely punctured on disc; fore tibiae stouter and shorter, their spur slender and almost straight; metasternal plate flat.

Material examined. EL SALVADOR: Monte Cristo, 23 Km N Matapan, m 2300, 08-10.V.1971, leg. Howden H. F. (12 exx., CMNO) . MEXICO: Hidalgo: 2.9 Km S Molango, 20.VII.1988. leg. Turnbow R. (2 exx., RHTC); 1 mi. S Santa Monica, 6100’, 12.VII.1973, leg. Newton A. (4 exx. CMNO, DCGI) ; 7 Km SW Tenango de Doria, VII.1971, leg. Newton A. (2 exx., CMNO) ; 6.4 mi. S Tenango de Doria, m 3000, 24-28.VII.1969, leg. Peck S. & J., cloud forest, oak (5 exx., CMNO) ; Tlanchinol env., m 1500, 10-24.VIII.2001, leg. Kabourek V. (2 exx., DCGI) ; 6.4 mi. S Tianguistengo, 6800’, 12.VII.1973. leg A. Newton (1 ex., DCGI) ; Zacualtipan, 27.V.1979, leg. Zunino M. (2 exx., DCGI) ; Jalisco: 10 mi W Atenquique, 6700’ 18.IX.1971, leg. Newton A. (5 exx. CMNO, DCGI) ; Oaxaca: 13 mi. N Nochixtlan, 7000’, 27.VI.1968, leg. Hardy A. R., Espinosa L. & Abrayaya J. P., Hardy Central American Expedition, (1 ex., USNM) ; Km 40 Camino Teotitlan, Hualta de Jiménez, m 2400, 09.XI.1968, leg. Reyes P. & Cabrera M, bosque nebular (2 exx., DCGI, FSCA) ; Km 133 carr. Tuxtepec to Oaxaca-Atepec, Llano de las Flores, 17°26’37.2"N – 96°30’13.2"W, m 2600, 14.IX.2003, leg. Martínez I. & Suarez T. (1 ex., DCGI) ; 30 mi. Valle Nacional, 17.VIII.1973, leg. Newton A. (2 exx., CNCI) ; Puebla: 5 mi. W Huauchimango, 5900’, 07.VII.1970, leg. Newton A. (3 exx., CNCI) ; 6 mi W Teziutlan, 4-6.VIII.1960, leg. Howden H. F. (1 ex., DCGI) ; Queretaro: 25 mi. E Landa de Matamoros, 5000’, 14-17.VII.1969, leg. Peck S. & J., liquidambar forest (2 exx., CNCI, 1 ex., DCGI) ; Veracruz: Mpio. Coscomatepec, Cuiyachapa, 06.II.1999, leg. Alvarado M. (5 exx., DCGI) ; Cuiyachapa, carr. á Potrero Nuevo, 15.IX.1994, leg. Arellano L. & Zunino M. (3 exx., DCGI) ; Mpio. Coscomatepec, carr. Cuiyachapa, Pueblo Nuevo, 08.XI.1999, leg. Dellacasa M. (1 ex., DCGI) ; El Duraznal, m 1600, 15.VII.1994, leg. Arellano L. & Sanchez R. (1 ex., DCGI) ; Mpio. Xico, Oxtlapa, m 2100, 16.XI.1999, leg. Dellacasa M. (1 ex., DCGI) ; carr. Xico-Tonalaco, 19°26’21"N – 97°06’10"W, m 2300, 09.XII.1999 - 09.II.2000, leg. Montes de Oca E. & Santiago Q. (4 exx., DCGI) ; idem, 06.VII.- 11.VIII.2000, leg. Montes de Oca E. & Santiago Q. (13 exx., DCGI) .

Distribution. El Salvador, Mexico (Hidalgo, Jalisco, Oaxaca, Puebla, Queretaro, Veracruz).

Bionomics. Probably all year round, coprophagous species found in several kinds of dung.