Gonaphodioides columbicus (Harold, 1880) new combination

(Fig. 78-83)

Aphodius columbicus Harold, 1880: 36; Blackwelder 1944: 212.

Aphodius (Gonaphodiellu s) columbicus; Schmidt 1913: 134; 1922: 121; Dellacasa 1988: 314.

Gonaphodiellus (s. l.) columbicus; Skelley et al. 2007: 4.

Type locality. Bogotá [Colombia].

Type repository. Muséum national d’Histoire naturelle, Paris, France (lectotype herein designated).

Redescription. Length 4.0-5.0 mm, elongate, moderately convex, shiny, glabrous. Chestnut-brown; clypeal margin, sides of pronotum, apex of elytra shadowy reddish; legs pale reddish; antennal club testaceous. Head with epistome feebly gibbous, finely regularly and rather sparsely punctured throughout; clypeus slightly sinuate at middle, obtusely angulate at sides, faintly bordered; edge distinct and somewhat upturned anteriorly only, glabrous; genae subangulose, not ciliate, protruding from the eyes; frontal suture finely impressed; front finely and sparsely punctured. Pronotum transverse, moderately convex, finely, regularly and rather sparsely punctured; punctation somewhat denser anteriorly; sides feebly curved, very finely bordered, edge glabrous; hind angles widely truncate, truncature slightly inwardly sinuate; base distinctly bisinuate. Scutellum ogival elongate, somewhat depressed and distinctly microreticulate basally. Elytra convex, elongate with humeral denticle very small; striae fine, superficially punctured, not crenulate; interstriae distinctly convex on disc and nearly imperceptibly punctured laterally, nearly flattened subapically. Hind tibiae upper spur as long as half of first tarsal segment; first tarsal segment longer than following three segments combined. Male: pronotum relatively more transverse and less convex, parallel-sided with anterior discal punctation finer; elytra more elongate, nearly parallel-sided; fore tibiae more slender and more elongate, their spur downward bent; aedeagus Fig. 82-83. Female: pronotum relatively less transverse and more convex, somewhat narrowed anteriorly, uniformly punctured throughout; elytra shorter and somewhat broadened backward; fore tibiae more stout and shorter, their spur straight.

Type material. Lectotype, male, herein designated (MNHN), bearing the following labels: 1) white, handwritten: Bogotá; 2) white, printed: ex Musaeo/E. Steinheil; 3) red, printed: Aphodius / columbicus / Harold, 1880; 4) red, printed: Lectotypus/ M. & G. Dellacasa des. 2001 .

Paralectotypes: 3 males and 4 females from Bogotá (paralectotypes, MNHN); 3 females from La Luzera [Colombia], ex Musaeo / E. Steinheil (paralectotypes, MNHN); 1 male and 1 female from Viani, ex Musaeo / Harold (paralectotypes,, MNHN) ; 1 female from Noescen. [Colombia?] (paralectotype, MNHN); 1 female from Colombia, ex Musaeo / E. Harold (paralectotype, MNHN) .

Material examined. COLOMBIA: Agnas Largas près Bogotá, 1877, leg. Thieme O. (1 ex., MNHN) ; Bogotá (1 ex., DCGI); Bogotá, 1877, leg. Thieme O. (3 exx., MNHN); Oberhalb Bogotá, 1000’, 1877, leg. Thieme O. (3 exx., MNHN) ; Papayan, 1899, leg. Abbé Gaujon (1 ex., MNHN) ; Santander, R. Natural Cachalú, m 2150-2250, III.1999, leg. Gonzales F. (2 exx., DCGI) . VENEZUELA: Merida, Mucuy Fish Hatchery, 7 Km E Tabay, 6600’, 10-13.II.1978, leg. Heppner J. B., black light (3 exx., DCGI) ; Merida, La Mucuy Estacion, 10-15.VIII.1976, leg. Bordon C. (4 exx., DCGI) ; Merida, La Mucuy, m 2300, 10.IX.1965, leg. Bordon C. (1 ex., DCGI) ; Trujillo, Paramo del Guaramacal, m 3000, 24.V.1980, leg. Bordon C. (5 exx., DCGI) .

Distribution. Colombia, Venezuela.

Bionomics. Poorly known; the specimens examined were collected in spring, summer and early autumn.