Bombus (Pyrobombus) bryorum Richards, 1930 stat. rev.

Figs 1–2, 10‒16

[ Bombus hypnorum] var. [not subsp.] bryorum Richards 1930: 650, but not infrasubspecific after Tkalců (1974: 328) (ICZN,1999: Article 45.6.4.1). Holotype by original designation: ♀ (not ♂) Kashmir, India (NHMUK). Examined.

B. [ombus] fletcheri Richards, 1934: 90 . Holotype by original designation: ♀ Gulmarg, Kashmir, India (NHMUK). Examined.

[ Bombus (Pyrobombus) hypnorum (Linnaeus, 1758) – Williams 1991: 70 — Williams et al. 2010: 127, misidentification.]

Bombus (Pyrobombus) bryorum Richards – Williams 2022a: 97.

Diagnosis

Female

Distinguished by the combination: labral lamella broad and shallowly rounded, almost rectangular; clypeus in the central area with scattered widely-spaced small punctures and the very large punctures spaced by more than 3× their own diameter; ocello-ocular area along the inner eye margin with only one row of larger medium punctures; thoracic dorsum with the hair brown in the centre between the wing bases without black hair.

Male

Distinguished by the combination: hair variable, from the thorax and T1‒4 yellow (western Himalaya), to thorax and metasomal T1‒2 brown and T3‒4 black (eastern Himalaya).

Material sequenced or examined

INDIA • 2 ♀♀ (queens); Kashmir, Gulmarg; 34.0343° N, 74.3678° E; alt. 3100 m; 1986; P. Williams leg.; PW • 2 ♀♀ (1 queen, 1 worker), 1 ♂; Kashmir, Gulmarg; 34.0343° N, 74.3678° E; alt. 3100 m; Aug. 1985; P. Williams leg.; PW • 1 ♀ (worker), 2 ♂♂; Kashmir, Gulmarg; 34.0343° N, 74.3678° E; alt. 2700‒3200 m; 16 Aug. 1980; P. Williams leg.; PW .

NEPAL • 1 ♀ (worker); Karnali, Maharigaon; 29.337° N, 82.387° E; alt. 3250 m; 8 Jul. 1999; M. Hartmann leg.; BOLD-1550B09-N539; NME • 1 ♂; Mahakali, Charmaliya Khola; 29.746° N, 80.801° E; alt. 3100 m; 10 Jun. 2005; J. Weipert leg.; BOLD-1550B12-N548; NME .

BHUTAN • 1 ♀ (queen); Thimpu, Phajoding Gompa; 27.479° N, 89.595° E; alt. 2850 m; 25 Apr. 2018; J. Smit leg.; sequence-RMNH-1092452; RMNH • 1 ♀ (queen); Wangdue, Phodrang, Dangchhu; 27.546° N, 90.203° E; alt 3380 m; 5 May 2018; J. Smit leg.; RMNH .

Distribution

Himalaya (from Pakistan to Arunachal Pradesh), at elevations of 2286‒3900 m (Williams 1991; Williams et al. 2010).