Cosmetra Diakonoff, 1977
Type species: Cosmetra anthophaga Diakonoff, 1977 .
Cosmetra includes three species restricted to the Afrotropical region: C. anthophaga Diakonoff, 1977 (type species) from Réunion Island, C. rythmosema Diakonoff, 1992 from Madagascar, and C. neka Razowski & Brown, 2009 from South Africa. We provisionally assign to the genus two new species described below and transfer Sycacantha nereidopa (Meyrick, 1937) to Cosmetra, new combination .
Based on recent work focused on the Afrotropical Region, Razowski et al. (2010) synonymized Polychrosis hendrickxi Ghesquière, 1940 (TL: Belgian Congo [Zaire]) with Eucosma nereidopa Meyrick, 1927 (TL: Kenya Colony [Kenya]) and its synonym Eucosma phylloscia (Meyrick, 1937) (TL: Uganda), and transferred them to Sycacantha (Olethreutini) . They provided illustrations of the holotype adult and male genitalia of P. hendrickxi (Razowski et al. 2010: figs. 37, 81). Illustrations of the holotypes of S. nereidopa and S. phylloscia can be found in Clarke (1958: 376). Below we describe two new species from Kenya reared from native fruit that are morphologically similar to nereidopa, the latter of which is a well-known pest of coffee in Africa (e.g., Coffee Research Services 1963; Evans 1968, 1970; Waller et al. 2007; Hill 2008). There is little question that these African species are congeneric, and based on the female genitalia they do not belong in Sycancatha or in Olethreutini . We provisionally describe our two new species in Cosmetra Diakonoff, 1977 and provisionally transfer nereidopa to that genus. However, it is likely that the three require a new genus.