Galeopsomyia crotonae Hansson sp.nov.

(Figs 358, 359, 1069)

Diagnosis (female). Antenna with solid clava (Fig. 1069); genal carina absent; mesoscutellum (Fig. 358) with weak reticulation on median part, lateral parts with strong reticulation, meshes elongate, submedian grooves ±parallel, with a weak median groove in anterior half; dorsellum without median carina; propodeum (Fig. 358) short, with strong reticulation without irregular carinae, callus with two setae; hind margin of hind coxa with a short longitudinal carina close to base; gaster (Fig. 358) 2.6× as long as wide, medio-basal Gt1 with a carina shaped like an inverted U (as in Fig. 26), tergites with strong reticulation.

Female holotype: length of body 2.0mm (paratypes 1.4–1.9mm).

Scape yellowish-brown, pedicel brown, flagellum dark brown. Face below level of toruli golden-green, clypeus dark brown, frons and vertex metallic bluish-green. Mesoscutum and mesoscutellum metallic bluish-green, propodeum golden-green. Legs with fore and hind coxae golden-green, mid coxa black; trochanters yellowish-brown; femora dark brown with apex yellowish-brown; tibiae yellowish-brown; T1–3 yellowish-brown, T4 pale brown. Petiole black. Gaster metallic bluish-green; gonoplac black.

Antenna with solid clava. Face below level of toruli with weak reticulation, clypeus smooth, frons with strong reticulation, antennal scrobes with weak reticulation; without genal carina. Vertex with strong reticulation.

Mesoscutum with strong reticulation, meshes isodiametric on sidelobes, elongate on midlobe; midlobe without median groove, with six scattered adnotaular setae. Mesoscutellum with weak reticulation on median part, lateral parts with strong reticulation, meshes elongate; submedian grooves straight and parallel; with ten scattered setae over lateral parts. Dorsellum with strong reticulation, without median carina. Propodeum with a triangular median carina; with strong reticulation; callus with two setae. Fore and hind coxae with strong reticulation, mid coxa with weak reticulation; hind margin of hind coxa with a short longitudinal carina close to base. Fore wing with two setae on dorsal part in basal half of submarginal vein; speculum closed; costal setal row broken.

Petiole very short, just a narrow band. Gaster elongate; medio-basal Gt 1 with a carina shaped like an inverted U; tergites with strong reticulation.

Relative measurements: head length, dorsal view 23; head length, frontal view 35; POL 9; OOL 4.5; lateral ocellus diameter 4.5; head width 44.5; mouth width 16; malar space 11.5; eye length 22; scape length 17; scape width 4.5; pedicel+flagellum length 48; pedicel length 6.5; pedicel width, dorsal view 3.5; F1 length 7.5; F1 width 4; F2 length 8.5; F2 width 4.5; F3 length 7; F3 width 5; clava length 16; clava width 5.3; C3 length 6; spicule length 1.5; mesosoma length 53; mesosoma width 41.5; midlobe of mesoscutum length 21; mesoscutellum length 21; mesoscutellum width 20.5; median part of mesoscutellum width (measured medially) 10; median part of mesoscutellum, width in anterior part10; median part of mesoscutellum, width in posterior part 10.5; lateral part of mesoscutellum, width (measured medially) 4.5; dorsellum length 2; propodeum length 7; costal cell length 29; costal cell width (measured at widest part) 2; marginal vein length 26; stigmal vein length 10; gaster length 84; gaster width 32; Gt 2 length (measured medially) 8; Gt 4 length (measured medially) 11; Gt 7 length (measured medially) 12.5; Gt 7 width (measured at base) 16; longest cercal seta length 11; shortest cercal seta length 5.

Male. Unknown.

Hosts. Reared from leaf gall induced by unidenitified Cecidomyiidae on Croton schiedeanus ( Euphorbiaceae). The gall is spherical, 3–7 mm in diameter, and detachable from the leaf.

Distribution. Costa Rica.

Material examined.

Holotype ♀ COSTA RICA, Heredia, Tirimbina, ii.2004, from spherical leaf gall on Croton Schn., Ley & Hanson (NHMUK) . Paratypes (3♀, MZLU, NHMUK): 1♀ COSTA RICA, Heredia, Estación Biológica La Selva, 10º26’N, 84º01’W, 100-200m, 30-31.iii.2002, swept, J.A. Azofeifa ; 1♀ from same locality as previous but collected 28-29.ii.2008; 1♀ COSTA RICA, Limón, Puerto Viejo, Hotel Punta Cocles, 27.ii.1991, J.S. Noyes.

Etymology. Named after the host plant of the gall from which this species was reared.