Physopelta (Neophysopelta) quadriguttata Bergroth, 1894

Physopelta quadriguttata Bergroth, 1894: 160–161 (description, distribution).

SYNTYPES: ♀♀, India, Sikkim (MZHF?) .

Physopelta quadriguttata: KIRKALDY & EDWARDS (1902): 165 (differences from Ph. indra); DISTANT (1903a): 98 (redescription, distribution; including Ph. indra); BERGROTH (1913): 167 (catalogue); ESAKI (1926): 157 (distribution); TAEUBER (1927): 176 and 177 (differences from Ph. lisae and Ph. roseni); HUSSEY (1929): 32 (catalogue, distribution); BLÖTE (1931): 100 (distribution); LIU (1981): 223–224, pl. 28 (key, redescription, figure); MANNA et al. (1985):621–630 (chromosomes, figures, distribution); Hu & Gui in ZHANG (1995): 111 (redescription, habitus, distribution, biology); HUA (2000): 187 (check-list, host plant, distribution); KERZHNER (2001): 246 (catalogue, distribution); STEHLÍK & JINDRA (2003): 7 (distribution); STEHLÍK (2004): 3 (distribution); MIYAMOTO et al. (2005): 11 –12 (habitus, distribution); STEHLÍK (2005a):144–145 (distribution); STEHLÍK (2007a): 127 (habitus, variability); RÉDEI et al. (2009): 13–14, 17, 48 (redescription, key, figures, distribution); VOTÝPKA et al. (2010): 246 (distribution); CHANDRA & KUSHWAHA (2012): 253 (distribution); ZHU et al. (2012): 195–198 (ecology, distribution).

Physopelta (Physopelta) quadriguttata: VOIGT (2006): 224 (key, distribution).

Material examined. CHINA: HAINAN: Jianfengling Mts., Tiachi Lake env., Bishu villa, 18°44′40″N 108°50′41″E, 950 m a.s.l., at light, 9.–11.v.2011, 1 ♂, M. Fikáček, V. Kubeček & L. Li lgt. (NMPC) ; Limushan Mts., first administr. Centre, 19°10′30″N 109°44′33″E, 630 m a.s.l., at light, 4.–6.v.2011, 1 ♀, M. Fikáček, V. Kubeček & L. Li lgt. (NMPC) . – SICHUAN: Baoxing, 100 km N of Yaan, 12. – 17.vii.l995, 1 ♀, Z. Jindra lgt. (ZJPC) ; 70 km NW Chengdu, Qingohenghou Shan Mts., 1,500 m a.s.l., 6.–13.viii.2010, 3 ♂♂ 2 ♀♀, S. Murzin lgt. (NMPC) ; Qingchenghou Shan Mts., 70 km NW Chengdu, 1500 m a.s.l., 6.–13.viii.2010, 3♂♂ 2 ♀♀, S. Murzin lgt. (NMPC) . – YUNNAN: Xima, Rk 12, 14°49′N 97°44′E, 2,080 m a.s.l., 11.vi.2001, 1♂, Deuve, Mantilleri, Rougerie & Tian lgt. (MNHN) . – TAIWAN: CHIAYI COUNTY: Fenchihu, 1,400 m a.s.l., 3.v.1977, 1 ♂, S. Klapperich lgt. (EHIA) . – HUALIEN COUNTY: S corner, Coastal Range SE of Fuli, 23°09′N 121°17′E, 500 m a.s.l., 12.–16.xi.2008, 36 ♂♂ 20♀♀, L. Dembický lgt. (MMBC) ; Hueisun, Experimental Forest Station, 700 m a.s.l., 3.viii.2002, 1 ♂, K. Schönitzer lgt. (ZSMC) . – NANTOU COUNTY: Meifeng, 24°05′25″N 121°10′21″E, 2,200 m a.s.l., 10.–11.v.2001, 1 ♂, K. Schönitzer lgt. (ZSMC) ; ditto, 2,100 m a.s.l., 10.–11.ix.2002, 3 ♂♂ 6 ♀♀, K. Schönitzer lgt. (ZSMC) . – PINGTUNG COUNTY: Pintung, Halen, Pettawushan, trail of 1,500 m a.s.l., 1.v.1992, 3 ♂♂, A. Smetana lgt. (MHNG) . – TAITUNG COUNTY: 14 km W of Chihshang, 23°11′09″N 121°04′21″, 900 m a.s.l., 16.xi.2008, 20 ♂♂ 42 ♀♀, L. Dembický lgt. (MMBC, NMPC, BMNH) . – VIETNAM: LAM DONG PROVINCE: Da Lat, 1,500 m a.s.l., 14.x.1988, 1♂ 1♀, 15.x.1988, 5♂♂ 3♀♀, T.Vásárhelyi lgt. (HNHM) . – LAO CAI PROVINCE: 17 km Dang Khao, 29.xi.1931, 1 ♀, Topál & Matskási lgt. (HNHM) ; Sa Pa, v.1990, 3 ♂♂ 2 ♀♀, Duong Tat Tu & J. Picka lgt. (NMPC) ; Tonkin, Chapa [= Sa Pa], v.1916, 1 ♂ 1 ♀, R. V. de Salvaza lgt. (BMNH) . – NGHE AN PROVINCE: Nghe An, 19°42.798′N 104°46.015′E, 800 m a.s.l., 8.–9.xi.1999, 1 ♂ 1 ♀, Kasai lgt. (HNHM) . – SON LA PROVINCE: Moe Chau [= Moc Chau], 800 m a.s.l., on light: 24.x.1986, 1 ♂, 25.x.1986, 1 ♀, 26.x.1986, 1 ♂, T. Vásárhelyi lgt. (HNHM). – VINH PHU PROVINCE: Tam Dao, 8 km N of Hanoi, xi.1986, 1 ♂, P. Bílek lgt. (ZJPC); Tam Dao, 900 m a.s.l., 17.–21.iii.1990, 1 ♀, 17.–21.v.1990, 1 ♀, J. Horák lgt. (NMPC) ; J. Horák lgt. (NMPC) ; 18.v.2007, 1 ♂, V. Socha lgt. (NMPC) . – YEN BAI PROVINCE: Luc Yen, 300 m a.s.l., 18.xi.1971, 1 ♀, Topál & Matskasi lgt. (HNHM) . – LAOS: ATTAPEU PROVINCE: Annam Highlands Mts., Dong Ampan NBCA, Nong Fa (crater lake) env., 15°05.9′N 107°25.6′E, ca. 1,160 m.s.l., 30.iv.–4.v.2010, 3 ♀♀, J. Hájek lgt. (NMPC). – HOUA PHAN PROVINCE: Ban Saluei, Phu Phan Mt., 20°13′N 103°59′E, 1,300–2,000 m a.s.l., 6.–18.iv.2005, 1 ♀, J. Bezděk lgt., Z. Jindra det. (NMPC); 30 km S of Xam Neua, Phou Pan Mt., 1,500 m a.s.l., 6.–17.v.2004, 1 ♀, P. Kresl & F. Kantner lgt. (ZJPC) ; Phu Phan Mt., 20°12′N 104°01′E, 1,500–1,900 m a.s.l., 17.v.–3.vi.2007, 7 ♂♂ 5 ♀♀, M. Brancucci & V. Kubáň lgt. (MMBC, NMPC) ; Ban Saluei → Phou Pane Mt., 20°12–13.5′N 103°59.5′–104°01′E, 1340–1870 m a.s.l., 15.iv.–15.v.2008, Lao collectors lgt. (NMPC) . – THAILAND: MAE HONG SON PROVINCE: Soppong, 19°27′N 98°20′E, 1,500 m a.s.l., 15.–17.xi.1996, 2 ♂♂ 2 ♀♀, S. Bečvář lgt. (MMBC).

Distribution. China (Anhui, Fujian, Guangdong, Guangxi, Henan, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Xizang, Zhejiang, Hainan) (LIU 1981, Hu & Gui in ZHANG 1995, KERZHNER 2001, VOTÝPKA et al. 2010; this paper), Taiwan (ESAKI 1926, KERZHNER 2001, RÉDEI et al. 2009; this paper), Japan (Ryukyus) (MIYAMOTO et al. 2005), India (Madhya Pradesh, Nagaland, Sikkim, West Bengal) (BERGROTH 1894, DISTANT 1903a, BLÖTE 1931, MANNA et al. 1985, CHANDRA & KUSHWAHA 2012), Nepal (STEHLÍK 2004), Thailand (STEHLÍK & JINDRA 2003; this paper), Laos (STEHLÍK 2005a; this paper), Vietnam (new record).

A record from Puducherry: Mayyazhi District (India) by DISTANT (1903a) belonging to Ph. indra, was erroneously considered as a junior synonym of Ph. quadriguttata in that paper.