SUBFAMILY SCIADOCERINAE

DIAGNOSIS: Pretarsus with empodium either minute or completely lost. Acrostichals long, in 1–3 rows (except Ulrichophora). Plesiomorphic features include: cells cup and bm present (closed); frons without median sulcus; tip of R 4+5 fully fused to C; sternites fully developed in both sexes.

COMMENTS: There are two living sister species of Sciadocerinae, Archiphora patagonica Schmitz in Chile and Sciadocera rufomaculata White in Austalia and New Zealand. Their close relationship is indicated by characters 14 and 15 (table 4): stem of M 1 -M 2 lost or spectral, and the basal flagellomere is enlarged in males (it is much larger in Archiphora). A Baltic amber (Eocene) species was transferred to its own genus, Hennigophora robusta (Meunier), by Brown (2007a), into which I am also transferring Archiphora pria Grimaldi and Cumming (see below). Two other genera of sciadocerines occur in Baltic amber: Ulrichophora Brown (monotypic: lobata), and Eosciadocera Hong (two species). I have not examined Ulrichophora, but the fly has some distinctive features, the most striking being the numerous acrostichals not arranged into one or a few rows (Brown, 2007a). Eosciadocera setosa and E. pauciseta, n. sp., are striking because of their large size, nearly 7 mm in body length.