Liphistius pinlaung sp. nov. Figs 4, 5

Type material.

Holotype: MYNAMAR · ♂; Shan State, Pinlaung Township, ca.14 km to Pinlaung from Pekon; 20.02N, 96.79E; alt. 1410 m; 19 July 2018; D. Li, F.X. Liu, X. Xu and L. Yu leg.; XUX-2018-164. Deposited in CBEE.

Paratype: MYNAMAR · 1 ♂, 5 ♀♀; same data as for holotype; XUX-2018-162, 167, 169, 169A, 169B, 169J; 19 July 2018. All specimens deposited in CBEE.

Diagnosis.

Males of L. pinlaung sp. nov. resemble those of L. birmanicus, L. lordae and L. lahu by the wide paraembolic plate, but can be distinguished from L. birmanicus by the lack of lateral process of paracymbium and by the cumulus with longer and stouter setae (Fig. 4C, D); from L. lordae by the wider tibial apophysis at base (Fig. 4D) and the tegulum with a dentated margin (Fig. 4C, F); from L. lahu by the narrower tegulum (Fig. 4C, F) and smaller paracybium (Fig. 4D, E). Females of L. pinlaung sp. nov. resemble those of L. birmanicus and L. hpruso sp. nov. by the poreplate with two pair of lobes, but can be distinguished from L. birmanicus by the wider posterior stalk, and sphere-shaped receptacular cluster (Fig. 5 D–F); from L. hpruso sp. nov. by the wider posterior stalk and larger anterior lobes of the poreplate (Fig. 5 A–F); from the other Liphistius by the poreplate with four anterior lobes (Fig. 5 D–F).

Description.

Male (holotype). Total length, excluding chelicerae, 12.71. Carapace 5.86 long and 5.47 wide, longer than wide, olive-green due to being fixed in ethanol immediately after molting, furnished with few short, scattered bristles (Fig. 4A). ALE>PLE>PME>AME, eye sizes and interdistances: AME 0.05, ALE 0.55, PME 0.31, PLE 0.48, AME-AME 0.10, AME-ALE 0.07, PME-PME 0.09, PME-PLE 0.09, ALE-PLE 0.09, ALE-ALE 0.11, PLE-PLE 0.38, AME-PME 0.09. Chelicerae robust, promargin of chelicerae groove with ten strong denticles of variable size. Labium 0.86 long and 0.89 wide, wider than long, fused with sternum and slightly pale olive-green (Fig. 4B). Sternum 2.94 long and 1.05 wide, longer than wide, and a few weakly spined setae on the anterior tip and many long spined setae on the posterior tip, elongated posterior tip (Fig. 4B). Opisthosoma 7.17 long and 4.92 wide, with 12 tergites, the fifth largest, eight spinnerets (Fig. 4B). Legs with strong hairs and spines. Measurements: leg I 16.99 (4.32 + 2.55 + 3.55 + 4.66 + 1.92), leg II 18.06 (4.32 + 2.41 + 3.74 + 5.18 + 2.41), leg III 18.46 (4.44 + 1.85 + 2.83 + 6.68 + 2.66), leg IV 20.40 (3.56 + 1.52 + 4.25 + 8.46 + 2.63).

Palp: Tibial apophysis with four long spines of different lengths (Fig. 4D, E), paracymbium large and wide, many setae situated at the tip and a row of several tapering spines one the indistinct cumulus (Fig. 4C, D, F); subtegular apophysis weakly developed (Fig. 4C); contrategulum with conical, tip blunt with a short process (Fig. 4C, F), distal edge widely arched, with a smooth and sharp edge (Fig. 4 F–H); tegulum small and the terminal apophysis with finely dentated margin (Fig. 4C, E, F); paraembolic plate short, widely rounded, embolic parts adjacent (Fig. 4D, F, H); embolus long and conical, basally sclerotized, with 3-4 longitudinal ridges that reach to tip (Fig. 4C, D, F).

Female. Total length, excluding chelicerae, 14.46. Carapace 6.70 long, 6.07 wide, light brown, furnished with few short, scattered bristles. Four thick setae on clypeus. Eight eyes on darkened ocular tubercle, ALE> PLE> PME> AME, eye size and interdistances: AME 0.09, ALE 0.61, PME 0.33, PLE 0.47, AME-AME 0.11, AME-ALE 0.16, PME-PME 0.13, PME-PLE 0.13, ALE-PLE 0.14, ALE-ALE 0.14, PLE-PLE 0.43, AME-PME 0.14. Chelicerae proximally glabrous, robust, dark brown; promargin of chelicerae groove with 14 strong denticles of variable size. Labium 0.75 long, 1.19 wide, slightly pale brown. Sternum 3.25 long, 1.59 wide, brown and weakly spined, a few setae on the outside of this area, elongated posterior tip. Opisthosoma 8.20 long, 5.73 wide, dark brown, with 12 tergites, the fifth largest, and eight spinnerets. Legs brown with strong hairs and spines, long and short black sparse setae, legs each with three tarsal claws. Measurements: palp 8.59 (2.01 + 1.67 + 2.65 + 2.27), leg I 11.75 (3.39 + 1.99 + 3.03 + 2.01 + 1.33), leg II 12.02 (2.69 + 2.05 + 3.14 + 2.45 + 1.68), leg III 13.47 (4.19 + 1.22 + 3.51 + 2.49 + 2.05), leg IV 22.4 (6.47 + 2.58 + 4.38 + 5.82 + 3.15).

Female genitalia: pore plate with a pair of large anterior lobes and a pair of small, strongly elevated anterolateral lobes, and anterior lobes larger than anterolateral lobes (Fig. 5 D–F); distinct transition between the pore plate and posterior stalk (Fig. 5 A–F); posterior stalk wide; receptacular cluster spherical and small; central dorsal opening small and circular (Fig. 5 A–C).

Entomology.

" pinlaung " refers to the type locality of this species.

Distribution.

Myanmar (Pinlaung Township, Shan State).

Variation.

Body measurements, see Table 1. The examined female genitalia differ from each other; for the specimen of XUX-2018-169A, the central part of anterior and anterolateral lobes of the pore plate are depressed in the dorsal view (Fig. 5B), whereas the depression is absent in the other two specimens (XUX-2018-167 and 169J); the shape and size of anterior and anterolateral lobes of the pore plate, as well as the shape of anterior margin of the pore plate are rather variable (Fig. 5 A–F). The size of the receptacular cluster is also slightly different (Fig. 5 D–F).

Relationships.

Liphistius hpruso sp. nov. and L. pinlaung sp. nov. belong to the birmanicus -group that currently contains L. birmanicus, L. lordae and L. lahu based on morphological characters (Schwendinger, 1998). The two new species are closer to L. birmanicus than to L. lordae and L. lahu since their female poreplates possess four anterior lobes (Figs 3 B–E; 5 D–F).