Pteropus mariannus Desmarest, 1822 . Mammalogie, Part 2, Suppl., p. 547.

TYPE LOCALITY: Marianna Isis., Guam (U.S.) .

DISTRIBUTION: Palau Isis.; Mariana Isis.; Caroline Isis.; Ryukyu Isis.

COMMENT: Includes loochooensis, pelewensis, ualanus, and yapensis; see Kuroda, 1938, A List of the Japanese Mammals, 122 pp.; Corbet and Hill, 1980:37-38, listed these forms as distinct species without comment.

ISIS NUMBER: 5301405001024025001.