Vermicularia milleti (Deshayes, 1850)

Figs 29A 1 –A 2, 29B 1 –B 2, 20C

* Vermetus Milleti Desh. — Deshayes, 1850: 43, pl. 70, figs 9–10.

Vermetus carinatus Hörn. — Hörnes, 1856: 486, pl. 46, fig. 17 [non Vermetus carinatus Quoy & Gaimard, 1834].

Turritella cf. terebralis Lmk. — Boettger, 1901: 153, no. 471.

Vermetus (Vermicularia) pseudoturritella n. sp. — Boettger, 1907: 171, no. 591.

Vermicularia Milleti (Deshayes) — Cossmann & Peyrot, 1922: 86, nr. 463, pl. 3, figs 14, 32.

Vermetus (Vermicularia) milleti pseudoturritella Boettger — Zilch, 1934: 220, pl. 7, figs 25a–25b.

Vermetus (Vermicularia) milleti Deshayes — Glibert, 1949: 128, pl. 7, fig. 10.

Vermetus (Vermicularia) milleti pseudoturritella Boettger — Strausz, 1966: 121, pl. 3, figs 22–23.

Vermicularia milleti (Deshayes) — Bałuk, 1975: 123, pl. 14, figs 6–7.

Vermetus (Vermicularia) pseudoturritella Boettger — Atanacković, 1985: 99, pl. 24, figs 1–2.

Vermicularia milleti (Deshayes) — Bałuk, 2006: 194, pl. 6, figs 5–6.

Vermicularia milleti (Deshayes, 1839) — Landau et al., 2004: 26, pl. 3, fig. 18.

milleti (Vermetus) Deshayes, 1850 — Bieler & Petit, 2011: 47.

Vermicularia milleti (Deshayes, 1839) — Landau et al., 2013: 65, pl. 5, fig. 17.

Vermicularia milleti (Deshayes, 1839) — Vaessen, 2013: 43, figs 4–7.

Vermicularia milleti (Deshayes, 1839) — Landau et al., 2018: 195, pl. 18, figs 1–4.

Type material. The Deshayes collection, which was part of the ‘Ecole des Mines Collection’, is stored at the Université Claude Bernard since 1978. Seven syntypes of Vermetus milleti are glued on a label; the syntype illustrated by Deshayes (1850) has the inventory number UCBL-EM 32115.

Studied material. 2 spec. NHMW 1856/0035/0193, Steinebrunn (Austria), syntypes of Vermetus carinatus Hörnes, 1856, 1 spec. NHMW 1876/0011/0077, 1 spec. NHMW 2016/0177/0350, Lăpugiu de Sus (Romania) .

Illustrated material. Figs 29A 1 –A 2: NHMW 1856/0035/0193, SL: 22.9 mm, MD: 16.1 mm, Steinebrunn (Austria), syntype of Vermetus carinatus Hörnes, illustrated in Hörnes (1856, pl. 46, fig. 17); Figs 29B 1 –B 2: NHMW 1856/0035/0193, SL: 17.5 mm, MD: 10.6 mm, Steinebrunn (Austria), syntype of Vermetus carinatus Hörnes; Fig. 29C: NHMW 1876/0011/0077, SL: 13.7 mm, MD: 8.5 mm, Lăpugiu de Sus (Romania), syntype of Vermetus carinatus Hörnes.

Discussion. Bałuk (1975, 2006) and Landau et al. (2004, 2018) described this species in detail. Most Paratethyan specimens are characterized by two prominent keel-like spiral cords on early teleoconch whorls, resulting in an Archimediella -like appearance. Specimens with three spiral cords and a weak adapical spiral cord are present as well. Therefore, we agree with Bałuk (1975, 2006) and Landau et al. (2004) in synonymizing the Paratethyan Vermetus carinatus Hörnes 1856 [non Quoy & Gaimard 1834] and Vermetus (Vermicularia) pseudoturritella Boettger, 1907 with Vermicularia milleti (Deshayes, 1850) .

Paleoecology. Saint Martin et al. (2000) mention the species from a Badenian micriobialitic coral reef in Hungary. Other occurrences from Korytnica (Poland), Steinebrunn (Austria) and Estepona (Spain) are not reef-associated but represent various clayey to sandy inner to middle neritic environments.

Distribution. Central Paratethys. Badenian (middle Miocene). Vienna Basin: Steinebrunn (Austria) (Hörnes 1856); Pannonian Basin: Várpalota, Érd-Diósd (Hungary) (Strausz 1966; István 1985; Saint Martin et al. 2000); southern Pannonian Basin: Hrvaćani (Bosnia and Herzegovina) (Atanacković 1985); Carpathian Foreland Basin: Korytnica (Poland) (Bałuk 2006); Transylvanian Basin: Coşteiu de Sus, Lăpugiu de Sus (Romania) (Boettger 1907; herein). Northeastern Atlantic. Burdigalian: Aquitaine Basin: Saucats (Peloua) (France) (Cossmann & Peyrot 1922); Langhian: Aquitaine Basin: Manciet, Salies-de-Béarn (France) (Cossmann & Peyrot 1922); Loire Basin: Paulmy- Pauvrelay, Ferrière Larson-La Placette, Ferrière Larfon-Les Verdaux (France); Serravallian: Loire Basin: Mirabeau, Ambèrre-Moulin Pochas (France); Tortonian and Messinian: Loire Basin: Sceaux-d’Anjou-La Presselière, Le Grand Chauvereau, St-Clément-de-la-Place, Renauleau, Maine-et-Loire (NW France) (Vaessen 2013; Landau et al. 2018). (proto-) Mediterranean Sea. Serrvallian: Karaman Basin: Akpinar (Turkey) (Landau et al. 2013); late Pliocene: western Mediterranean, Estepona (Spain) (Landau et al. 2004).