Ptychidia studeri (Rutsch, 1929)

T. [urritella] terebra— Studer, 1825: 332 [non Turritella terebra Linnaeus, 1758).

T. Studeri May — Mayer, 1853: 96, nr. 242 [nomen nudum].

Turritella turris— Kissling, 1890: 35, pl. 6, fig. 43 [non Turritella turris de Basterot, 1825].

* Turritella turris var. studeri Rutsch, 1929: 36, pl. 2, figs 5–6.

Turritella (Haustator) turris studeri Mayer — Sieber, 1960: 234, pl. 1, figs 9, 26.

Turritella studeri Rutsch, 1929 — Pfister & Wegmüller, 2007: 93, pl. 9, figs 7–15, pl. 10, figs 1–3.

? Turritella studeri Mayer — Kókay, 2002: 371, pl. 2, fig. 2 [non fig. 1].

Turritella studeri— Pfister et al., 2011: 162.

Discussion. A very dubious species, which was based by Rutsch (1929) on poorly preserved internal casts. One of the illustrated syntypes (Rutsch 1929, pl. 2, fig. 6) seems to represent Turritella terebralis Lamarck, 1822, in respect to its characteristic lateral sinus. Most of the specimens described by Pfister & Wegmüller (2007), however, may document a distinct species, typical for the Swiss early Miocene. Sieber (1969) and Pfister & Wegmüller (2007) stressed a relationship of T. studeri with ‘ Turritella eryna ’ (= Ptychidia erynella new sp.), which might suggest a placement of T. studeri in Ptychidia .

Distribution. Western Paratethys. North Alpine Foreland Basin: Burdigalian: (Eggenburgian/ Ottnangian) Switzerland (Bern region, St. Gallen, Belpberg), Austria (Bregenz, Wirtatobel) (Pfister et al. 2011).