Phrynobatrachus natalensis (Smith, 1849)

Snoring Puddle Frog (Fig. 27; Map 23)

Material (20 specimens): PEM A12637–8, Cuanavale River, -13.37406° 18.99269°, 1,297 m asl; PEM A12711, Lungwebungu River camp bridge crossing, -12.58347° 18.66598°, 1,304 m asl; PEM A12890–3, INBAC: WC-4599, INBAC (no number), Quembo River source lake, -13.13624° 19.04591°, 1,366 m asl; PEM A13738, Comba River, -12.62442° 18.65159°, 1,299 m asl; PEM A13745–8, Lungwebungu River old campsite, -12.58319° 18.66573°, 1,284 m asl; PEM A14690, PEM A14705–6, INBAC: WC-6740, Lungwebungu River camp, -12.58439° 18.66748°, 1,297 m asl; PEM A14707, wetland west of Lungwebungu River camp, -12.55855° 18.63770°, 1,308 m asl; PEM A14810–1, Luio River camp floodplains, -13.19711° 20.22194°, 1,181 m asl. Additional material (1 specimen): SAIAB 209104 (1 specimen), swamp near Cuanavale River source lake camp, -13.10750° 18.86089°, 1,386 m asl. Description: Medium sized Phrynobatrachus; dorsum with scattered elevated tubercles; reduced webbing; heel spine present; small outer metatarsal tubercle; large inner metatarsal tubercle; ridge running along outer toe; small tarsal ridge; mid-tarsal tubercle present; well-developed elevated subarticular tubercle. Dorsum coloration varies from grey to brown and even green; ventrum white. Female throats are speckled, while throats of males are uniformly grey to black. Adult females (n = 15) varied from 23.6–32.6 (28.1) mm (largest female: PEM A13745); adult males (n = 5) varied from 26.3–30.4 (28.2) mm (largest male: PEM A14705). Habitat and natural history notes: Found in flooded grasslands associated with miombo woodland. Comments: This species is widespread in Angola (Marques et al. 2018). Although there were no records for southeastern Angola prior to Conradie et al. (2016), they are widespread east of the Zambian border (Poynton and Broadley 1985b, 1991; Channing, 2001). This is another complex within Phrynobatrachus containing several cryptic species, and thus deserving of further investigation (Zimkus et al. 2010; Bittencourt-Silva 2019).