Drasteria cailino obscura (Staudinger, 1901)

(Figs. 1–3 (imago), 208, 209 (male genitalia))

Staudinger in Staudinger et Rebel, 1901: 243 ( Leucanitis Cailino Lef. v. Obscura). Type locality: “Ross. mer. or. (Narün), Tarb. et Ala Tau (trans.)” (by original description); “Narün” (by the lectotype designation, designated here) [“nordöstlich von Astrachan zwischen Wolga- und Ural-Fluss, … etwa 15 deutsche Meilen ostwärts von der Wolga” (Staudinger, 1879: 315)].

Type material: Lectotype male (designated here in accordance with ICZN Art. 74) (Figs. 1–3), labelled: “Narün | Henke” (green paper, handwritten), “Origin.” (pink paper, printed), “ex coll. 1/6 | Staudinger” (white paper, printed with handwritten numbers), “ LECTOTYPUS ♂ | obscura | Stgr. | S.K. Korb des. 2016” (red paper, printed with handwritten taxon name and author) (ZMHU) . Paralectotypes 3 males, 2 females (ZMHU) .

Butler, 1892: 315 [Asia Minor, Caucasus, Schuscha]; Culot, 1913: 184 [la Russie méridionale]; Aibasov, 1974: 122 [Shevchenko, Mangyshlaksky Karatau: Syir-Su dwell]; Kumakov, Korshunov, 1979: 153 [4 th Dacha, Saratov]; Aibasov, Zhdanko, 1982: 30 [Aksu in Zelinograd Prov.]; Ishkov, Sukhareva, 1986: 124 [Aksu-Dzhabagly Nature Reserve: Novonikolaevka, Dzhabagly, Aksu river canyon, Kshi-Kaindy, Ulken-Kaindy, Aksay]; Poole, 1989: 329; Anikin et al., 2000: 332 [Saratov]; Hacker, Miatleuski, 2001: 814 [Bisen]; Nupponen, Fibiger, 2002: 135 [Arkaim, Bajmak, Kidriasovo, Kuvandyk, Moskovo, Sakmara river, Verbljushka]; Poltavsky, Ilyina, 2002: 17 [Gunib, Tshervlennie Buruni]; Poltavsky, Nekrasov, 2002: 26 [Rostov-on-Don Region, Daghestan Republic]; Goater et al., 2003: 79 [South Europe, in Portugal, Spain, south France, Italy, Sicily and the Balkan countries. Recorded also from South Russia: Saratov District and Kazakhstan: Bisen. Outside Europe, in Middle East, Asia Minor, and Central Asia]; Poltavsky, Ilyina, 2003: 171 [Leninaul, Tchervlennie Buruni]; Poltavsky et al., 2010: 93 [Botkul]; Kononenko, 2010: 89 [Kazakhstan]; Gorbunov, 2011: 56 [steppes and northern deserts of West Kazakhstan]; Lehmann & Zahiri, 2011: 150 [Iran: Gate Dehh, Yush]; Volynkin, 2012: 27 [Kazakhstan part of Altai]; Poltavsky, 2016: 19 [Rostov-on-Don Province]; Shovkoon, Trofimova, 2016: 606 [Aktobe Prov., near Kompressorniy vill. N 48°30’; E 59°19’; Aktobe Prov., Mugodzhary Mts., N 48°26’; E 58°24’; Aktobe Prov., Bolshie Barsuki sands, N 46°58’; E 59°13’; Aktobe Prov., Shagyrai Plateau, N 46°44’; E 57°30’; Kyzyl-Orda Prov., Karatup penin., Aral lake, N 46°20’; E 59°43’]; Titov et al., 2017 a: 148 [Pavlodar region, vic. of Pavlodarskoe vill., 52°22.084’N 76°52.161’E; Pavlodar region, Tuz lake, 51°17’58.24”N 78°37’57.06”E; Pavlodar region, Bayanaul Mts., Birzhankol lake, 50°49’9.97”N 75°20’52.44”E; Pavlodar region, Bayanaul Mts., Kurkeli natural landmark, 50°44’34.00”N 75°38’16.80”E; Pavlodar region, Bayanaul Mts., vic. of Kempirtas Mt., 50°51’24.65”N 75°34’37.21”E; Pavlodar region, Bayanaul Mts., vic. of Toraighyr lake, 50°51’54.01”N 75°40’21.29”E; Pavlodar region, Bayanaul Mts., Kirgichi natural landmark, 50°50’26.05”N 75°44’15.03”E; Pavlodar region, Zhartas natural landmark, 51°38’7.49”N 74°39’54.86”E].

Habitat (Fig. 299). Steppes, deserts and semideserts. Elevation: 200–2700 m.

Distribution. Astrakhan, Saratov, Orenburg and Volgograd Prov., Krasnodarsky Kray, Daghestan (Russia), Ukraine, North and Central Kazakhstan, Armenia, Azerbaijan, North-West Iran, North-West Turkey.

Material examined (in total: 221 specimens). Armenia: 1 ♂, 1.08.1934, Alagez, southern slope, 8500’, [leg. M. Ryabov] (ZISP) ; 1 ♂, 18.07.1936, Alagez vic., Inaklu river, [leg. M. Ryabov] (ZISP) ; 1 ♂, 8.07.1937, Arpa river, [leg. M. Ryabov] (ZISP) ; 1 ♀, 16.07.1939, Gedzhanan, h=7000–8000’, meadow, [leg. M. Ryabov] (ZISP) ; 1 ♀, 28.05.1960, Erevan, leg. Vardikian (ZISP); 1 ♂, 15.07.1963, Daralagez, Ger-Ger, [leg. E. Milyanovsky] (ZISP) ; 1 ♂, 18.06– 7.07.1996, Gegamsky range, Vedy river, right bank, Khosrov reserve, h= 2000 m, leg. A. Danchenko, ex coll. А. Nekrasov, slide ffi2748 Nekrasov (ZISP) ; 1 ♂, 20.06.2002, Gegamsky range, Khosrov res., leg. M. Ka- lashian, ex coll. А. Nekrasov (ZISP) ; 3 ♂, 18.07.1936, Aragaz Mt., Antarut ( Inaklyu), leg. A.V. Zvetaev (ZMMU) ; 1 ♂, 10.07.1965, Aragaz Mt., Kara-Gel’ lake, 3300 m, leg. A.V. Zvetaev (ZMMU) ; 1 ♂, 4.06.1970, Nakhichevan, Daralagezsky Mts., Buzgov, 1400 m, leg. A.V. Zvetaev (ZMMU) . Azerbaijan: 1 ♂, Helendorf, ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♂, Ordubad, leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich (ZISP); 1 ♂, 4 ♀, 15.06.1881, 7.05– 15.06.1883, Ordub [ad], leg. Chr [istoph], ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♂, 9.07.1937, Sultanbek, [leg. М. Ryabov] (ZISP) ; 1 ♀, 20.08.1939, Kupudzhik, upper forest zone, h=8000’, [leg. М. Ryabov] (ZISP) ; 1 ♂, 14.06.1967, Ordubad, Pazmara, h=1800 м, leg. R. Effendi (ZISP) ; 1 ♂, 22.07.1986, Nakh- ichevan ASSR, Buzgov, leg. A. Ivanov (ZISP) ; 1 ♂, 16.05.2014, Talysh, Yardymly vic., Deman, leg. V. Tikhonov (OPB) . Kazakhstan. 1 ♀, 18.05.1899, Turgaisk Prov., Kaban-Kulak Mt. (20 verst SW Irgiz), leg. Sushkin (ZISP) ; 1 ♀, 25.05.1907, Turgaisk Prov., Bolshie Barsuki, Chelkala lake, leg. Volman (ZISP) ; 1 ♂, 29.06.1907, Uralsk, leg. B. Uvarov (ZISP) ; 1 ♂, Karaganda Prov., 19.06.1958, 40 km S Zhana-Arka, Koksengir Mt., leg. A. Zagulyaev (ZISP) ; 1 ♂, 4– 5.05.2005, Akkum-Sagyz sands, leg. O. Novikov, slide OP 3141m (OPB) ; 1 ♂, 19.06.2010, Kara- ganda Province, Zhaksy-Arganaty Mts., Karaatan, h= 500 m, 49 ° 14’ N 67 ° 01 E, leg. P. Gorbunov (ZISP) ; 1 ♂, 30.06– 01.07.2010, 33 km N of Panfilov, Boro-Khoro Mts., N 44 29.765 E 80 03.848, 1830 m, leg. S.K. Korb (SKK) ; 2 ♂, 10.07.2011, Aktyubinsk Prov., Mugodzhary, N48° 32.172’ E58° 29.345’, 520 m, leg. S.K. Korb (SKK) ; 11 ♂, 30.07.2011, Akmola Prov. near Zhaksy, N51° 52.769’ E67° 29.952’, leg. S.K. Korb (SKK) ; 1 ♀, 26.04.2012, West Kazakhstan, Kumzhargan sands at Zhagabulak vill., 47 km SW of Emba town, 48º33’N, 57º37’E, leg. P. Gorbunov (OPB) ; 1 ♀, 12.06.2012, Aktyubinsk Prov., Mugodzhary, Shauyryk river, 48°30’36.6” N 58°50’26.6” E, leg. Т. Trofimova and D. Shovkun (ZISP) ; 1 ♂, 1 ♀, 30.06– 1.07.2012, Alma-Ata Prov., Panfilof distr., 25 km SW of Koktal, Ili river, leg. S. Rybalkin (CKO) ; 1 ♂, 7– 8.05.2013, Alma-Ata Prov., 28 km NW of Bakanas, Ili river, leg. S.A. Knyazev (CKO) ; 7 ♂, 3 ♀, 2.05.2014, Bol. Barsuki sands, 10 km E of Chelkar, 47°49’ N 59°46’ E, leg. P. Gorbunov (ZISP) ; 2 ♂, 1 ♀, 1– 3.07.2015, Ili river valley near Kok-Tal bridge, N43°57.818’, E79°36.322’, 500 m, leg. S.K. Korb (SKK) ; 2 ♂, 1 ♀, 4– 6.07.2015, Boro-Khoro Mts., Usek valley, N44° 28.082’ E79° 49.760’, 1260 m, leg. S.K. Korb (SKK) ; 2 ♂, 07.07.2015, Boguty Mts., Chingilsu valley, N43° 34.825’ E78° 39.847’, 851 m, leg. S. K. Korb (SKK) . Ukraine: 3 ♂, 17.05.1982, Kherson Prov., Chenromorsky Nature Reserve, Rybalchinsky Kordon, leg. I. Plyustsh, ex coll. A. Nekrasov, slide ffi1208 Nekrasov (ZISP) . Russia. 4 ♂, 2 ♀, Narün, Henke (lectotype and paralectotype) (ZMHU) ; 1 ♂, Sarepta, leg. unknown, ex coll. O. John (ZISP) ; 4 ♀, Guberli, leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♂, 25.06.[?], Guberli, leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♂, 1 ♀, Spask, [leg. Eversmann], ex coll. Eversmann (ZISP) ; 12 ♂, 4 ♀, 2.05.2010, Astra- khan Prov., Dosang, leg. S.K. Korb (SKK) ; 1 ♂, 4.05.2010, Saratov Prov., Melovoe, leg. S.K. Korb (SKK) ; 2 ♂, 7.05.2010, Volgograd Prov., Eruslan river valley, leg. S.K. Korb (SKK) ; 1 ♂, 12.05.2019, Volgograd Prov., 17 km SSE Frolovo, 49°39’39”N 43°28’23”E, leg. A. Samus (SKV) ; 1 ♂, 1 ♀, Caucasus, leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♂, 1885, Caucasus, leg. Lederer. ex coll. Acad. Petrop (ZISP) ; 1 ♀, 22.06.1893, Alupka, leg. Melioransky (ZISP) ; 1 ♂, 2 ♀, 12.05.1904, 22– 29.06.1905, Ai-Petri, slope near Yalta, leg. S. Tshet- verikov (ZISP) ; 1 ♂, 3.06.1905, St [ary] Kr [ym], [leg. Pliginsky] (ZISP) ; 7 ♂, 4 ♀, 1.06.1905, 20.04– 22.05.1906, Agarmysh, [leg. Pliginsky], ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♀, 11.06.1905, distr. Jaltensis, loc. Oreanda, leg. S. Tshetverikov (ZISP) ; 1 ♂, 8 ♀, 13.06.1905, 28.05.1907, 26.05– 26.06.1908, distr. Jaltensis, mons Ai-Petri, southern slope, leg. S. Tshetverikov, ex coll. S. Tshetverikov (ZISP) ; 1 ♂, 26.05.1907, Simferopol, leg. G. and K. Khristoforov, ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♀, 3.06.1907, distr. Jaltensis, loc. Mis- chor, lower border of pine forest, leg. S. Tshetverikov (ZISP) ; 2 ♂, 4 ♀, 26.05– 24.06.1908, distr. Jaltensis, loc. Koreiz, stony scree near Alupka, leg. S. Tshetverikov, ex coll. S. Tshetverikov (ZISP) ; 1 ♂, 1 ♀, 4.06.1910, distr. Jaltensis, mnt. Ai-Petri, screes over Alupka, leg. S. Tshetverikov, ex coll. S. Tshetverikov (ZISP) ; 1 ♂, 1 ♀, 23– 25.06.1913, distr. Theodosia, Staryi Crim, leg. S. Tschetverikov (ZISP) ; 1 ♀, 31.07.1924, Karadag, leg. A. Dyako- nov (ZISP) ; 2 ♂, 29.06.1926, 20.06.1944, Daghestan, Chodzhalmachi, leg. M. Rjabov (ZISP) ; 1 ♀, 3.08.1926, Kumtorkale, leg. M. Ryabov (ZISP) ; 1 ♀, 15.06.1992, Daghestan, Rutul, leg. O. Pekarsky, slide OP3140f (OPB) ; 1 ♂, 27.07.2001, 25 km SW Buynaksk, Balakhani vill., leg. A. Magomedova (ZISP) ; 2 ♂, 24.06.2002, Ay-Petri yayla, h=1300 м, leg. S. Sinev (ZISP) ; 1 ♂, 18.06.2008, Krasnodar Prov., Seversky distr., Papay Mts., arid semiforest, southern slope steppes, h=700–800 м, leg. V. Stshurov (ZISP) ; 1 ♀, 28.05.2009, Crimea, Chiginitra, leg. V. Savchuk (OPB) ; 1 ♂, 29.06.2010, Old Crimea, Agarmish Mt., leg. V. Savchuk (OPB) . Tajikistan. 1 ♂, 1 ♀, 16.07.2011, Shugnansky Mts., Gunt river valley, 2670 m, N 37 41.887, E 71 55.821, leg. A. Nikolaev, slide OP 3171m, OP3172f (OPB) ; 1 ♂, 2 ♀, 17– 18.06.2012, Gissar Mts., distr. Varzob, vill. Kondara, 1150–1200 m, leg. E. Rutjan, slide OP 3171m, OP3172f, OP3182f (OPB) . Turkey. 1 ♀, Asia m[inor], leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♀, 26– 27.07.1984, Prov. Ağrý, 39º57´N, 43º14´E, Asgüneyi Dağlarý 2000 m, 3 km nö Çu- maçay, leg. H. Hacker (HNHM) ; 1 ♂, 4.07.2002, Prov. Hakkari, 30 km NE of Hakkari, 1500 m, 37º41´N, 43º55´E, leg. B. Benedek and T. Csővári (HNHM) ; 2 ♂, 1 ♀, 7– 8.07.2002, Prov. Agri, 2000 m, Karasu-Aras mts, 5 km SE of Sarican, 39º47´N, 42º28´E, leg. B. Benedek and T. Csővári (HNHM) . Turkmenistan. 1 ♂, Askh [abad], [leg. Chris- toph], ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♂, Aschab [ad], ex coll. Bramson (ZISP) ; 2 ♂, Tekke (ZMHU) ; 1 ♂, 1 ♀, 29.05– 10.06.1882, Nochur, leg. Chr [istoph]. ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♂, 2 ♀, 20.05.[?], Aidara, leg. Eylandt, ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♀, 4.07.1892, Aidere, [leg. Christoph], ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 5 ♂, 6 ♀, 15.06– 20.08.1894, Transcaspien, Sumbar, leg. Herz, ex coll. Grand Duke Nikolai Mikhailovich (ZISP) ; 1 ♂, 29.04.1952, 55 km E Kara-Kala, Ai- Dere, leg. V. Kusnetzov (ZISP) ; 1 ♀, 27.06.1952, Kara-Kala, garden of VIR, leg. V. Kuznetsov (ZISP) ; 5 ♂, 2 ♀, 7.04– 6.07.1953, 15 km NO Kara-Kala, Iol-Dere, leg. V. Kuznetsov (ZISP) ; 5 ♂, 5 ♀, 18.06– 18.07.1954, 22.06.– 7.08.1956, mts. Kopet-Dag, valles Boboso, leg. W. Potopolskij (ZISP) ; 1 ♂, 3 ♀, 7.07.1954, 16.05.1955, Aschabad circ., Tschuli, leg. W. Potopolskij (ZISP); 1 ♂, 1 ♀, 12.06.1960, Askhabad vic., leg. Potopolsky (ZISP) ; 1 ♂, 3 ♀, 1.05– 11.06.1986, Aidere, h=850 м, leg. M. Falkovich (ZISP) . Iran. 1 ♀, 31.05– 2.06.2014, Prov. Mazandaran, near Gaschar vill., N 36º6’36.48”, E 51º20’1.85”, leg. D. Kasatkin, I. Shokhin, slide OP3182f (OPB) .