Drasteria kusnezovi kusnezovi (John, 1910)

(Map 8) (Figs. 70–78 (imago), 226, 227 (male genitalia), 276 (female genitalia))

John, 1910b: 606; Tab. 16, figs. 6 b, c, 7 a) ( Leucanitis kusnezovi). Type locality: “aus dem Syr-darja Gebiet: Perovsk… und den ihm nahegelegenen Ortschaften: Aj-darle… Baigakum… und Dshulek… aus dem Semiretshje Gebiet: Tsharyn… Transcaspien, Sumbar” (by the original description); “St. Baigakum, Syr-Darja” (by the lectotype designation, designated here).

Type material. Lectotype female (designated here in accordance with ICZN Art. 74) (Figs. 70–72), labelled: red square; golden circle; “specimen | typicum” (white paper, printed); “ St. Baigakum | Syr-Darja 12.V | 1908 Malyschew” (white paper, printed with handwritten date and month); “ Leucanitis | kusnezovi John. | O. John det.” (ZISP) . Paralectotypes: 14 males, 6 females (ZISP) .

Poole, 1989: 329; Klyuchko, Kreuzberg, 1998: 26 [Nuratinsky Nature Reserve]; Lehmann et al., 1998: 526 [33.37’N, 79.21’O, Sarytogai, Tal des Flusses Shaiyn, ca 20 km vor der Mündung in den Ili, 666m]; Poltavsky et al., 1998: 108 [Kara-Kala]; Gorbunov, 2011: 57 [southern part of Caspian depression; Mangyshlak; Ustyurt; Shagyrai plateau; Aral sea coast); pl. 8, fig. 3 (Onere)]; Gorbunov et al., 2011: 76 [Ustyurt Nature reserve, Onere dwell, 42°36’ N, 54°08’ E; Ustyurt Nature reserve, Kendyrli dwell, 42°57’ N, 54°41’ E]; Benedek, Bálint, 2013: 139 [Province Almaty, 15 km N of Masak, Ayak-Kalkan, 500 m, N 43° 45’, E 078° 30’]; Korb, 2013: 28 [Baschi]; Shovkoon, Trofimova, 2016: 607 [Aktobe Prov., Shagyrai Plateau, N 46°52’; E 58°02’; Aktobe Prov., Shagyrai Plateau, N 46°50’; E 57°47’; Aktobe Prov., Shagyrai Plateau, N 46°44’; E 57°30’; Mangystau Prov., N 46°05’; E 54°35’; Mangistau Prov., Mangistau Mts., N 44°18’; E 51°35’; Mangistau Prov., Ustyurt Plateau, N 43°55’; E 53°34’].

Habitat (Figs. 301, 303, 309). Deserts and semideserts, dry stony meadows in middle mountains. Elevation: 200– 2100 m.

Distribution. Deserts and semideserts of Central Asia and Kazakhstan, very local. It is possible that the species inhabit the sands near Astrakhan in Russia.

Material examined (in total: 52 specimens). Kazakhstan. 1 ♀, 2.07.1907, Semirechye Prov., Sary-Togai on Charyn river, leg. А. Yakobson, ex coll. O. John (ZISP) ; 3 ♂, 1 ♀, 18.04– 12.05.1908, Syr-Darja, St. Baigakum, leg. Malyschew, ex coll. O. John (paralectotypes) (ZISP) ; 1 ♂, 1 ♀, 7.06– 2.07.1908, Baigacum, leg. Androsow, ex coll. O. John (ZISP) ; 1 ♂, 1 ♀, 6.05.1908, St. Baigakum, Syr-Darja, Malyshew (paralectotypes) (ZMHU) ; 1 ♂, 1 ♀, 24.04.1909, prov. Syr-Darjensis, Perovsk circ., city tree station, leg. O. Miller (ZISP) ; 1 ♂, 27.04.1909, Syr- Darja, Dshulek, leg. Koshantshikoff, slide Matov 0531 (lectotype) (ZISP) ; 1 ♂, 23.04.1909, Syr-Darja, Dshulek, leg. Koshantshikoff, ex coll. O. John (paralectotype) (ZISP) ; 9 ♂, 4 ♀, 9.05– 6.08.1909, Syr-Darja, Aj-Darle, leg. Koshantshikoff, ex coll. O. John (paralectotypes) (ZISP) ; 1 ♂, Sir [darya], ex coll. Eversmann (ZISP) ; 1 ♀, 3.07.1911, Aral Sea, Yanu-Su river, leg. Molchanov (ZISP) ; 1 ♂, 2 ♀, 17.07– 7.08.1911, Syr-Daria, Baigacum, leg. Koshantschikoff (ZISP) ; 1 ♂, 1 ♀, 1915, Semirechye Prov., Dzharkent, leg. unknown, ex coll. Dyakonov (ZISP) ; 1 ♂, 1 ♀, 1– 10.04.1915, Semitechye, Dzhar-Kuduk S of Dzharkent, leg. V. Kavrigin (ZISP) ; 1 ♂, 20– 30.04.1915, Semirechye, Chulokai S of Dzharkent, leg. V. Kavrigin (ZISP) ; 1 ♂, 11.06.1916, Perovsky Uezd, Kara-Uzyan, leg. unknown, ex coll. Dyakonov (ZISP) ; 1 ♀, Kazaly distr., Basykara env., near r. Syr Daria, h= 67 m, 45.755º N 62.303º E, leg. A. Rokhletzova (ZISP) ; 1 ♂, 1 ♀, 15.07.1950, Aksu-Dzhabagly (ZMMU); 4 ♂, 18.07.1950, Dzhusaly, leg. unknown (ZISP) ; 1 ♂, 1 ♀, 14.07.1952, Syr-Darya, Dzhalagash, leg. Bundel (ZISP) ; 1 ♀, 3.08.1982, [ Central Of- fice of the Barsakelmes Nature Reserve, Barsakelmes island], leg. D. Piryulin (ZISP) ; 2 ♂, 1.05.2011, Altyn-Emel Nature Reserve, 30 km from Baschi, 43.96147N, 78.76941E, 645 m, leg. S.K. Korb (SKK) . Turkmenistan. 1 ♂, 1 ♀, 6.07.1910, Borly-Kul valley, leg. Zarudny (ZISP) . Uzbekistan. 1 ♀, 26.VII.1911, Nukus, leg. Molchanov (ZISP) .

Drasteria rada (Boisduval, 1848) (= D. roda (Herrich-Schäffer, [1851])

(Map 9)

Boisduval, 1848: xxx ( Microphisa rada). Type locality: “dans la Russie méridionale et surtout au pied du Caucase” (by the original description).

Type material. Not found.

Poole, 1989: 330; Beck, 1996: 25.

Distribution. South Russia (Volgograd and Orenburg Prov.), Kazakhstan, Caucasus and Transcaucasia, Asia Minor, Central Asia (deserts and semideserts), South Siberia and Mongolia.

Variability. There are four species group taxa described within this species: rada Boisduval, 1848, altivaga Alphéraky, 1894, schlumbergeri Fuchs, 1903, sibirica Kozhantshikov, 1924 . The status of only two of them is valid: nominate and altivaga. First one is widely distributed in South Europe, North and Central Kazakhstan and inhabits deserts and semideserts only; the second subspecies flies in the mountainous regions of Central Asia, Mongolia and South Siberia.