Disophrys Foerster, 1862

(Figs 3 B, 4B, 7B, 10A & B)

Disophrys Foerster, 1862: 246 . Type species: Agathis caesa Klug, by ICZN designation 1989: 76; Turner, 1918b: 225 [key, description]; Parrott, 1953: 200 [catalogue]; Shenefelt, 1970b: 391 [catalogue]; Bhat & Gupta, 1977: 267 [notes, catalogue]; Nixon, 1986: 190 [key, descriptions]; Chou & Sharkey, 1989 [key, description]: 183; Sharkey, 1992: 441 [in tribe Disophrini]; Yu et al., 2005 [catalogue]; Sharkey et al., 2006: 558 [notes, phylogeny, in tribe Disophrini]; Sharkey et al., 2009: 40 [key, description].

Pseudagathis Kriechbaumer, 1894: 65. Type species: Pseudagathis calabarica Kriechbaumer, 1894, by monotypy; Shenefelt, 1970b: 392 [synonym of Disophrys]; Chou & Sharkey, 1989: 183 [synonym of Disophrys].

Pseudocremnops Szépligeti, 1915: 152 . Type species: Pseudocremnops atripennis Szépligeti, 1915, by monotypy; Shenefelt, 1970b: 423 [catalogue]; Sharkey et al., 2006: 558 [synonym of Disophrys].

Platyagathis Turner, 1918a: 113 . Type species: Platyagathis leaena Turner, 1918 a, by monotypy; Parrott, 1953: 200; Shenefelt, 1970b: 423. syn. nov.

Diagnosis. Length: 5–12 mm; colour: commonly yellow and dark brown (Fig. 10 A), rarely with BROW colour pattern (Fig. 10 B); inter-antennal region with medial elevated mound that may be broadly rounded to carinate, paired carinae or protuberances may be present (Fig. 7 B); frons with lateral carinae (Fig. 3 B); notauli present, deeply impressed and scrobiculate (Fig. 3 B); tarsal claws cleft but base not pectinate (e.g., Fig. 2 A); mid tibial preapical spines absent; fore tibial spur not elongate (e.g., Fig. 2 F); pair of longitudinal carinae present on ventral surface of hind trochantellus (Fig. 2 G), although can be lacking in non-Australian members of the genus (e.g. Sharkey et al. 2009); fore wing cell 1–RS always present and sessile, distinctly quadrate, vein 2–2RS present (Fig. 4 B); ovipositor short, length less than 0.5 metasoma length (e.g., Fig. 9 D).

Comments. Most members of this genus exhibit variation in the yellow and dark brown colour pattern associated with a putative mimicry complex. However, an interesting aspect of this genus in Australia is the presence of two species exhibiting the BROW colour pattern. One species, D. leaena (Turner) (Fig. 10 B), represents the most southerly occurrence for Disophrys, occurring in the Southwestern region of the continent. It is nearly identical in colour and pattern to Therophilus latibalteatus (Cameron) (e.g., Fig. 8 C), including possessing a distinct propodeal setal field. The other Disophrys species exhibiting the BROW pattern occur on the opposite side of the continent in the Torresian region.

Species richness and distribution. Disophrys is well represented in Australia with over 20 species recognised, seven of which are described. Most of the species diversity occurs in the northern parts of the continent in the Timorian, Torresian and northern Kosciuskan regions. However, one species occurs in the temperate Southwestern region (see under Generic distribution and Comments above).