SCARABAEIDAE
Cetoninae
Gymnetini
Amithao Thomson, 1878
Amithao incertum (Gory & Percheron, 1833) . Blackwelder, 1944b:262 *Hisp.
Paragymnetis Schürhoff, 1937
Paragymnetis chalcipes undata Olivier, 1789 . Wolcott, 1927b:236; Blackwelder, 1944b:263
Paragymnetis lanius burmeisteri Schürhoff, 1937:63 . Blackwelder, 1944b:263
Dynastinae
Agaocephalini
Democrates Burmeister, 1847
Democrates croesus (Newman, 1836) . Blackwelder, 1944b:260
Cyclocephalini
Chalepides Casey, 1915
Chalepides barbata (Fabricius, 1787) . Blackwelder, 1944b:253
Cyclocephala Latreille, 1829
Cyclocephala laevis Arrow, 1937:40 . Blackwelder, 1944b:251
Cyclocephala notata (Illiger, 1806) . Wolcott, 1927b:233; Blackwelder, 1944b:252; Endrödi, 1966:267
Cyclocephala quisqueya Joly, 1998:50 *DR
Cyclocephala sanguinicollis Burmeister, 1847 . Endrödi, 1966:301
Cyclocephala striata haitiensis Endrödi, 1963:328 . Endrödi, 1966:315 *H
Dyscinetus laevicollis Arrow, 1937:41
Dyscinetus picipes (Burmeister, 1847) . Leng & Mutchler, 1914:442; Peck, 2005:73
Mimeoma Casey, 1915
Mimeoma nigra Endrödi, 1979:216 *DR
Dynastini
Dynastes Kirby, 1825
Dynastes hercules (Linnaeus, 1758) . Wetherbee, 1985a:83 [Probably extinct on the island]
Oryctini
Bothynus Hope, 1837
Bothynus bituberculatus (Palisot de Beauvois, 1805) . Blackwelder, 1944b:254 *DR
Hispanioryctes Howden & Endrodi in Howden, 1978
Hispanioryctes wittmeri Howden & Endrödi in Howden, 1978:389 *DR
Strategus Hope, 1837
Strategus aeneobarbus (Fabricius, 1775) . Blackwelder, 1944b:257 *H, *DR
Strategus barbigerus Chapin, 1932 . Schmutterer, 1990:92
Strategus inermis Arrow, 1947:223 *H
Strategus oblongus (Palisot de Beauvois, 1807) . Blackwelder, 1944b:257; Schmutterer, 1990:125
= Strategus quadrifoveatus (Palisot de Beauvois, 1807) . Blackwelder, 1944b:257
Strategus syphax (Fabricius, 1775) . Blackwelder, 1944b:257; Endrödi, 1985:613
Xyloryctes Hope, 1837
Xyloryctes jamaicensis (Drury, 1773) . Blackwelder, 1944:b255
Pentodontini
Endroedianibe Chalumeau, 1981
Endroedianibe bozzolai Chalumeau, 1981:221 . Chalumeau, 1983a:98 *DR
Tomarus Erichson, 1847
Tomarus cuniculus (Fabricius, 1801) . Cartwright & Chalumeau, 1978:27
Tomarus ebenus (De Geer, 1774) . Wolcott, 1927b:233
Phileurini
Hemiphileurus Kolbe, 1910
Hemiphileurus dispar (Kolbe, 1910) . Blackwelder, 1944b:259 *Hisp.
Hemiphileurus phratrius Ratcliffe & Ivie, 1998:205 *DR
Hemiphileurus ryani Ratcliffe & Ivie, 1998:201 *DR
Hemiphileurus scutellatus Howden & Endrödi in Howden, 1978:390 *DR
Homophileurus Kolbe, 1910
Homophileurus quadrituberculatus (Palisot de Beauvois, 1806) . Blackwelder, 1944b:258
Phileurus Latreille, 1807
Phileurus complanatus Palisot de Beauvois, 1809 . Blackwelder, 1944b:258 *DR
Melolonthinae
Key reference: Woodruff and Sanderson (2004).
Phyllophaga Harris, 1826
Phyllophaga aceitillar Woodruff in Woodruff & Sanderson, 2004:31 *DR
Phyllophaga alcoa Woodruff in Woodruff & Sanderson, 2004:33 *DR
Phyllophaga aliada Sanderson, 1951:279 . Woodruff & Sanderson, 2004:36 *H, *DR
Phyllophaga androw Woodruff in Woodruff & Sanderson, 2004:39 *DR, *H
Phyllophaga approxima Woodruff & Sanderson, 2004:42 *DR
Phyllophaga audanti (Wolcott, 1928c):27. Woodruff & Sanderson, 2004:45 *H, *DR
Phyllophaga baoruco Woodruff in Woodruff & Sanderson, 2004:45 *DR
Phyllophaga barrosa Sanderson, 1951:280 . Woodruff & Sanderson, 2004:50 *H
Phyllophaga bonfils Woodruff & Sanderson, 2004:52 *H
Phyllophaga canoa Sanderson, 1951:279 . Woodruff & Sanderson, 2004:54 *DR
Phyllophaga carnegie Woodruff in Woodruff & Sanderson, 2004:57 *DR
Phyllophaga cartaba Sanderson, 1951:274 . Woodruff & Sanderson, 2004:60 *DR
Phyllophaga eladio Woodruff in Woodruff & Sanderson, 2004:65 *DR
Phyllophaga espina Sanderson, 1951:264 . Woodruff & Sanderson, 2004:68 *DR
Phyllophaga esquinda Sanderson, 1951:268 . Woodruff & Sanderson, 2004:68 *H
Phyllophaga fossoria Sanderson, 1951:258 . Woodruff & Sanderson, 2004:68 *DR
Phyllophaga n. sp. nr. fossoria —Woodruff & Sanderson (2004):73
Phyllophaga garrota Sanderson, 1951:262 . Woodruff & Sanderson, 2004:73 *DR
Phyllophaga haitiensis Woodruff in Woodruff & Sanderson, 2004:76 *DR
Phyllophaga hogardi (Blanchard, 1850):137. Wolcott, 1927b:227; Woodruff & Sanderson, 2004:79 *H, *DR Phyllophaga imprima Sanderson, 1951:276 . Woodruff & Sanderson, 2004:81 *DR
Phyllophaga jaragua Woodruff in Woodruff & Sanderson, 2004:84 *DR
Phyllophaga jimenezi Woodruff & Sanderson, 2004:87 *DR
Phyllophaga kenscoffi (Wolcott, 1928c):24. Woodruff & Sanderson, 2004:90 *H, *DR
Phyllophaga larimar Woodruff in Woodruff & Sanderson, 2004:90 *DR
Phyllophaga latiungula (Wolcott, 1928c):28. Woodruff & Sanderson, 2004:95 *H
Phyllophaga leptospica Sanderson, 1951:262 . Woodruff & Sanderson, 2004:95 *H, *DR
Phyllophaga mali (Wolcott, 1928c):25. Woodruff & Sanderson, 2004:97 *H, *DR
Phyllophaga marcano Woodruff in Woodruff & Sanderson, 2004:97 *DR
Phyllophaga mella Sanderson, 1951:278 . Woodruff & Sanderson, 2004:103 *DR
Phyllophaga minutissima (Wolcott, 1928a):76. Woodruff & Sanderson, 2004:106 *H, *DR
Phyllophaga neglecta (Blanchard, 1850):137. Woodruff & Sanderson, 2004:108 *H, *DR
Phyllophaga nunezi Woodruff in Woodruff & Sanderson, 2004:110 *DR
Phyllophaga ortizi Woodruff in Woodruff & Sanderson, 2004:113 *DR
Phyllophaga panicula Sanderson, 1951:275 . Woodruff & Sanderson, 2004:117 *H, *DR
Phyllophaga pedernales Woodruff in Woodruff & Sanderson, 2004:117 *DR
Phyllophaga permagna (Moser, 1918):57. Woodruff & Sanderson, 2004:121 *H, *DR
Phyllophaga probaporra Sanderson, 1951:263 . Woodruff & Sanderson, 2004:123 *DR
Phyllophaga pseudocalcaris (Saylor, 1940):309. Woodruff & Sanderson, 2004:123 *DR
Phyllophaga rawlinsi Woodruff in Woodruff & Sanderson, 2004:126 *DR
Phyllophaga recorta Sanderson, 1951:268 . Woodruff & Sanderson, 2004:130 *H, *DR
Phyllophaga rex Woodruff & Sanderson, 2004:130 *DR
Phyllophaga romana (Saylor, 1946):1. Woodruff & Sanderson, 2004:137 *DR
Phyllophaga rustica Woodruff in Woodruff & Sanderson, 2004:137 *DR
Phyllophaga santachloe Woodruff in Woodruff & Sanderson, 2004:141 *DR
Phyllophaga toni Woodruff in Woodruff & Sanderson, 2004:143 *DR
Rutelinae
Anomalini
Anomala Samouelle, 1819
Anomala insularis Laporte, 1840 . Blackwelder, 1944b:244
Rutelini
Pelidnota MacLeay, 1819
Pelidnota sanctidomini Ohaus, 1905 . Leng & Mutchler, 1914:441; Blackwelder, 1944b:238 *Hisp.
Rutela Latreille, 1802
Rutela dorcyi (Olivier, 1789) . Leng & Mutchler, 1914:441; Blackwelder, 1944b:239 *H
Scarabaeinae
Aphodiini
Aphodius Illiger, 1798
Aphodius cuniculus Chevrolat, 1864 . Blackwelder, 1944b:212
Aphodius lividus Olivier, 1789 . Blackwelder, 1944b:212
Aphodius sallei Harold, 1863 . Blackwelder, 1944b:213
Rhyparus Westwood, 1845
† Rhyparus sp. —Wu (1997):187 [ Dominican amber]
Termitodius Wasmann, 1894
Ataeniopsis Petrovitz, 1973
Ataeniopsis armasi (Chalumeau, 1982) . Stebnicka, 2003a:109
Ataenius Harold, 1867
Ataenius aciculus Hinton, 1937:187 . Blackwelder, 1944b:213; Stebnicka, 2002:271 *H, *DR
= Ataenius apicecoloratus Balthasar, 1941:168 *H
Ataenius beattyi Chapin, 1940 . Stebnicka, 2006:107
Ataenius brevicollis (Wollaston, 1854) . Stebnicka, 2007:58
Ataenius elisaensis Stebnicka, 2002:270 *DR
Ataenius elongatus Palisot de Beauvois, 1805:104 . Blackwelder, 1944b:213; Stebnicka, 2002:276
Ataenius gracilis (Melsheimer, 1844) . Blackwelder, 1944b:214
Ataenius heinekeni (Wollaston, 1854) . Stebnicka, 2004:216
= Ataenius rhyticephalus (Chevrolat, 1964) . Stebnicka, 2004:216; Peck, 2005:85
Ataenius hispaniolae Chalumeau, 1982:325 . Stebnicka, 2002:269 *DR
Ataenius imbricatus (Melsheimer, 1844) . Blackwelder, 1944b:214; Stebnicka, 2003b:225
Ataenius jelineki Chalumeau, 1982:323 *H
Ataenius klapperichi Howden, 1978:378 . Stebnicka, 2002:279 *DR
Ataenius liogaster Bates, 1887 . Stebnicka & Lago, 2005:60
Ataenius luteomargo Chapin, 1940 . Blackwelder, 1944b:215
Ataenius morator Harold, 1869 [By synonymy of species below]
= Ataenius picipes Fleutiaux & Sallé, 1890 . Stebnicka, 2003b:243
= Ataenius tenebrosus Arrow, 1903 . Blackwelder, 1944b:216; Stebnicka, 2003b:243
Ataenius picinus Harold, 1867 . Cartwright, 1964:103; Peck & Thomas, 1998:59; Stebnicka, 2004:224
= Ataenius darlingtoni Hinton, 1937 . Blackwelder, 1944b:214
= Ataenius edwardsi Chapin, 1940 . Blackwelder, 1944b:214
Ataenius raccurti Chalumeau, 1978:50 . Stebnicka, 2002:276
Ataenius scabrellus Schmidt, 1909 . Stebnicka, 2003b:227
Ataenius scutellaris Harold, 1867 . Cartwright, 1978:14; Stebnicka, 2007:48
= Ataenius frater Arrow, 1903 . Blackwelder, 1944b:214
Ataenius strigicauda Bates, 1887 . Blackwelder, 1944b:215. Stebnicka, 2007:214
Ataenius versicolor Schmidt, 1916 . Blackwelder, 1944b:216; Stebnicka, 2002:273
= Ataenius haitianus Hinton, 1937:184 . Blackwelder, 1944b:216
Ataenius vincentiae Arrow, 1903 . Blackwelder, 1944b:216
According to Z. Stebnicka (pers. comm, 2007) Ataenius abditus (Haldeman, 1848), recorded by Blackwelder, 1944b:213 and Ataenius haroldi Steinheil, 1872, recorded by Blackwelder, 1944b:214 do not occur in Hispaniola.
Onthophagini
Digitonthophagus Balthasar, 1959
Digitonthophagus gazella (Fabricius, 1787) . MacRae & Penn, 2001:49; Ivie & Philips, 2007, in press [Introduced] Onthophagus Latreille, 1802
Onthophagus albicornis albicornis Palisot de Beauvois, 1805 . Blackwelder, 1944:211; Matthews, 1969:114 Onthophagus albicornis capitatus Laporte, 1840 . Matthews, 1969:114
Scarabaeini
Canthidium Erichson, 1847
† Canthidium sp. —Poinar & Poinar (1999):205 [ Dominican amber]
Canthochilum Chapin, 1934
Canthochilum ciboney Matthews, 1969:120 *H
Canthochilum darlingtoni Matthews, 1969:117 *DR
Canthochilum sp. —Matthews (1969):121
Canthon Hoffmannsegg, 1817
Canthon callosus Harold, 1868 . Wolcott, 1927b:227; Blackwelder, 1944b:198 *DR, *H
Canthon signifer Harold, 1868 . Wolcott, 1927b:227; Blackwelder, 1944b:201 *DR, *H
Canthon violaceus (Olivier, 1789) . Wolcott, 1927b:227; Blackwelder, 1944b:202 *DR, *H
Canthonella Chapin, 1930
Canthonella howdeni Ivie & Philips, 1990:432 *DR
Canthonella isabellae Matthews, 1966 . Ivie & Philips, 1990:432 *DR