Heterophrynus batesii (Butler, 1873)

Figs 3 A – F, 4 A – D, 15

Material examined.

Colombia • ♀ ♂, 1 juv.; Guaviare Department; Cueva Casa de Piedra, Serranía La Macarena / La Lindosa; 02°34'41.52"N, 72°52'21.67"W; altitude 195 m a. s. l. (IAvH -I-7874) . Caquetá Department • 2 ♀, 1 ♂, 4 juv.; Cueva Necaeridagoda (Cueva de Los Sueños), subcuenca del río Yarí; 02°34'41.52"N, 72°52'21.67"W; altitude 195 m a. s. l.; Carlos A. Lasso leg. (IAvH -I-7875) .

Records.

Colombia: Department of Huila, S. O. Acevedo. P. N. N. Cueva de los Guácharos, Cedros sector, Indio Cave; 1850 m a. s. l. (01°36'58.68"N, 76°06'15.12"W) (Víquez et al. 2014).

Remarks.

Heterophrynus batesii is widely distributed across the Amazon basin, and this record represents its westernmost known occurrence in Colombia, extending its range to the Andean foothills. The species appears to be troglophilic, as supported by previous records from Colombian caves. The morphology of the female gonopods (Fig. 4 C, D) largely corresponds to the descriptions and illustrations provided by Giupponi and Kury (2013). Unfortunately, the microsculpture of the male gonopods could not be confirmed due to the lack of SEM images, precluding detailed comparisons of this taxonomically critical feature.